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PROCEEDINGS. 


PROCEEDINGS 

OF   THE 

Jlogd  Sorittn   of  ilctona. 

VOL.   V    (New   Series). 

Edited    under    the    Authority    of    the    Council. 
ISSUED    MAY   1893. 


THE   AUTHORS  OK  THE  SEVERAL  PAPERS   ARE   SOLELY  RESPOKSIBLE   FOR    THE    SOUNDNESS  OF 
THE  OPINIONS   GIVEN  AND  FOR  THE  ACCURACY  OF  THE  STATEMENTS   MADE   THEUEIN. 


MELBOUENE : 
STILLWELL    AND    CO.,    PKINTEES,    195a    COLLINS    STREET. 


AGENTS    TO    THE   SOCIEIY: 

WILLIAMS  &  NORGATE,   14  HENRIETTA  STREET,   OUVENT  GARDEN,   LONDON 

To  wliom  all  communications  for  transmission  to  the  Royal  Society  of  Yicto:  ia, 
fiom  all  parts  of  Europe,  should  be  sent. 

1893. 


lanal  $0fictD  of  ^Tittorhi. 


18  9  2. 


fatron. 


HIS    lOXCELLENCY   THE   RIGHT    HON.    JOHN    ADRIAN    LOUIS   HOPE,    G.C.M.G. 
SEVENTH    EARL   OF   HOPETOUN. 

;|1rfsiDcnt. 

PROFESSOR   W.    C.    Ki:UN()T,    M.A.,  C.E. 
K.    J.    WHITE,    F.R.A.S.  |  H.    K.    RUSDCN,    F.R.G.S. 

|jon.   (Lrcasurcr. 

C.    R.    BLACKETT,    F.C.S. 

|)ou.  Secretaries. 

PROFESSOR   W.    BALDWIN   SPENCER,    JI.A. 
A.    SUTHERLAND,   M.A. 


I^on.  ICibradau. 

A.    DENDY,    D.SC. 

douiuil. 

W.  H.   ARCHER,  F.L.S.,  F.I.A.  A.   H.  S.  LUCAS,  M.A. 

.).  W.  BARRETT,  M.D.  PROFESSOR   L\LE,  M.A. 

K.   L.  J.  ELLERY,  C.M.G.,  F.R.S.,  PROFESSOR  ORME  MASSON,  M.A., 

F.R.A.S.  D.Sc. 

G.  S.  GRIFFITHS,  F.R.G.S.  H.  MOORS. 

.IAS.  JAMIESON,  M.D.  REV.   R.  H.  SUGDEN,   B.A.,  B..S.  . 

PROFESSOR   LAURIE,  LL.D.  C.   A.  TOPP,  M.A.,  LL.B. 


CONTENTS    OF    VOLUME    V. 


Akt.  I.— Preliminary  Notice  of  Victorian  Earthworms.  Part  II.  The 
genus  Peiichfeta  (with  Plates  II,  III,  IV,  V,  VI,  VII). 
By  W.  Baldwin  Spencer,  M.A.  ..  ..  -•  1 

I[.-  Fm'ther  Notes  on  the  Oviparity  of  the  larger  Victorian 
Peripatus,  generally  known  as  P.  Leuckartii.  By 
Abthuk  Dendy,  D.Sc.  . .  - .  . .  . .       27 

III.— Nest   and    Egg   of    Queen   Victoria's    Rifle    Bird    (Ptilorhis 

Victoriffi)  (with  Plate  I).     By  D.  Le  Souef    ..  ..       3fi 

IV.— Notes  on  the  Lilydale  Limestone  (with  Plates  VIII  and  IX). 

By  Rev.  A.  W.  Ckesswell,  M.A.      . .  . .  .  •        '^>i 

V. — Preliminary  Account  of  the  Glacial  Deposits  of  Bacchus 
Marsh  (with  Plates  X,  XI,  XII).  By  Graham  Officer, 
B.Sc,  and  Lewis  Balfour..  ..  ..  ..45 

VI. — Synopsis  of  the  Australian  Calcarea  Heterocosla ;  with  a 
proposed  Classification  of  the  Group  and  Descriptions 
of  some  New  Genera  and  Species.     By  A.  Dendy,  D.Sc.       (59 

VII.— On  Two  New  Tertiary  Stylasterids  (with  Plate  XIII).      By 

T.  S.  Hall,  M.A.  ..  ..  ..  ..  ..117 

VIII.— Three  rare  Species  of   Eggs  hitherto  only   described   from 

the  Oviduct  of  the  Bird.     By  A.  J.  Campbell,  F.L.S...     128 

IX.— Notes  on  the  Mode  of   Reproduction  of   Geonemertes  aus- 

traliensis.     By  Arthur  Dendy,  D.Sc.  ..  ..     127 

X.— The  Bluff  at  Barwou  Heads  (with  Plate  XIV).      By  G.  S. 

Griffiths,  F.G.S.  . .  . .  . .  . .  . .     131 

XI. — On  the  Conductivity  of  a  Solution  of  Copper  Sulphate  (with 

Plates  XV  and  XVI).      By  W.  Huey  Steele,  M.A.    . .      184 

XII.— The    Lichens    of    Victoria.      Part    I.      By    Rev.  F.   R.   M. 

Wilson      . .  . .  . .  . .  . .  . .      141 

XIII. — On  a  New  Species  of  Leueosoleuia  from  the  neighbourhood 

of  Port  Phillip  Heads.     By  Arthur  Dendy,  D.Sc.       . .     178 

XI  v.- The  Present  Position  of   the  Snake-bite  Controversy.      By 

James  W.  Barrett,  M.D.,  M.S.,  F.R.C.S.  Eng.  ..      181 

XV. — Sneezing:  Fallacious  Observations.     Bj'  James  W.  Barrett, 

M.D.,  M.S.,  F.R.C.S.  Eng.  . .  ..  ..  ..187 

XVI.— Physical  Constants  of  Thallium  (with  Plate  XVH).     By  W. 

Huey  Steele,  M.A.  . .  . .  . .  . .     193 

XVII. — On  "  Confocal  Quadrics  of  Moments  of  Inertia"  pertaining 
of  all  Planes  in  Space,  and  Loci  and  Envelopes  of 
Straight  Lines  whose  "Moments  of  Inertia"  are  Con- 
stant.    By  Martin  Gardiner,  C.E.  ..  ..     20(» 


40H0 


vi        Proceedings  of  the   Hu/ja(  tiocietij  of  Vicforia. 

XVIII. — Notes    on    a    PoiaououB    Species    of    Homeiia    (H.    colliua,    i.ai;i; 
Vent. — var.    miniata),    found    at    Pascoc    Vale,    causing 
death  in  cattle  and  other  animals  feeding  upon  it.     By 
D.  McAlpine  and  P.  W.  Farmer,  M.B.,  Ch.B  ..     209 

XIX.— Report  of  the  Committee  of  the  Royal  Society  of  Victoria, 
consisting  of  Prokessors  Kerxot,  Lylk  and  Masson, 
and  Messrs.  Elleky,  Love  and  White,  appointed  to 
arrange  for  the  carrying  out  of  the  Gravity  Survey  of 
Australasia  . .  . .  . .  . .  . .     21B 

XX. — Report  of  the  Cremation  Committee  of  the  Eoyal  Society 
of  Victoria,  appointed  to  enquire  into  and  report  upon 
"  Cremation "  and  other  methods  of  disposing  of  the 
dead,  with  particular  regard  to  hygiene  and  economy..     222 

XXL— Report  of   Port  Phillip   Biological   Survey  Committee,  1892  229 

Anntai.  Meeting,  Report  anp  Balance  Sheet     ..             ..  ..  230 

Report  of  Ordinary  Meeting.s                 . .             . .             . .  . .  239 

Laws  of  the  Royal  Society  of  Victoria              . .             . .  . .  289 

List  of  Members          . .             . .             . .             . .             . .  . .  299 

List  of  Institutions  and  Learned  Societies  which  receive 
Copies  of  the  "  Transactions  and  Proceedings  of  the 
KoYAL  Society  of  Victoria"  ..  ..  ..  ..     306 


Art.   l.—Prdlmlnarii  Xotice   of    Victorian    Kartlnvorni'i, 

Part  II.     Tha  (icnuH  Pevichceta.  _,„„-«-.,_ 

(With  Plates  II,  III,  IV,  V,  VI,  VII.)  /'^'y^^'^^  '^^^ 


Profi 


By  W.  BaldwixX  Spencer,  M.A.    i,^.j  h  I  B  R  A  R  Y  •  - 

essor  of  Biology  in  the  University  of  Melbouru'^II^N.    ^^     ^  '  y'c^ 
[Read  March  10,  1892.]  ^^^_J--^ 

This  ficcouiit  includes  tlie  description  of  twenty-two 
species  of  the  genus  Perichfeta,  which  have  up  to  the  present 
time  been  collected  in  Victoria.  Two  of  these,  Perichceia 
dorsalis  and  baker i,  have  been  previously  described  by 
Mr.  J.  J.  Fletcher,  who  obtained  them  from  Gippsland.  My 
own  collection  has  been  made  in  different  parts  of  Victoria, 
and  especiall}^  in  the  South  Eastern  district,  where  Gippsland 
is  peculiarly  rich.  As  in  the  case  of  the  genera  Cryptodrilus 
and  Megascolides,  so  in  that  of  Perichseta,  the  forms 
described  are  at  present,  for  the  sake  of  convenience, 
referred  to  the  one  genus,  though  this  will  undoubtedly 
have  to  be  broken  up,  and  at  the  same  time,  certain  forms 
provisionally  as  yet  referred  to  it,  may  have  to  be  placed 
under  other  existing  genera.  With  this,  Mr.  Fletcher  and 
myself  will  deal  in  our  extended  monograph.  Sufficient 
details  only  are  now  given  to  serve  for  the  identification  of 
the  species. 

We  have  in  Victoria  only  one  species  which  is  reall_y 
widely  distributed — this  is  P.  dorsalis,  which  was  first 
described  by  Fletcher  from  Warragul,  Gippsland.  Since 
then  it  has  been  obtained  in  West  and  South  Gippsland, 
from  the  Otway  district,  from  the  Grampians,  and  from 
Creswick  and  Castlemaine.  It  is  not  only  widely  spread, 
but  is  also  abundant  in  numbers,  almost  always  forming  the 
majority  of  specimens  of  any  collection  in  West  or  South 
GipjDsland  especially.  It  is  interesting  to  note  that  an  allied 
form,  Peric/iceta  stirlingi,  is  apparently  prevalent  in  South 
Australia. 


2  Proceedings  of  the  Royal  Society  of   Victoria. 

The  diNtiibution  of  the  former  species  is  in  marked  contrast 
to  that  of  most.  Certain  forms,  such  as  P.  tanjilensis  and 
yarraensis,  are  characteristic  of  the  Upper  Yarra  Valley 
especially.  Perichcvta  fielderi,  a  veiy  well  marked  form, 
has  only  come  from  Fern  Ti-ec  Gully  and  Sassafras  Gully 
and  the  hills  outside  Narre  Wan-en,  all  of  which  localities 
lie  within  a  small  compass.  P.  hakeri,  copelandi,  'and 
ohscura,  are  characteristic  of  the  Warragul  district  in 
Cippsland,  and  P.  deiulyi  is  an  intei'esting  form  recorded  as 
yet  only  from  Healesville.  P.  lateralis  has  been  found  only 
in  North  West  Victoria  and  is  closely  allied  to  the  Soutli 
Australian  species  P.  stirlingi. 

For  valuable  assistance  in  collecting,  I  am  again  indebted 
to  Dr.  Dendy,  Rev.  W.  Fielder,  and  Messrs.  French,  Frost, 
Shephard,  Hall,  Steel,  Mann,  Copeland,  Brittlebank,  D.  h 
Souef,  R.  H.  Anderson  and  H.  Giles. 

Unless  otherwise  stated,  the  description  alway?,  refers  to 
spirit  specimens.  Such  structures  as  the  accessory  copulatory 
ones  are  only  evident  after  preservation,  and  examination  of 
numerous  specimens  shows  that  spirit  exerts  a  uniform 
action  upon  these. 

(I)  Perlchceta  copelandi,  sp.  n.  (Figs.  52,  53,  54,  76).  Length 
of  spirit  specimen  o  inclies,  |  inch  broad.  Number  of 
segments  about  175, 

Dark  purple  colour  dorsally,  with  a  darker  median  line. 

Prostomium  completely  dovetailed  into  the  peristomiuni, 
and  marked  by  a  median  groove. 

Clitellum  not  strongly  marked,  occupying  segments  13-17, 
but  not  alwa^^s  the  anterior  part  of  1.}  or  the  posterior  of  17. 

Setpe.  The  first  setigerous  segment  has  10  on  each  side, 
after  this  and  to  the  posterior  end  of  the  clitellum  the 
number  varies  from  lo-]7.  Behind  segment  20,  it  varies 
from  23-25.  On  the  last  6  or  7  segments  the  setae  are 
difficult  to  see.  Dorsal  and  ventral  median  lines  free  of 
seta3. 

Male  poi'es  on  papillae  in  segment  18,  at  the  level  of  the 
interval  between  the  two  inner  seta^  of  each  side. 

Oviduct  pores  on  segment  14  anterior  to,  and  at  the  level 
of  the  interval  between  the  two  inner  seta;  of  each  side. 

Spermatl)ecal  pores,  5  pairs  placed  on  the  line  between 
segments  4-9,  very  slightly  dorsal  of  the  level  of  the  innei'- 
most  setse. 


PreUmiiiai'j/  Notice  of  Victorian  Earthtvornis.         3 

Accessoiy  copalatory  structures.  A  pair  of  elliptical  tumid 
patches  between  segments  16  and  17,  at  the  level  ot"  the 
interval  between  the  inner  two  setse  of  each  side.  A  pair 
between  segments  17  and  18,  at  the  level  of  the  interval 
between  the  second  and  third  sehB  of  each  side  ;  a  pair  at 
the  same  level  between  segments  18  and  19,  and  nioother 
between  segments  19  and  20.  The  male  openings  lie  ventral 
of  these  structures  and  not  dorsal,  as  in  the  case  of  the 
similar  ones  present  in  P.  hakeri.  A  series  of  ]iaii-s  of 
small  elliptical  patches  marked  by  distinct  pores  on  the  veiy 
anterior  margins  of  segments  9-13,  each  one  slightl}'  dorsal 
of  the  innei-most  setfe  of  its  side.  These  patches  in  segment 
9  are  enlarged  and  include  the  openings  of  the  spermatheca3. 
An  additional  pair,  with  similar  relationships,  may  be  present 
on  segment  8. 

Dorsal  pores  present,  the  first  between  segments  4  and  5. 

Kephridiopores  not  visible  externall}". 

Alimentary  canal.  Gizzard  present  in  segment  5.  No 
true  calciferous  glands,  but  vascular  swellings  are  present  in 
segments  9-15.     Large  intestine  connnences  in  segment  17. 

Circulatory  system.  Single  dorsal  vessel,  with  the  last  pair 
of  hearts  in  segment  12.     No  supra-intestinal  vessel  present. 

Excretory  system.     Meganephric. 

Reproductive  .system.  Testes,  two  pairs  in  segments  10 
and  11,  into  which  the  ciliated  rosettes  open. 

Prostates  long,  coiled,  and  tubuhir,  occupying  segments  1 8-22. 

Sperm  sacs,  three  pairs  attached  respectively  to  the 
posterior  wall  of  segment  9  and  the  anterior  of  segments 
12  and  13.     Saccular  in  foi-ni. 

Ovaries  in  segment  13,  with  oviducts  opening  into  the 
same  segment. 

Spermatheca3,  5  pairs  present  in  segments  5-9,  each 
consisting  of  a  large  sac  and  short  diverticulum. 

Habitat.  Warragul  district.  I  have  associated  with  this 
characteristic  Gippsland  ])ericha3te  the  name  of  Mr.  Hugii 
Copeland,  to  whom  I  am  much  indebted  for  frequent  and 
valuable  assistance  in  collecting. 

(2)  Pcrichwta  ohsciivd,  sp.  n.  (Figs.  4,  5,  G,  70).     Length  of 
spirit    specimen     2^    inches,    aljout     |    inch     br<)a<l. 
Number  of  segments  90-100. 
Prostomium  completely  dovetailed  into  the  peristomiuin. 
Clitellum  complete,  and  extending  over  segments  14-lG, 
together  with  the  posterior  part  of  13. 

B 


t  P I'oceediwjs  of  the  Royal  Society  of  Victoria. 

Seta',  from  U-11  on  each  side  in  front  of  the  clitellum  ; 
behind  this,  10-12  eacli  side.  Dorsally  and  ventrall}'  there 
is  a  median  space  free  of  setfe. 

Male  pores  on  segment  18  on  papillie,  on  a  level  with  the 
second  seta  on  each  side. 

Oviduct  pores  on  segment  14  anterior  to,  and  very 
slightly  ventral  of,  the  level  of  the  innermost  setfe. 

Spermathecal  pores,  five  pairs,  betv/een  segments  4?  and  5, 
5  and  G,  G  and  7,  7  and  8,  8  and  '.),  at  the  level  ot  the  second 
setfie  of  each  side. 

Accessory  copulatory  structures.  A  pair  of  elliptical 
tumid  patches  between  segments  IG  and  17,  at  the  level  of 
the  interval  between  the  two  inner  setse  of  each  side.  A 
pair  at  the  same  level  on  the  anterior  part  of  segment  18^ 
and  anocher  between  segments  17  and  18,  at  the  level  of  the 
interval  between  the  second  and  third  setse  of  each  side.  A 
swollen  tumid  ridge  occupies  the  posterior  part  ventrally  of 
.segment  18,  and  the  anterior  of  segment  19,  extending 
outwards  as  far  as  the  level  of  the  third  seta  of  each  side. 
A  ridge  on  the  posterior  part  of  segment  1 9,  and  anterior  of 
segment  20,  extending  outwards  as  far  as  the  level- of  the 
second  seta  of  each  side.  These  structures,  and  especially 
the  two  on  each  side  immediately  in  front  of  the  male 
openings,  are  very  characteristic. 

Dorsal  pores  present,  the  first  between  segments  4  and  5. 

Alimentary-  canal.  Gizzard  in  segment  5.  Ino  true 
calcifeious  glands  present,  but  vascular  swellings  in 
segments  12-15  ;  those  in  14  and  15  being  especially  large. 
Large  intestine  commencing  in  segment  17. 

Circulatory  system.  Dorsal  vessel  single  ;  last  heart  in 
segment  ]'2.  Those  in  segments  10,  11,  12  large;  those  in 
segments  7,  8,  9  small.  Supi'a-intestinal  vessel  in  segments 
9-12. 

Excretory  system.     Meganephric. 

Reproductive  system.  Testes,  two  pairs  in  segments 
10  and  11,  into  which  open  the  rosettes. 

Prostates  tubular,  coiled,  extending  through  segments 
17,  18,  and  19. 

Sperm  sacs,  two  pairs  attached  respectively  to  the  anterior 
wall  of  segment  12,  and  the  posterior  of  segment  9. 

Ovaries  in  segment  13,  the  oviducts  opening  into  the  same 
seo-ment. 


Preliminary  Notice  of  Victorian  Eavthivorms.         5 

Sperinathecae,  five  pairs,  in  segments  5-9.     Each  consisting 
of  a  sac,  with  a  diverticulum  half  the  length  of  the  sac. 
Habitat.     Warragiil,  Fern  Tree  Gully. 

(3)  Perichceta  sylvatica,  sp.  n.  (Figs.  34,  35,  36,  68).  Length 
of  spirit  specimen  3  inches,  less  than  -J  inch  broad. 
Number  of  segments  about  100. 

Prostouiium  not  completely  dovetailed  into  the  peris- 
tomium  (about  three-quarters).  The  peristomium  marked 
by  a  fairl_7  distinct  median  ventral  cleft. 

Clitellum  occupying  segments  14,  15,  16,  with"  the 
posterior  part  of  1  3,  and  the  anterior  of  17. 

Setse,  12  on  each  side,  except  in  the  first  two  setigerous 
segments,  where  there  are  10. 

Male  pores  on  segment  18,  slightly  ventral  of  the  level  of 
the  second  set?e. 

Oviduct  pores  on  segment  14  ventral  of,  and  very  slightly 
anterior  to,  the  innermost  setfe. 

Spermathecal  pores,  five  pairs,  between  segments  4  and  5, 
5  and  6,  6  and  7,  7  and  8,  8  and  9,  each  at  the  level  of  the 
innermost  seta. 

Accessory  copulatory  structures.  Two  pairs  of  fiintly 
marked  circular  patches  on  segments  16  and  17,  at  the  level 
of  the  interval  between  the  two  inner  setee  of  each  side.  A 
pair  of  well  marked  elliptical  patches  on  segment  10, 
posterior  to,  and  at  the  level  of,  the  interval  between  the 
second  and  third  setse  of  each  side. 

Dorsal  pores  present,  the  first  between  segments  5  and  6. 

Alimentary  canal.  Gizzard  in  segment  5.  Three  pairs  of 
well  marked  calciferous  glands  present  in  segments  10,  11, 
and  12.     Large  intestine  commencing  in  segment  15. 

Circulatory  system.  Dorsal  vessel  single,  the-  last  heart 
in  segment  12. 

Excretory  system.     Plectonephric. 

Reproductive  system.  Testes  in  segments  10  and  11, 
rosettes  opening  into  the  same  segments. 

Prostates  small,  flattened  and  bilobed,  but  with  a  single 
duct.     In  segment  18. 

Sperm  sacs,  two  pairs,  one  attached  to  the  anterior  wall 
of  segment  12,  the  other  to  the  posterior  wall  of  segment  9. 
Saccular  in  form. 

Ovaries  in  segment  10,  the  oviducts  opening  into  the 
same  segment. 


G  Proceedings  of  the  lioyal  Society  of  Victoria. 

Spermathecce,  five  pairs,  in  segments  5-9.  Each  consisting 
of  a  sac,  with  a  diverticukim  about  the  same  length  as  the 
latter. 

Habitat.     Fern  Tree  Gully. 

(4)  Ferichceta  ho(jrjll,  sp.  n.  (Figs.  28,  29,  80,  80).  Length 
of  spirit  specimen  4-J  inches,  ^  inch  broad.  Number 
of  segments  about  12o. 

Prostomium  not  completely  dovetailed  into  the  peristo- 
mium  (about  one-half).  The  latter  has  a  distinct  median 
ventral  clett. 

Clitelluui  occupying  the  posterior  part  of  segment  13, 
together  with  the  segments  14,  15,  \Q,  and  the  anterior  part 
of  segment  17. 

Seta3,  except  the  first  two  segments,  12  on  each  side. 

Male  pores  on  papillae  in  segment  18,  at  the  level  of  the 
innermost  seta  on  each  side. 

Oviduct  pores  on  segment  14  ventral  of,  and  anterior  to, 
the  le\'el  of  the  innermost  setse. 

Spermathecal  pores,  five  pairs,  between  segments  4  and  5, 
5  and  G,  6  and  7,  8  and  9,  at  the  level  of  the  innermost 
setif. 

Accessory  copulatory  structures.  Paired  tumid  patches 
slightly  anterior  to,  and  at  the  level  of  the  innermost  set?e 
in  segments  20,  21  and  22.  A  median  ventral  ridge,  occupy- 
ing the  space  between  the  two  innermost  papillse,  is  present 
on  segn:ients  17  and  19.  On  segments  20,  21,  and  22,  median 
ventral  ridges  are  usually  present,  connecting  the  cii'cular 
patches  across  the  median  line. 

Dorsal  pores  present,  the  first  between  segments  4  and  5. 

Alimentary  canal.  Gizzard  in  segments.  Truecalciferous 
glands  in  segments  10,  11,  12.  Large  intestine  commencing 
in  segment  15. 

Circrdatory  system.  Single  dorsal  vessel.  Last  heart  in 
segment  1 2.     Small  hearts  in  segments  G-9. 

Excretory  system.     Plectonephric. 

Reproductive  system.  Testes  in  segments  10  and  11, 
with  rosettes  opening  into  the  same  segments. 

Prostates  small,  flattened,  and  bilobed  with  a  single  duct. 

Sperm  sacs,  two  pairs ;  one  attached  to  the  anterior  wall 
of  segment  12,  the  other  to  the  posteiior  wall  of  segment  9. 

Ovaries  in  segment  13,  oviducts  opening  into  the  same 
seo-ment. 


Preliminary  Notice  of  Victorian  Eartkvjormn.         7 

Spermathecse,  5  pairs,  in  segments  5-9.  Each  witli  a  sac 
and  divevticulun)  more  tlian  three-quarters  the  length  of  the 
former. 

Habitat.     Mt.  Macedon  and  Healesville  (Dr.  Dendy). 

The  first  specimens  were  found  at  Mount  Macedon,  whilst 
collecting  with  Mr.  H.  R.  Hogg,  to  whom  I  am  indebted  jfor 
valuable  assistance,  and  whose  name  is  associated  with  this 
form. 

(5)  Pevichaita  haUii  (Figs.  40,  41,  42,  69).  Length  of  spirit 
specimen  If  iuclies,  a  little  more  than  ^t  irich  broad. 
Number  of  segments  about  100. 

Prostomium  not  completely  dovetailed  into  the  peris- 
tomium  (about  three-quarters).  The  peristomium  with  a 
distinct  median  ventral  cleft. 

Clitellum  not  \ery  strongly  developed,  lighter  coloured 
than  the  surrounding  parts,  and  occupying  segments  13-17. 

Seta3  somewhat  difficult  to  see.  In  front  of  clitellum, 
12  or  13  on  each  side  ;  behind,  12-16  on  each  side. 

Male  pores  on  very  strongly  marked  pa])ill8e  on  segment 
18,  the  openings  being  slightly  dorsal  to  the  level  of  the 
innermost  setre.  Between  the  papillae  is  a  deep  depression 
extenfling  on  to  segments  17  and  19. 

Oviduct  pores  (m  segment  14. 

Spermathecal  pores,  five  pairs. 

Accessory  copnlatory  structures.  Three  pairs  of  well 
marked  elliptical  tumid  patches,  each  with  a  median  linear 
depression  on  the  posterior  portions  of  segments  9,  10,  and 
1 1 ,  and  extending  over  a  space  equal  to  that  between  setee 
1  and  3  on  each  side.  A  smaller  i)atch  on  the  anterior  part 
of  the  segments  9  and  1 0,  confiuent  with  the  larger  posterior 
one,  but  not  so  strongly  marked. 

Dorsal  pores  present,  the  first  between  segments  4  and  5. 

Alimentaiy  canal.  Gizzard  in  segment  5.  True  calci- 
ferous  glands  present  in  segments  10,  11,  and  12.  Large 
intestine  conunencing  in  segment  17. 

Circulatory  system.  Dorsal  vessel  single.  Last  heart  in 
segment  12. 

Excretory  system.     Plectonephric. 

Reproductive  system.  Testes  in  segments  10  and  11,  into 
which  the  rosettes  open.  The  same  segments  are  filled  with 
sperm. 

Prostates_small,  fiattened,  and  bilobed,  but  with  a  single 
duct  on  each  side,  in  segment  18. 


8         Proceedincjs  of  the  Royal  Society  of  Victoria. 

Sperm  sacs  attached  to  the  anterior  wall  of  segment  12, 
and  tlie  ])osterior  wall  of  segment  9,  with  a  smaller  ])air 
attached  to  the  anterior  wall  of  segment  14. 

Ovaries  in  segment  13,  into  which  the  oviducts  open. 

Sj)ermatheca3,  5  pairs,  in  segments  5,  G,  7,  8,  and  9.  Each 
con.sisting  of  a  sac,  with  a  diverticulum  more  than  half  as  long 
as  the  former. 

Habitat.     Castlemaine.     Collected  by  Mr.  T.  S.  Hall,  M.A. 

(G)  Fenchceta  rubra,  sp.  n.  (Figs.  25,  26,  27).  Length  of 
spirit  specimen  2-|  inches,  breadth  about  -^  incli. 
Number  of  segments  80-90. 

Prostomium  not  completely  dovetailed  into  the  peristo- 
miuni  (about  |).  The  peristoinium  with  a  distinct  median 
ventral  cleft. 

Clitellum  distinct  and  complete,  occupying  segments  1 4-l(i. 

Setae,  in  front  of  the  clitellum  10  each  side,  behind  J  2. 

Male  pores  on  distinct  papilhe  on  segment  18,  at  the  level 
of  the  third  seta  of  each  side. 

Oviduct  pores  on  segment  1-i  anterior  to,  and  ventral  of, 
the  tirst  setse, 

Spermathecal  pores,  5  ]mirs,  between  segments  4  and  5, 
5  and  G,  G  and  7,  7  and  8,  8  and  9,  at  the  level  of  the  second 
seta  of  each  side. 

Accessory  copulatory  structures.  Median  ventral  ridges 
occupying  the  anterior  portions  of  segments  17,  19,  20,  21, 
22  and  23.  On  segment  10,  two  strongly  marked  circular 
patches,  with  central  depressions,  placed  posterior  to  the  seta:; 
and  at  the  level  of  the  interval  between  the  second  and 
third  setse.  On  segments  G,  7,  8  and  9  in  the  median  ven- 
tral space  devoid  of  setse,  and  anterior  to  the  level  of  the 
latter,  are  pairs  of  closely  apposed  circular  tumid  patches, 
each  with  a  central  pore-like  depression.  On  segment  5,  a 
median  ventral  patch. 

Dorsal  pores  present,  the  tirst  between  segments  4  and  5. 

Alimentary  canal.  Gizzard  in  segment  5.  True  cal- 
ciferous  glands  in  segments  10,  1 1  and  12.  Large  intestine 
commencing  in  segment  1 5. 

Circulatory  system.  Doi'«al  vessel  single.  Last  heart  in 
segment  12. 

Exeretor}^  system.     Plectonephric. 

Reproductive  system.  Testes,  two  pairs  in  segments 
10  and  11,  into  which  the  rosettes  open. 


Preliminary  Notice  of  Victorian  Earthworms.         9 

Prostates  small,  flattened,  and  bilobed,  but  with  a  single 
duct  in  segment  18. 

Sperm  sacs  attached  to  the  posterior  wall  of  segment  9, 
the  anterior  of  segment  12,  and  a  small  pair  to  the  anterior 
of  segment  14. 

Ovaries  in  segment  13,  Avith  oviducts  opening  into  the 
same  segment. 

Spermathecpe,  five  pairs,  in  segments  5-9.  Each  consisting 
of  a  large  sac,  with  a  diverticulum  of  about  the  same  length 
as  the  sac. 

Habitat.  Tallarook,  Goulburn  River.  Collected  by 
Mr.  A.  Ji.  S.  Lucas.  This  is  locally  known  as  the  "  red 
worm." 

(7)  Perlchceta  frencJdi,  sp.  n.  (Figs.  31,  32,  33,  79).  Lengtl) 
of  spirit  specimen  2-4  inches,  breadth  about  ^  inch. 
Nam  ber  of  segments  110. 

Spirit  specimens  are  dark  purple  colour  in  frorit  of  the 
clitellum,  save  on  the  mid-ventral  surface ;  dark  purple 
median  dorsal  line  ;  the  rest  of  the  body  a  dirty  wliite. 

Prostomium  not  completely  dovetailed  into  the  peris- 
tomium  (about  half).  The  peristomium  marked  by  a 
distinct  median  ventral  cleft. 

Clitellum  occupying  segments  13-16,  together  with  some- 
times the  anterior  part  of  .  7  dorsally. 

Setse,  10  each  side  in  front  of  the  clitellum,  behind  this 
12  each  side. 

Male  poi'es  on  papilhe  on  segment  18,  at  the  level  of  Uu' 
interval  between  the  two  inner  sette  of  each  side.  A 
distinct  depi'ession  between  the  two  papillae. 

Oviduct  pores  on  a  small  elliptical  patch  on  segment  14, 
anterior  to,  and  slightly  ventral  of,  the  innermost  setse. 

Spermathecal  poi'es,  five  pairs,  between  segilients  4  and  5, 
5  and  0,  G  and  7,  7  and  8,  8  and  9.  at  t!ie  level  of  the 
innermost  setse  on  each  side. 

Accessory  copulatory  structures.  Median  ventral  ridges, 
with  linear  depressions  on  the  anterior  parts  of  segments 
19,  20,  21  and  22.  Median  ventral  ridges  on  the  posterior 
parts  of  segments  9  and  10  ;  the  anterior  of  the  two,  small. 

Dorsal  pores  present,  the  first  between  segments  4  and  5. 

Alimentary  cana!.  Gizzard  in  segment  o.  True  calciferous 
glands  in  segments  10,  11  and  1 2.  Large  intestine  com- 
mencino;  in  seo-ment  1 5. 


10        Proceedlngis  of  the   lioyal  Societij  of  Victoria. 

Circulatory  system.  Dorsal  vessel  single,  the  last  heart  in 
segment  12. 

Excretory  system.     Ple3tonephric. 

Reproductive  system.  Testes  in  segments  10  and  11, 
rosettes  iu  the  same  segments. 

Prostates  flattened,  bilobed,  one  half  in  segment  18,  the 
other  in  segment  19  ;  each  half  with  a  .short  separate  duct, 
the  two  uniting  on  each  side  in  segment  18. 

Speim  sacs  attached  to  the  posterior  wall  of  segment  9, 
and  the  anterior  of  segment  12.     Saccular  in  form. 

Ovaries  in  segment  13,  into  which  the  oviducts  0[)en. 

Spermathecse,  five  pairs,  in  segments  5-9.  Each  consisting 
of  a  sac,  with  a  diverticulum  longer  than  the  latter,  and 
terminating  in  a  slightl}^  swollen  part. 

Habitat.  Loch,  8.  Gippsland  ;  Narre  Warren,  Waratah 
Bay  (Mr.  W.  Mann). 

Found  abundantly  under  logs  at  Nari'e  Warren  by  Mr. 
French  and  myself  I  have  much  pleasure  in  associating  with 
this  the  name  of  Mr.  French,  the  Government  Entomologist 
of  Victoria,  to  whom  I  am  much  indebted  for  frequent  and 
valuable  assistance.  This  is  one  of  those  forms  which  make 
burrows,  coming  to  the  su'-face  under  logs  and  stones,  in 
which  position  the  buri'ow  lies  open,  the  upper  surface  being 
closed  in  by  the  log  or  stone. 

(8)  Feric/urta  steelii,  sp.  n.  (Figs.  ."^7,  o8,  39).  Length  of 
spirit  specimen  2|  inches,  breadth  about  |  inch. 
Number  of  segments  about  120. 

Prostouiium  incompletely  dovetailed  into  the  peristomium 
(about  f).  The  peristomium  marked  by  a  distinct  median 
ventral  cleft. 

Body  dark  purple-ljrown  at  the  anterior  end,  with  a  dark 
median  dorsal  hue  along  the  body.  Light  brown  behind  the 
clitellum,  except  the  dorsal  portion  close  to  the  lattei'. 

SetiB.  The  first  3  segments  have  1 1  on  each  side,  after 
this  tl.tere  are  1 2  seta3  on  each  side. 

Male  poi'es  on  segment  12,  at  the  level  of  the  innermost 
setse,  but  not  on  papillae. 

Oviduct  pores  on  segment  14. 

Spermathecal  pores,  5  ])airs,  between  segments  4  and  5,  5 
and  6,  6  and  7,  7  and  8,  8  and  9,  at  the  level  of  the  second 
seta  of  each  side. 

Accessory  copulatoiy  structures  not  developed,  except  a 
slight  median  ventral  ridge  on  segment  17. 


PreliminaTy  N'otlce  of  Victorian  Earthivorms.       11 

Dorsal  pores  present,  the  tirst  between  segments  3  and  4. 

Alimentar}^  canal.  Gizzard  in  segment  5.  True  calci- 
ferous  glands  in  segments  10,  11  nnd  12.  Large  intestine 
commencing  in  segment  15. 

Circulatoiy  system.  Dorsal  vessel  single.  Hearts  in 
segments  5-12,  the  last  thiee  large. 

Excretory  system.     Plectonephric. 

Repi-oductive  system.  Testes  in  segments  10  and  11,  into 
which  the  rosettes  open. 

Prostates  small,  flattened,  l)ilobed,  but  with  a  single  duct  in 
.segment  18. 

Sperm  sacs  attached  to  the  anterior  wall  of  segment  1  2, 
and  the  posterior  of  segment  9. 

Ovaries  in  segment  1 3,  into  which  the  oviducts  o[/en. 

Spermathecae,  five  [)aii-s,  in  segments  5-9.  Each  consisting 
of  a  sac,  with  a  diverticulum  about  half  the  length  of  the 
formei-. 

Habitat.     Woodend.     Collected  by  Mr.  T.  Steel. 

(9j  Pericliceta  lateralis,  sp.  n.  (Figs.  55,  56,  57,  7S).  Length 
of  spirit  specimen  3-3^  inches,  breadth  less  than  ^ 
inch.     Number  of  segments  126. 

Prostomium  not  completely  dovetailed  into  the  peris- 
tomium  (about  |).  The  prostomium  marked  by  a  median 
ventral  cleft. 

Clitelluui  complete,  lighter  coloured  than  the  surrounding- 
parts,  occupying  segments  14,  15  and  16. 

Seta;  in  front  of  the  ciitellum,  10  or  11  each  side;  behind 
the  ciitellum,  10-12  each  .side. 

Male  pores  on  papillae  on  .segment  18,  at  the  level  of  the 
interval  between  the  second  and  third  setse  of  each  side. 

Oviduct  pores  on  segment  14  ph.ced  on  an  elliptical  patch, 
each  opening  almost  at  the  same  level  as,  and  very  slightly 
in  front  of,  the  innermost  seta  of  each  side. 

Spermathecal  pores,  three  pairs,  between  segments  6  and  7, 
7  and  S,  8  and  9,  at  the  level  of  the  flfth  seta  of  each  side. 

Accessory  copulatory  sti'uctures.  A  pnir  of  small  papillse 
each  immediately  in  front  of,  and  confluent  with,  one  of 
the  papiihe  bearing  the  male  opening.  In  addition  to  these, 
which  are  very  characteristic,  two  small  pairs  of  tumid 
patches  may  be  present  at  the  level  of  the  innermost  setae, 
one  half  on  segments  18  and   19,  the  other  half  on  segments 


12       Proceedings  of  the  Royal  Society  of  Victoria. 

]9  and  20,  and  another  pair  ma}'  be  present  at  tlie  level  of 
the  interval  between  the  two  inner  rows  of  setse  on  eacli 
side,  half  on  segments  9  and  10. 

Dorsal  pores  present,  the  first  between  segments  4  and  ■'). 

Alimentary  canal.  Gizzard  in  segnrient  5.  No  true 
calciferous  glands  present,  but  vascular  swellings  present 
in  segments  9-12.  Large  intestine  commencing  in  segment 
17. 

Circulatory  system  Dorsal  vessel  single.  Hearts  in 
segments  6-1  2. 

Excretor}^  system.     Plectonephric. 

Reproductive  system.  Testes,  in  [)airs,  in  segments  10  and 
1 1,  with  rosettes  opening  into  the  same  segments. 

Prostates  flattened  and  elongate,  with  a  somewhat 
mammillate  surface.  Each  is  leaf-shaped,  the  single  duct 
running  up  the  centre  in  the  position  of  a  mid-iib.  Extend- 
ing through  segments  18-21. 

Sperm  sacs,  three  pairs,  one  pair  on  the  anterior  wall  of 
segments  11  and  12;  another  on  the  posterior  wall  of 
segment  9.     Sac-like  in  form. 

Ovaries  in  segment  13,  into  which  the  oviducts  open. 

Spermathecce,  three  pairs,  in  segments  7,  8  and  9.  Each 
consisting  of  a  sac,  with  a  long  coiled  tubular  diverticulum. 

Habitat.  Castlemaine  (collected  by  Mr.  T.  S.  Hall). 
Tallarook,  Goulburn  Valley  (collected  by  Mr.  A.  H.  S.  Lucas.) 

(10)  Perichoita  dendyi,  sp.  n.  (Figs.  49,  50,  51,  77).  Length 
of  spirit  specimen  2i  inches,  breadth  about  -J 
inch.  Number  of  segments  about  IGO.  Colour 
yellowish  when  alive. 

Prostomium  completely  dovetailed  into  the  peristomium, 
the  former  being  distinctly  wedge-shaped. 

Clitellum  complete,  occupying  segments  14-16. 

Setee,  as  far  back  as  segment  19,  there  ai-e  6  on  each  side, 
arranged  in  pairs  ;  segment  20  has  8  each  side,  in  pairs, 
behind  this  increased  to  10  each  side,  and  at  the  posterior  end 
vary  from  7-10.  The  inner  two  on  each  side  I'emain  regular 
along  the  whole  length  of  the  body. 

Male  pores  on  slight  papillse  on  segment  18,  at  the  level 
of  the  interval  between  the  two  inner  seta?  of  each  side. 

Oviduct  ]wres  on  segment  14. 

Spermathecal  pores,  four  pairs,  between  segments  5  and  6, 
6  and  7,  7  and  8,  8  and  9,  at  the  level  of  the  innermost 
setae. 


Preliminary  Notice  of  Victorian  Earthworms.       13 

Accessory  copulatoiy  structures.  Narrow  tumid  ridges 
placed  on  the  mid-ventral  lines  between  segments  17  and 
18,  IS  and  19,  19  and  20.  The  single  ridge  may  be  divided 
into  two  halves,  the  centre  of  each  half  corresponding  with 
the  level  of  one  of  the  innermost  setce.  Special  small  tumid 
patches  are  constantly  present,  surroundin-g  the  openings  of 
the  two  posterior  pairs  of  spermathecae. 

Dorsal  pores  present,  the  first  between  segments  4  and  5. 

Alimentary  canal.  Gizzard  in  segment  5.  No  true  cal- 
ciferous  glands  present.  Vascular  swellings  in  segments 
9-12.     Large  intestine  connnencing  in  segment  IS. 

Circulatory  system.  Dorsal  vessel  single.  Hearts  in 
segments  8-12.  A  supra  -  intestinal  vessel  present  in 
.segments  9-12. 

Excretory  S3^stem.      Meganephric. 

Reproductive  s_ystem.  Two  pairs  of  testes  in  segments 
10  and  11,  into  which  open  the  rosettes. 

Prostates,  flattened  bodies  folded  over  the  intestine  on 
each  side  in  segment  18. 

Sperm  sacs,  two  pairs,  one  attached  to  the  anterior  wall 
of  segment  12,  the  other  to  the  posterior  wall  of  segment  9. 
Sac-like  in  form. 

Ovaries  in  segment  13,  with  oviducts  opening  into  the 
same  segments. 

Spermathecse,  four  pairs,  in  segments  G,  7,  S  and  9.  Each 
consisting  of  a  long  sac,  with  a  very  short  diverticulum  at 
its  base. 

Habitat.  Healesville  (collected  by  Dr.  Dendy),  living  in 
rotten  logs. 

I  liave  much  pleasure  in  associating  with  this  the  name  of 
Dr.  Dendy. 

(11)  Fericliwta  locheusis,  sp.  n.  (Figs.  1,  2,  3).     Length  of 
spirit  specimen  8  inches,  breadth  about  ^  inch. 

Prostomium  not  completely  dovetailed  into  the  peristomium 
(about  f). 

Clitellum  well-marked  and  complete,  extending  over  seg- 
ments 14-16,  and  including  also  the  posterior  part  of  segment 
13. 

Setae ;  the  usual  number  on  each  side,  as  far  back  as  the 
17th  segment,  is  9;  there  may  occasionally  be  11  ;  after  and 
including  the  19tli  segment  there  are  10.     A  small  posterior 


14        Proceediiujs  of  the  Royal  Soc'ietij  of  Victoria. 

part  of  the  body,  distinct  from  the  rest  by  its  ligiiter  colour 
and  flattened  shape,  dorso-ventrally,  has  16-Jl)  setiB  on  each 
side. 

Male  pores  on  slight  papillas  on  segment  18,  at  the  level  of 
the  interval  between  the  two  inner  set?e  of  each  side. 

Female  pores  on  segment  11  anterior  to,  and  ventral  of, 
the  level  of  the  two  innermost  seta?. 

Spermathecal  pores,  Ave  paii-s,  between  segments  1  and  5, 
5  and  6,  6  and  7,  7  and  8,  8  and  i),  difficult  to  see. 

Accessory  copulatory  structuies.  Segments  17  and  18  have 
their  mid-ventral  parts  tumid.  A  special  swollen  part  lies 
immediately  in  front  of  the  male  opening  on  each  side,  and 
there  are  tumid  ridges  ventrall}^  between  segments  18  and 
19,  19  and  20,  but  these  are  not  strongly  marked. 

Dorsal  pores  present. 

Alimentary  canal.  Gizzard  in  segment  5.  No  true  calci- 
ferous  glands,  but  vascular  swellings  are  present  in  segments 
12-15.     Large  intestine  commencing  in  segment  17. 

Circulatory  system.  Dorsal  vessel  single.  Hearts  in 
segments  8-1  2,  the  first  two  being  small. 

Excj-etory  system.  Meganephric.  Each  nephridium  has 
a  large  sac. 

Reproductive  .system.  Two  pairs  of  testes  in  segments  10 
and  11,  into  which  the  rosettes  open. 

Prostate"  coiled,  tubular,  occupying  segments  18-21. 

Sperm  sacs,  two  pairs,  one  attached  to  the  anterior  wall  of 
f5egment  12,  the  other  to  the  posterior  wall  of  segment  9. 

Ovaries  in  segment  13,  into  which  ti)e  oviducts  open. 

Spermathecae,  5  pairs,  in  segments  5,  6,  7,  8,  and  9,  each 
consisting  of  a  sac  and  diverticulum  less  than  half  the  length 
of  the  former-. 

Habitat.     Loch,  S.  Gip})sland,  under  logs. 

(12)  Perichcda  duhia,  sp.  n.  (Figs.  40,  47,  48,  07).  Length 
of  spirit  specimen  If  inches,  breadth  ^  inch.  Number 
of  segments,  about  100. 

Prostomium  completely  dovetailed  into  the  peristomium. 

Clitellum  extending  over  segments  13-17  ;  lighter  colour 
than  the  surrounding  parts  ;  not  thick  and  glandular,  and 
scarcely  noticeable  ventral ly. 

Sette.  The  first  setigerous  segment  has  Q  on  each  side, 
the  15  following  ones  have  8;  behind  this  the  number  varies 
from  9-12. 


Preliminary  Notice,  of  Victorian  Earthivorms.       15 

Male  pores  on  papillfe  on  segment  18,  at  the  level  of  the 
interval  between  the  two  inner  sette  of  each  side. 

Oviduct  pores  on  segment  14  slightl}^  anterior  to,  and  very 
nearly  on,  the  same  level  as  the  innermost  setfe  of  each  side. 

Spermathecal  pores,  five  pairs,  betv/een  segments  -i  and  5, 
5  and  6,  G  and  7,  7  and  8,  8  and  9,  at  the  level  of  the  intervals 
between  the  two  inner  set?e  of  each  side. 

Accessory  copulatory  structures.  These  resemble  somewhat 
those  of  P.  hakari,  but  the  two  forms  may  be  distinguished 
by  the  position  of  the  spermathecal  pores  and  the  form  of  the 
prostate.  Between  segments  17  and  18,  18  and  19,  and  19 
and  20,  are  three  pairs  of  elliptical  tumid  patches,  nearer  to 
the  median  line  than  the  male  pores. 

Dorsal  pores  present,  the  fii'st  between  segments  4  and  o. 

Alimentary  canal.  Gizzard  in  segment  5.  No  true 
caiciferous  glands,  but  vascular  swellings  in  segments  9-14. 
Large  intestine  commencing  in  segment  1 8. 

Circulatory  system.  Single  dorsal  blood-vessel.  Hearts 
in  segments  5-12,  those  in  segments  5-8,  small.  No 
continuous  supra -intestinal  vessel,  but  one  in  each  of  the 
segments  1 2-8  (?),  which  is  connected  with  the  dorsal  vessel 
in  the  posterior  part  of  the  segment,  and  ends  blindly  in  the 
anterior  part. 

Excretory  system.     Meganephric. 

Reproductive  system.  Two  pairs  of  testes  in  segments 
10  and  11,  into  which  the  rosettes  open. 

Prostates  coiled  and  tubular,  extending  thi-ongh  segments 
18  and  19,  the  blind  end  being  in  segment  18. 

Sperm  sacs,  two  pairs,  one  attached  to  the  anterior  wall 
of  segment  12,  the  other  to  the  })osterior  wall  of  segment  9. 

Ovaries  in  segment  13,  into  which  the  oviducts  open. 

Spermathecse,  five  pairs,  in  segments  5-9.  Each  consisting 
of  a  long  sac  with  a  short  diverticulum,  about  J  the  length 
of  the  former. 

Habitat.     S.  Warragul  (collected  by  Mr.  W.  Mann). 

(13)  Perichceta  ivalhalke,  sp.  n.  (Figs.  43,  44,  45,  6Q).    Length 

of  spirit  specimen  1   inch,  width  slightly  less  than 

^  inch.     Number  of  segments  88. 

The  form  of  the  body  is  bluntly  tapering  at  both  ends. 

Tliere  is  a  median  broad  dark  purple-brown  band  stai'ting 

immediately  behind  the  clitellum,  and  running  back  half 

way  to  the  posterior  end.     Tn  the  median  third  of  the  body 


16        Proceedings  of  the  Royal  Society  of  Victoria. 

the  lateral  surfaces  are  of  the  same  colour,  but  chequered 
with  little  rectangular  light  areas. 

The  prostomium  is  not  completely  dovetailed  into  the 
peristomium  (about  I,). 

Clitellum  well  marked,  tumid  and  complete,  occup3dng 
segments  1-i-lG. 

Setie,  in  front  of  the  clitellum,  10  on  each  side  ;  behind, 
12. 

Male  pores  on  segment  IS,  at  the  level  of  the  interval 
between  the  two  inner  setfe  of  each  side.  Between  the  two 
openings  a  ridge  runs  across  the  mid-ventral  surface,  with  a 
de]5ression  both  in  front  of,  and  behind,  it. 

Oviduct  pores  on  a  small  elliptical  ])atcli  on  segment  1 4, 
anterior  to  the  level  of  the  setse. 

Spermathecal  pores  difficult  to  see  externally. 

Accessory  copulatory  structures.     None  present. 

Alimentary  canal.  Gizzard  in  segment  5.  JS!o  true  cal- 
ciferous  glands.     Large  intestine  commencing   in    segment 

Circulatory  system.  Dorsal  vessel  single.  Last  heart  in 
segment  1 2. 

Excretory  system.     Meganephric. 

Reproductive  system.  Two  pairs  of  testes  in  segments 
10  and  1 1  ;  rosettes  opening  into  the  same  segments. 

Prostates  small,  tubular,  coiled,  in  segment  18. 

Sperm  sacs,  one  pair,  attached  to  the  anterior  wall  of 
segment  12.     Sac-like  in  form. 

Ov^aries  in  segment  13,  with  oviducts  opening  into  the 
same  segment. 

Spermathecre,  five  pairs,  in  segments  5-9.  Each  con- 
sisting of  a  long  sac  with  a  short  diverticulum. 

Habitat.     Walhalla  (collected  by  Dr.  Dendy). 

I  have,  unfortunately,  only  one  specimen  (a  mature  one) 
of  this  form.  But  the  shape  and  colouration  of  the  body, 
together  with  the  absence  of  accessory  copulatory  structures, 
render  it  so  distinct  from  other  forms  that  I  have  ventured 
to  distinguish  it  specifically. 

(14)  Perichwta  dicksonia,  sp.  n.  (Figs.  7,  8,  \)).  Length  of 
spirit  specimens  2  inches,  width  less  than  ^  inch. 

Prostomium  completely  dovetailed  into  the  peristomium, 
which  is  marked  by  a  median  ventral  cleft. 

Clitellum  well  marked,  complete,  occupying  segments 
14-16. 


Prelbnimtrij  Notice  of  Vidoritui  Earthworms.       17 

Setge,  10  on  each  side  in  front  of  the  cliteHum  ;  11  in  the 
middle  of  the  body.  The  rows  are  regnlarly  arranged  along 
the  body. 

Male  })ores  on  well  marked  i)a])illa3  on  segment  IS,  at  the 
level  of  the  second  seta  of  each  side. 

Oviduct  pores  on  segment  14-  anterior  to,  and  slightl}^ 
ventral  of,  the  level  of  the  innermost  setse. 

Spermathecal  pores,  five  pairs,  between  segments  4  and  o, 
5  and  6,  G  and  7,  7  and  8,  8  and  9  ;  slight!}^  dorsal  of  the 
level  of  the  innermost  setse  of  each  side. 

Accessory  copulator}'  structures.  Two  well  marked 
elliptical  tumid  jiatches,  one  immediately  in  front  of,  and 
the  other  imniediatel}^  behind,  the  male  openings.  The 
first  half  on  segments  17  and  18,  ard  the  second  half  on 
segments  18  and  19. 

Dorsal  pores  present,  the  first  between  segments  -i  and  5. 

Nephridiopores  on  the  anterior  margin  of  the  segments, 
at  the  level  of  the  sixth  seta  of  each  side,  commencing  on 
the  third  segment. 

Alimentary  canal.  Gizzard  in  segment  5.  No  true 
calciferous  glands.  Large  intestine  commencing  in  segment 
17. 

Circulatory  .system.  Dorsal  vessel  single.  Hearts  in 
segments  6-12. 

Excretory  system.     Meganephric. 

Reproductive  system.  Testes,  two  pairs,  in  segments 
JO  and  11,  into  which  the  rosettes  open.  The  testes  and 
rosettes  on  each  side  in  each  segment,  enclosed  by  a  mem- 
braneous bag  tilled  with  sperm. 

Prostates  tubular  and  coiled,  in  segments  18  and  19. 

Sperm  sacs,  two  pairs,  one  on  the  anterior  wall  of 
segment  12,  the  other  on  the  posterior  wall  of  segment  9. 

Ovaries  in  segment  13,  into  which  the  oviducts  open. 

Spermathecfe,  five  pairs,  in  segments  5-9.  Each  consisting 
of  a  large  sac  and  small  diverticulum,  not  more  than  one- 
third  the  length  of  the  former. 

Habitat.     Fern  Tree  Gully,  under  logs. 

(15)  Ptrichcda  alsojjhila,  sp.  n.  (Figs.  U),  11,  12).     Length 

of  spirit  specimen  1^-2  inches,  breadth  neai'l}^  |  inch. 

Number  of  segments  about  104?. 

Prostomium    not    completely    dovetailed    into    the    peris- 

tomium    (about    half).      The    peristomium    marked    by   a 

median  ventral  furrow. 

c 


]8       Proceedings  of  the  Royal  Society  of  Victoria. 

Clitellum  well  marked,  complete,  occupying  segments 
14,  15,  16. 

Setae,  in  front  of  the  clitellum,  10-11  each  side  ;  behind 
the  clitellum,  13  each  side.     The  rows  regular. 

Male  i)ores  on  two  fairly  well  marked  papillae  on  segment 
18,  at  the  level  of  the  interval  between  the  second  and  third 
setfe  of  each  side. 

Oviduct  pores  on  segment  14  slightly  anterior  to,  and 
ventral  of,  the  level  of  the  setae. 

Spermathecal  pores,  four  pairs,  between  segments  5  and  6, 
6  and  7,  7  and  8,  8  and  9,  at  the  level  of  the  innermost  seta3 
of  each  side. 

Accessory  copulatory  structures.  Two  ridges  with  swollen 
ends,  one  immediately  in  front  of,  the  other  immediately 
behinil,  the  male  openings.  Both  have  their  swollen  ends 
at  the  level  of  the  intei  val  between  the  two  inner  setae  of 
each  side.  One  is  placed  half  on  each  of  the  segments 
17  and  18,  the  other  half  on  each  of  the  segments  18  and  JO. 
A  smaller  ridge  with  two  confluent  swellings,  half  on  each  of 
segments  16  and  17,  and  situated  in  the  mid-ventral  space 
between  the  innermost  rows  of  setae.  Two  elliijtical  tumid 
patches  in  the  mid-ventral  space,  one  on  the  posterior  half  of 
segment  7,  the  other  on  the  posterior  half  of  segment  8. 

Dorsal  pores  present,  the  first  between  segments  4  and  5. 

Nephridiopores  very  prominent,  at  the  level  of  the  seventh 
seta  of  each  side,  and  placed  at  the  anterior  margin  of  each 
segment,  commencing  with  the  third.  On  contraction  in 
s]3irit,  the  body  wall  in  transverse  section  has  the  form  of  an 
upper  and  lower  half,  meeting  on  each  side  at  an  angle 
which  corresponds  in  position  to  the  nephridiopore. 

Alimentary  canal.  Gizzard  in  segment  5.  No  true 
calciferous  glands,  but  large  vascular  swellings  in  segments 
14  and  15,  and  smaller  ones  in  segments  9-13.  Large 
intestine  commencing  in  segment  17. 

Circulatory  system.  Dorsal  vessel  single.  Last  heart  in 
segment  12. 

Excretory  system.     Megan ephric. 

Reproductive  system.  Two  pairs  of  testes  in  segments 
1 0  and  1  ] ,  into  which  open  the  rosettes. 

Prostates  tubular,  coiled,  in  segment  18. 

Sperm  sacs,  two  pairs,  one  attached  to  the  anterior  wall 
of  segment  12,  the  other  to  the  posterior  wall  of  segment  9. 
Sac-like  in  form. 


Pveliininarij  Notice  of  Vlctorla>i  Earihw<>Tins.       IJ) 

Ovaries  in  seo-ineiit  13,  into  whieii  clie  oviducts  o|)en. 
Spennathecw,  four  pairs,  in  se^^ments  G-9.      Each  consist- 
ing of  a  large  sac  and  short  soniewliat  thin  diverticuhun. 
Habitat.     Fern  Tree  Gnlly,  under  logs. 

(IG)  PevhcludaJieUleri,  ^^.  n.  (Figs.  19,  20.  21,  G-t).  Length 
in  spirits  nearly  6  inclies,  breadth  \  inch. 
Both  when  alive  and  when  in  spirits,  the  worm,  has  not 
the  sliglitest  resemblance  in  appearance  to  an  ordinary 
pericha3te  form.  It  is  only  provisionally  referred  to  this 
genus.  The  body  is  cream  coloured,  with  a  thick  bright 
pink  coloured  clitellum,  and  is  quite  smooth,  there  being- 
no  indication  of  setse,  except  in  the  clitellar  region  and 
perhajjs  an  odd  one  here  and  there  posteriorly  ;  to  see  the 
setse,  it  is  neces.sary  to  cut  sections.  Its  general  ai)pearance 
is  closely  similar  to  that  of  a  Megascolides,  to  which  genus 
I  took  it  to  belong  when  collecting  it. 

Prostomium  possibly  completely  dovetailed  into  the  peris- 
tomium,  but  the  latter  is  strongly  ribbed,  two  grooves  being 
continuous  with  the  edges  of  the  prostomium,  which  has 
also  a  median  furi'ow  continued  on  to  the  peristomium. 

Setfe,  about  six  on  each  side,  irregularly  arranged  behind 
the  clitellum.  In  segments  13-lG,  a  pair  can  often  be  seen 
on  either  side  ventrall}^ 

Clitellum  well  marked  and  thick,  extending  over  segments 
13-18  ;  complete,  save  for  two  small  depressed  patches 
ventrally,  one  in  the  middle  of  segment  16,  another  occupy- 
ing the  hinder  part  of  segment  17  ventrall}^,  and  the  anterior 
of  18.  These  depressed  patches  ma}"  be  absent,  and  the 
clitellum  complete,  in  some  specimens. 

Male  pores  on  two  prominent  papilla?,  which  may  have 
their  inner  sides  confluent,  on  segment  18.  The  pores  at 
the  level  of  the  interval  between  the  two  inner  setfe  of  each 
side. 

Oviduct  pores  in  a  small  linear  depression  on  the  anterior 
half  of  segment  14,  each  pore  slightly  ventral  of  the  level 
of  the  innermost  setfe. 

Spermathecal  pores,  two  pairs,  one  on  the  posterior  margin 
of  segment  7,  anothei-  on  the  posterior  margin  of  segment  8. 
Each  pore  is  placed  on  a  small,  tumid,  elliptical  patch. 

Accessor}^  copulatory  structures.  An  elliptical  patch  ven- 
trally, half  on  each  of  segments  19  and  20,  a  similar  one  half 
on  each  of  segments  20  and  21.  Only  one  of  these  may  be 
present. 

C  2 


20       Proceedings  of  tite  Roijal  Society  of  Victoria. 

Dorsal  por-es  present. 

Alimentary  canal.  Gizzard  in  segment  15.  Vascular 
swellings  in  segments  11  and  12.  Calciferous  glands  some- 
what ventraliy  placed  in  segment  18.  Lai'ge  intestine 
commencing  in  segment  17.  Prominent  glandular  tufts 
(pepto-nephridia  ?)  attached  to  the  pharynx. 

Circulatory  system.  Dorsal  vessel  single.  Hearts  in 
segments  8-13.  A  continuous  supra -intestinal  vessel  in 
segments  9-14.     A  lateral  vessel  in  segments  7-13. 

Excretory  system.     Plectonephric. 

Reproductive  system.  Testes  not  visible,  but  a  large 
membranous  sac  on  each  side  in  segment  1 1  tilled  with 
sperm,  and  enclosing  a  prominent  rosette.  Probably  this 
encloses  also  the  testes. 

Prostates  flattened,  rather  small;  mammiilated  surface;  in 
segment  18.  A  large  whitish  swelling  close  to  each  duct, 
containing  penial  setae. 

Sperm  sacs,  one  pair,  attached  to  the  anterior  wall  of 
segment  12. 

Ovaries  in  segment  13,  into  which  the  oviducts  open. 

Spermathecse,  two  pairs,  in  segments  8  and  9.  Each  con- 
sisting of  a  large  sac  and  divei'ticulum. 

Habitat.  Navre  Warren.  Fern  Tree  Gully  (collected  by 
Rev.  W.  Fielder  and  Mr.  Mann).  Sassafras  Gully  (collected 
by  Mr.  She})hard).  Under  logs,  in  burrows  partly  exposed 
when  the  log  is  lifted,  and  })artly  penetrating  to  a  depth  of 
one  or  two  feet  beneath  the  surface. 

The  first  specimens  of  this  were  found  by  the  Rev.  W. 
Fielder  and  Air.  Shephard,  and  subsequently  Mr.  French 
and  myself  found  it  abundantly  at  Narre  Warren.  •  Its  area 
of  distribution  appears  to  be  very  limited,  as  I  have  never 
found  it  elsewhere,  or  received  it  from  other  districts. 

(17)  Ferichcvta  frost i,  sp.  n.  (Figs.  13,  14,  15,  71).     Length 
of  spirit  specimen   G   inches,   breadth  about  ^  incli. 
Number  of  segments  about  220. 
As  in  the  case  of  F.fielderi,  the  worm  has  not  the  slight- 
est resemblance  in  appearance  to  an  ordinary  pericha^te.     It 
resembles  closely  in  general  appearance  the  group  of  forms  at 
present  classed  together  under  the  genus  Qyptodrilus.     In 
spirit  the  body  is  bleached,  and   the   clitellum    of  a    light 
brown  colour. 

Prostomium  not  at  all  dovetailed  into  the  peristomium. 


Preliminary  Notice  of  Viclorion  Earthworms.      21 

Setre,  save  an  odd  one  here  and  there,  are  invisible. 

Clitellnra  strongly  marked,  saddle-shaped,  incomplete 
ventrally,  except  in  the  middle  of  segments  IG  and  17, 
extending  over  segments  14-17. 

Male  pores  on  very  prominent  papillae  on  segment  18. 

Oviduct  pores  placed  on  a  ridge  which  runs  across  the 
anterior  part  of  segment  14>. 

Spermathecal  pores,  five  pairs,  between  segments  4  and  5, 
5  and  C,  6  and  7,  7  and  8,  8  and  9. 

Dorsal  pores  present,  the  first  between  segments  3  and  4. 

Alimentary  canal.  Gizzard  in  segment  (3.  Calciferous 
glands  in  segments  8,  9  and  10.  Vascular  swellings  in 
segments  11,  12  and  13.  Large  intestine  commencing  in 
segment  15. 

Circulatory  S3'stem.  Dorsal  vessel  single,  as  far  back  as 
segment  13.  In  segment  14  and  succeeding  ones  to  the 
posterior  end,  it  is  double — that  is,  there  is  a  loop  in  each 
segment,  the  two  parts  uniting  at  the  septum.  Hearts  in 
segments  6-13.  A  lateral  vessel  is  present  on  each  side  in 
segments  8,  9  and  10. 

Exci-etory  system.     Plectonephric. 

Reproductive  system.  A  single  pair  of  testes  and  rosettes 
in  segment  1 1. 

Prostates  small,  flattened  ;  bi-lobed  ;  in  segment  18. 

Sperm  sacs.  A  pair  attached  to  the  anterior  wall  of 
segment  12,  and  a  smaller  pair  to  the  anterior  wall  of 
segment  13.     Sac-like  in  form. 

Ovaries  in  segment  13,  into  which  the  oviducts  open. 

Spermathecse,  five  pairs,  in  segments  5-9.  Each  consist- 
ing of  a  short  sub-spherical  sac,  with  a  blunt  rounded 
diverticulum  about  quarter  the  size  of  the  sac. 

Habitat.  Croajingolong,  E.  Gi]>psland.  Collected  during 
an  expedition  of  the  Field  Naturalists'  Club  of  Victoria  to 
Eastern  Gippsland.  I  have  associated  with  this  the  name 
of  Mr.  F]-ost,  to  whom  I  am  indebted  for  much  valuable  aid. 

(I8j  Perichcfita  goonriiiirh,  sp.  n.  (Figs.  16,  17,  18).  Length 
in  spirits  4i  inches,  breadth  ^e  ^^  '^^^  inch.  Number 
of  segments  about  150. 

The  body  is  dark  purple  colour  dorsally.  Laterally  it  is 
dark  purple,  but  chequered  with  little  rectangular  cream 
coloured  areas,  in  the   centre   of   each  of  which  is  a   seta. 


22        Proceedings  of  the  Royal  Society  of  Victoria. 

Cream  white  on  the  veiitiMl  surface.  The  colour  is  niucii 
the  same  in  spirit-preserved  animals,  as  in  the  fresh  state. 
This  form  is  provisionally  referred  to  the  genus  Perich^eta. 

Prostomiura  veiy  slio-htl y  dovetailed  into  the  peristomium. 

Clitellum  well  marked,  complete,  light  giey  in  colour, 
extending  over  segments  13-19. 

Seta3.  The  first  setigerous  segment  has  4  sette  on  each 
side.  The  second  5,  then  up  to  the  clitellum  there  are  (i. 
The  2()th  and  remaining  segments  have  8.  The  setfe  are 
irregularly  arranged,  save  the  inner  two  of  each  side. 

Male  pores  on  papilla?  on  segment  18,  each  slightly  ven- 
tral of  the  level  of  the  second  setse  of  each  side.  There  is 
a  marked  depression  immediately  in  fi-ont  of,  and  behind,  a 
median  ventral  ridge  on  segment  18. 

Oviduct  pores  on  segment  14  anterior  to,  and  very  slightly 
ventral  of,  the  level  of  the  iimermost  setee. 

Spermathecal  pores,  five  pairs,  between  segments  4  and  5, 
5  and  G,  6  and  7,  7  and  8,  8  and  9.  Each  slightly  ventral 
of  the  level  of  the  innermost  seta. 

Dorsal  pores. 

Alimentary  canal.  Gizzard  in  segment  5.  No  true  calci- 
ferous  glands,  but  vascular  swellings  in  segments  8-14,  those 
in  segments  13  and  14  smaller  than  the  others.  Large 
intestine  commencing  in  segment  16. 

Circulatory  system.  Dorsal  vessel  double  as  far  forward 
as  the  sixth  segment,  where  the  two  halves  do  not  unite 
anteriorly,  but  pass  forwards  on  to  the  surface  of  the  gizzard. 
In  each  segment  the  two  halves  unite  where  they  pass 
through  the  septum.  In  addition  to  the  dorsal,  there  is  a 
double  supra-intestincd.  vessel  in  segments  9-12.  Hearts 
in  segments  8-11.  In  segment  8,  the  hearts  arise  from  the 
dorsal  vessel.  In  segments  9-11,  they  arise  from  the 
supra-intestinal. 

Excretory  system.  Plectonephric,  associated  witli  large 
nephridia  with  internal  funnels  at  the  posterior  end  of  the 
body. 

Reproductive  system.  Two  pairs  of  testes  in  segments 
10  and  11.     Rosettes  doubtful. 

Prostates  flattened,  somewhat  fan-shaped  structures  in 
segment  18. 

Sperm  sacs,  a  single  small  pair  attached  to  the  anterior 
wall  of  seoinent  12.     Sac-like  in  form. 


Prelimlmcri/  Notice  of  Vlc'or'mn  EarthworrtiH.       28 

Ovaries  in  seo'ment  13,  into  which  the  oviducts  open. 

Spermathecpe,  five  pairs,  in  segments  -5-9,  gradually 
increasing  in  size  from  before  backwards.  Each  consisting 
of  a  sac,  with  a  short  blunt  rounded  diverticulum. 

Habitat.  Mt.  Goonmurk,  Croajingolong.  Whilst  collect- 
ing in  Croajingolong  I  only  found  this  interesting  form,  the 
colouring  of  which  renders  it  at  once  noticeable,  under  logs 
at  the  head  of  a  fern  gully  on  iVlt.  Goonmurk,  at  an  elevation 
of  about  3500  feet.  Mt.  Goonmurk  forms  part  of  the 
Dividing  Range  which  runs  from  east  to  west  across 
Victoria. 

(19)  Perlchceta  yarraensis,  sp.  n.  (Figs.  61,  62,  63,  74). 
Length  of  spirit  specimen  5i  inches,  of  living  form 
7  or  8  inches,  breadth  h  inch. 

In  life  the  body  is  of  a  dull  purple  colour,  darker  dorsally 
than  ventrally.  The  setpe  are  placed  on  little  lighter-coloured 
.spots.  The  clitellum  stands  out  very  clearl}^  in  the  living- 
form,  being  thick  and  cream  white  in  colour. 

P]-ostomium  completely  dovetailed  into  the  peristouiiuni. 

Clitellum  thick  and  strongly-marked,  and  extending  over 
segments  13-17.  The  ventral  surface  of  segments  15,  16 
and  17  is  not  always  white  and  glandular,  the  clitellum  here 
being  then  incomplete  and  saddle-shaped.  In  other  specimens 
it  is  complete. 

Setae.  The  first  two  setigerous  segments  have  four  on  each 
side,  arranged  in  two  couples.  Back  to  the  clitellum  there 
are  two  couples  on  each  side,  and  in  addition  a  fifth  one 
external  to  these.  Occasionally,  but  rarelj^  an  additional 
one  may  be  developed,  but  the  worm  can  be  recognised  by 
the  presence  of  five  setae  on  eajh  side,  regularly  an-anged  so 
far  back  as  the  clitellum  and  including,  at  any  rate,  the  two 
first  segments  of  this.  Worms  from  four  localities  all  show 
this  feature.  Behind  the  clitellum  the  number  increases  to 
10-14?  on  each  side,  arranged,  save  the  innei-most  one.  very 
irregularly.  There  is  left  a  broad  very  iiTegular  doisal  space 
fiee  from  set?e. 

Male  pores  on  slight  papilltB,  from  which  penial  settv  may 
iDe  seen  protruding,  on  segment  18,  at  the  level  of  the 
interval  between  the  two  inner  setae  of  each  side. 

Oviduct  pores  on  segment  14  anterior  to,  and  ventral  of, 
the  level  of  the  innermost  seta?. 


24-       Proceedings  of  the  Roijdl  Society  of  Victoria. 

Spei'uiatbecal  i)ores,  five  ])airs,  between  vsegiuents  4  and  5, 
5  and  (>,  G  and  7,  7  and  8,  8  and  !J,  at  the  level  of  tlie  inner- 
most setae. 

Accessory  copulatory  structures.  Three  pairs  ot  elliptical 
tumid  patches  in  front  of  the  male  openings,  and  three  or 
four  behind.  One  pair  placed  half  on  segment  17,  and  half 
on  segment  18,  at  the  level  of  tiie  interval  between  the 
second  and  third  seta  on  each  side.  The  others  placed  at 
the  level  of  the  interval  between  the  two  inner  set<e  of  each 
side,  and  placed  respectively  half  on  each  of  the  following 
segments,  15  and  16,  IG  and  17,  1.9  and  20,  20  and  21,  21 
and  22,  22  and  23.  Each  is  marked  by  a  median  linear 
depression. 

Doi'sal  pores  present,  the  tir.st  between  segments  5  and  G, 

Alimentary  canal.  Gizzard  in  segment  5.  No  true  calci- 
ferous  glands  present,  but  vascular  swellings  in  segments  13, 
14  and  lo.     Large  intestine  in  segment  17. 

Circulatory  system.  Dorsal  vessel  single.  A  supra-intes- 
tinal vessel  present  in  segment.-:,  10-13.  The  last  heart  in 
segment  12. 

Excretory  system.     Megan ephric. 

Reproductive  system.  Two  pairs  of  testes  in  segments  10 
and  1  I,  into  which  the  rosettes  open.  These  segments  are 
filled  with  mas.ses  of  sperm,  but  the.se  are  not  enclosed  in 
sacs. 

Prostates  coiled,  tubular,  oceupving  segments  18,  19  and 
20. 

Sperm  sacs,  three  pairs.  Two  huge  ones  attached  to  the 
anterior  walls  of  segments  12  and  13,  a  smaller  pair  attached 
to  the  posterior  wall  of  segment  9.     Sac-like  in  form. 

Ovaries  in  segment  13,  into  which  the  oviducts  open. 

SpermathecEe,  five  pah'S,  in  segments  5-9.  Each  consist- 
ing of  a  long  sac,  witli  a  short  diverticulum  about  one-fifth 
the  length  of  the  former. 

Habitat.  Tanjil  Ti'ack,  near  Wood's  Point.  Warragul. 
Warburton. 

(20)  Periclicda  tavjilensis.  Length  of  spirit  specimen  S-^- 
inches,  breadth  |  inch.  Tlie  worm  contracts  very 
much  in  spirits.  When  alive,  it  has  a  dull  grey 
purple  colour. 

Prostomium  completely  dovetailed  into  the  peristomium, 
and  marked  by  a  median  groove  continuous  with  one  which 
runs  alony;  the  mid-dor.sal  Tine  of  the  bod  v. 


Pveliminary  Notice  of  Victorian  Eartlnvorms.      25 

(Jlitellum  not  strongl}'  marked,  occupj'ing  segments  14—17, 
and  slightly  darker  than  the  surrounding  parts. 

Setfe.  The  number  and  arrangement  varies  slightl}',  hut 
the  following  description  of  a  particular  specimen  may  be 
taken  as  fairly  repi'eseutative  : —  The  first  six  setigerous 
segments  have  lour  on  each  side,  an-anged  in  couples.  The 
seventh  has  four  on  one  side  and  five  on  the  other.  T'he 
eighth,  four  on  each  side  ;  and  the  ninth,  four  on  one  side 
and  six  on  the  other.  The  tenth,  six  on  each  side.  Tlie 
eleventh,  four  on  one  side  and  six  on  the  other.  The  twelfth, 
four  on  one  side  and  seven  on  the  other.  The  thirteenth, 
five  on  each  side.  The  fourteenth,  six  on  each  side.  The 
lifteentli  and  sixteenth,  five  on  one  side  and  six  on  the  other. 
The  seventeenth,  .six  on  each  side.  The  eighteenth,  nine- 
teenth, twentieth,  twenty-fii-st,  and  twenty-second  segments 
have  eight  on  each  .side,  except  on  the  right  of  the  twentieth, 
wliere  there  are  siw  Behind  this,  the  number  varies  from 
(5-10  oii  each  si<le.  Tiie  two  inner  rows  on  eacli  side  are 
regularly  arranged,  except  during  the  posterior  third  of  the 
body.  The  doi'sal  interval  free  from  setpe  is  broad  and 
irregular. 

Male  pores  not  on  papilhie,  at  the  level  of  the  interval 
between  the  two  inner  setse  of  each  side,  on  segment  IS. 

Oviduct  pores  on  segment  14-  anterior  to,  and  slightly 
ventral  of,  the  level  of  the  innermost  seta3. 

Spermathecal  pores,  five  pairs,  between  segments  4  and  5, 
5  and  6,  G  and  7,  7  and  8,  8  and  9,  at  the  level  ot"  the 
innermost  set*. 

Accessory  copulatory  structures.  The  whole  of  the  ventral 
surface  of  segments  16-21  is  deeply  depressed  in  spirit 
specimens.  Four  pairs  of  small  elliptical  tumid  patches  are 
[)resent,  each  at  the  level  of  the  second  seta.  The  first  are 
placed  half  on  segment  16,  half  on  segment  17,  and  the 
remaining  three  respectively,  half  on  segments  17  and  18, 
1 9  and  20,  20  and  2 1 . 

i)or.sal  pores  present,  the  first  between  segments  4  and  5. 

Alimentaiy  canal,  Circulatoiy  system.  Excretory  system 
and  Reproductive  organs  similar  to  those  of  P.  yarraensis. 

This  form  is  closely  allied  to  the  latter,  but  the  presence 
in  F.  yarraeihsis  of  5  seta3  regularly  arranged  on  each  side  in 
front  of  the  clitellum,  renders  it  distinct  and  easy  to  recognise. 

Habitat.  Gembrook  (Mr.  D.  le  Souef),  Warlnnlon,  Tanjil 
Track  (near  Wood's  Point),  Fern  Tree  Gully,  and  Dandenong, 


26 


Froceedings  of  the  Royal  ^oc'ieti/  of  Victoria. 


(21)  Perichcda  hakeri'^  (Fletcher).     (Figs.  22,  23,  24,  75). 
Thi.s    form    was    first    obtained    by    Mr.    Fletcher   from 

Warragul.  It  is  ver}'-  abundant  there  under  logs,  and  is  a 
characteristic  Gipp.s]and  form  I  have  since  collected  it  in 
abundance  at  Warragul,  and  the  surrounding  district.  Fern 
Tree  Gully  and  Nane  Warren,  and  have  received  it  from 
Gembrook  (Mr.  D.  le  Souef),  and  Healesville  (Dr.  Dendy). 

(22)  Perichceta  dorsalisf  (Fletcher).     (Figs.  58,  59,  (50). 
This    form   was    tirst    obtained    by    Mr.    Fletcher   from 

Warragul.  It  is  present  in  a  much  greater  proportion  than 
any  ')ther  single  species,  and  has  come  to  me  from  very 
many  pai-ts  of  the  colony.  We  did  not  however  secure  it  in 
Croajingolong,  and  it  does  not  appear  to  extend  into  the 
east  and  north-east  parts  of  the  colony. 

Specimens  have  been  secured  at  Warragul,  Fern  Tree 
Gully,  Gembrook  (Mr.  D.  le  Souef),  Narre  Warren,  Dande- 
nong,  Creswick  (Mr.  J.  Fiddian),  Castiemaine  (Mr.  T.  S. 
Hall),  Myrniong  (Mr.  0.  Brittlebank),  Grampians  (Mr.  C. 
Frost),  Gerangamete  (Mr.  K.  L.  Anderson),  and  Waihalla 
(Mr.  H.  R.  Hogg). 


DESCRIPTION  OF  PLATES  II,  III,  IV,  V,  VI  &  VII. 
In  the  case  of  each  s[)ecies  one  drawing  represents  the 
external  anatomy,  a  second  the  alimentaiy  canal,  circulatory 
system  and  disposition  of  nephridia,  and  a  third  the  repro- 
ductive system.  On  Plate  VII  the  spermathecse  are  drawn 
in  outline  (under  the  camera  lucida  x  4).  Lines  represent 
the  position  of  the  two  inner  rows  of  setse  on  each  side. 

Reference  Letters. 
Ace.      Accessory   copulatory        Ocd.    Oviduct. 


structui 

es 

Pr. 

Prostate  gland. 

Cede. 

Calciferous 

glands. 

It 

Sperm  rosette. 

at. 

Clitellum. 

Sptk. 

Sperniatheca?. 

D.Bv. 

Dorsal  blood- 

vessel. 

T. 

Testis. 

Gz. 

Gizzard. 

ViiHC. 

Vascular  swellings  on 

Hts. 

Hearts. 

oesophagus. 

L 

Intestine. 

v.s. 

Sperm  sacs. 

Ov. 

Ovary. 

*  Proc.  Limi.  Soc,  N.S.W.,  Vol.  II  (Series  -indj,  Sootcmber  28,  1887, 
p.  61G. 

t  Proc.  Linn.  Soc,  N.S.W.,  Vol.  II  (Series  2nd),  September  28,  188/. 
p.  G18. 


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Art.    II. — Further  Notes  on  the    Ovipariti/  of  the    larger 
Victorian  Peripatus,  generally  liwivn  as  P.  leuckartii. 

By  Arthur  Dendy,  D.Sc. 

[Bead  May  12,  1832.] 


My  observations*  on  the  oviparous  habit  of  the  larger 
Victoi-iau  Perlpatus  (hitherto  generally  regarded  as  identical 
with  the  Peripatus  leuckartii  of  Sanger)  have  excited  a 
good  deal  of  hostile  criticism,  chiefly  emanating  from  the  pen 
of  Mr.  J.  J.  Fletcher.  On  three  different  occasions  since  the 
publication  of  my  notes  Mr.  Fletcher  has  brought  the 
question  before  the  Linnean  Society  of  New  South  Wales 
and  his  I'emarks  have  been  ))ublished  (I  do  not  know 
whether  in  full  or  not)  in  the  Abstracts  of  Proceedings  of 
the  Society.f 

I  have  already  replied  to  the  eai-lier  criticism^s  in  a  short 
paper  read  at  the  Hobait  meeting  of  the  Australasian 
Association  for  the  Advancement  of  Science,  which  will,  I 
am  informed,  be  published  shortl^^  Mr.  Fletcher's  latest 
observations,  however,  compel  me  to  return  to  the  question 
and  I  am  the  more  willing  to  do  so  as  I  have  some  further 
information  to  communicate  in  support  of  my  views. 

The  object  of  Mr.  Fletcher's  latest  contribution  to  the 
literature  of  the  subject  is  explained  in  the  opening  para- 
graph, which  runs  as  follows  : — "  This  paper  is  a  reply  to 
certain  views  expressed  by  Dr.  Dendy  with  regard  to  the 
reproduction  of  the  New  South  Wales  Peripatus,  which  on 
the  ipse  dixit  of  Dr.  Dendy  himself  is  P.  leuckartii,  Sang.; 
the  questions  at  issue  being  not  whether  or  no  the  Victorian 
Peripatus  is  oviparous,  but  whether,  firstly.  Dr.  Dendy  was 

*  Proc.  Eoval  Soc.  Victoria  for  1891,  p.  31;  Nature  and  Zoologischer 
Anzeiiier,  No.":-580,  1891. 

t  Septemljer  80,  1891;  February  24,  1892  ;  April  27,   1892. 


28        Proceed hig.s  of  the  lioyd.i  Society  of  Victoria. 

Justitied,  on  the  evidence  before  biui  and  in  the  absence  of 
an}''  pei'sonal  knowledge  of  the  reproduction  of  tlie  I^ew 
South  Wales  Fervpatu.s,  in  contradicting  statements  which 
were  quite  in  order  ;  and  secondly,  as  Dr.  Dend3''s  views 
were  published  in  September  ]891,  and  as  certain  informa- 
tion on  the  subject  was  subsequently  brought  under  his 
notice,  whether  it  is  not  now  nearly  time  that  Dr.  Dendy 
took  ste])s  to  explain  that  his  views  apply  wholly  and  solely 
to  the  Victorian  Peripatus,  and  to  withdiaw  his  insinua- 
tions respecting,  and  his  erroneous  interpretation  of,  '  Mr. 
Fletcher's  observations,'  because  already  Dr.  Dendy's  state- 
ments are  finding  their  wa^y  into  the  records  of  zoological 
literature,  and  confusion  and  nnsap})rehension  may  result 
therefrom." 

In  reply  to  Mr.  Fletcher's  indictment  I  wish  to  make  the 
following  remarks  : — 

(1)  I  di)  not  understand  the  meaning  of  the  statement 
that  the  New  South  Wales  Peripatus  is,  "on  the  ipm  dixit 
of  Dr.  Dendy  himself,"  P.  leuckaitii,  I  (;ertainly  am  not 
responsible  for  this  identification,  which  was,  I  believe,  first 
made  by  Mr.  OUiti*  who  remarks,*  on  first  recording  the 
animal  from  New  South  Wales,  that  "  the  species  is 
identical  witli  that  recently  recorded  by  Mr.  Fletcher  from 
Gippsland  and  is  probably  the  Peripatus  leiiclauiii  of 
Sanger."  I  need  scarcely  point  out  tiiat  the  name  leackartii 
has  since  been  applied  by  Mr.  Fletcher  himself  to  the  New 
South  Wales  species. 

Possibly  Mr.  Fletchei  means  to  i-efer  to  tlie  larger 
Victorian  species,  of  whicli  the  first  i-ecorded  specimen  was 
identified  by  himself  f  as  "  in  all  probability  an  example 
of  P.  leiiclurtii,  Sanger."  If  Mi-.  Fletcher  will  refer  to  my 
earliest  communication  on  the  subject:|:  he  will  find  that  in 
recording  the  discovery  of  two  specimens  at  Warburton 
(only  one  specimen  having  been  previously  recorded  from  this 
colony)  I  made  the  following  statement,  "after  carefully  study- 
ing Professor  Sedgwick's  full  description  of  P.  leuchartii,  I 
am  fairly  certain  that  they  do  not  belong  to  that  species,  but 
to  a  new  one,  which  I  for  the  present  refrain  from  naming," 
basing  my  conclusion  on  the  )-emarI<:able  pattern  of  the  skin. 

*  Proc.  Limi.  Soc.  N.S.W.,  Vol.  il,  p.  981. 
t  Proc.  Linn.  Soc.  N.S.W.,  Vol.  II,  p.  450. 
t    Victorian  Satnralixt,  January  ]889. 


The  Ovlparitjj  of  the  larger    Victorian  Peripatus.    29 

Professor  Sedgwick,  however,  in  reply  to  my  observations, 
expressed  the  opinion*  tliat  the  species  probably  was 
subject  to  a  considerable  range  of  variation  in  colour. 
Having  studied  more  specimens  I  myself  came  to  the  same 
conclusion^  and  liave  since  tlien  followed  Mr.  Fletcher  in 
calling  the  larger  Victorian  species  l\  leiickartii.  This 
use  of  tlie  name  leuckartii  on  my  part  seems  to  be  Mr. 
Fletcher's  chief  grievance  against  me  but  I  would  ask  him 
to  remember  that  I  have  onl_y  followed  his  own  lead  in  this 
respect. 

(2)  I  am  not  aware  that  I  have  contradicted  any  state- 
ments for  the  simple  reason  that  I  cannot  find  that  there 
wei'e  any  definite  statements  as  to  the  mode  of  reproduction 
of  the  New  South  Wales  Peripatus  for  me  to  contradict. 
There  was  merely  the  assumption  by  Mr.  Fletcher  (which  I 
quoted  and  characterized  as  very  natural)  that  the  3^oung 
animals  which  he  found  in  comj)any  with  the  parent  had 
been  born  alive. 

(3)  I  consider  that  I  was  fully  justified  in  assuming  that 
the  mode  of  i-eproduction  of  the  New  South  Wales  Peripatun 
was  the  same  as  that  of  the  Victorian  one,  as  at  the  time 
when  I  wrote  there  were  no  definite  observations  published 
as  to  the  mode  of  rejiroduction  of  the  former,  and  it 
was  almost  inconceivable  that  different  individuals  which 
Mr.  Fletcher  liimself,  in  common  with  all  other  writers  on 
the  subject,  regarded  as  belonging  to  one  and  the  same 
species  should  be  oviparous  in  the  one  colony  and  viviparous 
in  the  other.  I  have  no  doubt  now  that  the  New  South 
W •a\q?,  Peripatus  is  viviparous,  as  maintained  by  Mr.  Fletcher 
and  Professor  Haswell,  but  I  would  ask  Mr.  Fletcher  to 
remember  that  when  I  wrote,  the  only  publisiied  observations 
as  to  the  mode  of  reproduction  of  the  New  South  Wales 
species  were — {a)  the  finding  of  the  young  in  company 
with  the  mother,  though  there  was  notliing,  so  far  as  the 
published  account  goes,  to  show  that  they  had  not  been 
hatched  from  eggs  laid  for  some  time  ;  and  (/))  a  footnote:|: 
to  one  (jf  Mr.  Fletcher's  observations,  stating  that  a  female 
had  been  dissected  and  found  to  Ije  pregnant ;  the  term 
pregnant   is    not   defined    and    might,    in    my   opinion,    be 

*  Nature,  February  28,  1889. 

t  "  Observations  on  the  Australian  Species  of  Peripatus,"  Proc.  Royal 
See.  Victoria,  July  11,  1889. 

t  Proc.  Linn.  Soc.  N.S.W.,  Vol.  Ill,  p.  892. 


00  FroceecUuf/s  of  the  Royal  Society  of  Victoria. 

correctly  applied  to  a  female  containing  lai'ge  but  un- 
developed eggs  in  the  uterus  ;  nothing  is  said  l:)y  Mr. 
Fletcher  about  the  embiyos. 

Mr.  Fletcher  may  personally  have  had  abundant  evidence 
that  the  New  South  Wales  Feripatiis  was  viviparous,  but 
that  evidence  was  not  published  and  not  known  to  me  when 

1  wrote,  and,  therefore,  I  consider  that  I  was  quite  justified 
in  stating  that  the  mode  of  reproduction  of  F.  leuckartii 
was  unknown,  and  in  placing  my  own  interpretation  upon 
the  only  i-ecorded  facts  as  to  the  life  history  of  the  New 
South  Wales  form.  Naturally  1  interpretated  them  in  the 
Ught  of  my  own  observations  on  the  Victorian  species. 
That  interpretation  I  now  fully  admit  to  be  incoi-rect  and  I 
congratulate  myself  that  if  my  observations  have  had  no 
other  good  result  they  have  at  least  elicited  some  definite 
information  as  to  the  mode  of  reproduction  of  the  New 
South  Wales  Feripatiis. 

(4)  Mr.  Fletcher  seems  to  be  very  greatly  troubled  because 
my  statements  are  already  "  finding  their  way  into  the 
records  of  zoological  literature,  and  confusion  and  mis- 
apprehension may  result  therefrom."  There  is  not  the 
slightest  need  for  confusion  now  that  we  have  at  length  a 
definite  statement  as  to  the  reproduction  of  the  New  South 
Wales  species.  It  must  be  perfectly  obvious  to  ever}^  reader 
that  my  own  observations  were  based  entirel}^  on  Victorian 
specimens,  as  stated  distinctly  in  the  paper,  and  that  my 
suggestion  as  to  the  New  South  Wales  form  was  a  perfectly 
justifiable,  though,  as  it  turns  out,  incorrect  deduction  from 
the  only  published  facts.  It  is  perhaps  unfortunate  that 
both  the  New  South  Wales  and  Victorian  forms  should 
have  been  included  under  the  name  leuckartii,  but  for  this 
Mr.  Fletcher  himself  is  at  least  as  much  responsible  as  any 


(5)  Mr.  Fletcher  states  that  the  question  at  issue  is  not 
whether  or  no  the  Victorian  species  is  oviparous.  Herein  I 
must  beg  to  differ  from  him,  as  this  is  the  real  question 
which  I  have  been  all  along  trying  to  solve  and  compared 
with  which  the  mere  question  of  nomenclature  is,  in  my 
opinion,  insignificant  In  concluding  his  observations  he 
also  indulges  in  certain  offensive  and  unjustifiable  person- 
alities, which  I  need  not  ([uote.  It  is  greatly  to  be  regretted 
that  he  should  have  considered  such  a  proceeding  advisable 


The  Oviparlty  of  the  larger   Victorian  Perijyatus.    81 

and,  for  my  own  ]iart,  I  entirely  fail  to  see  the  advantage  to 
be  derived  therefrom  and  must  refuse  to  follow  his  example 
in  this  respect. 

Probably  the  solution  of  the  whole  difficulty  will  be 
found  to  lie  in  the  fact  that  my  original  opinion  was  correct 
after  all,  and  that  our  larger  Victorian  Feripatus  is  specifi- 
cally distinct  from  F.  leuclurtii.  For  the  present,  however, 
I  still  refrain  from  giving  it  a  distinctive  name,  as  I  have  had 
very  few  specimens  from  other  localities  to  compare  it  with 
and  do  not  wish,  if  it  can  be  helped,  to  create  a  new  species 
merely  on  account  of  the  oviparous  habit.  This  question, 
however,  is  discussed  in  my  communication  to  the  Austra- 
lasian Association  already  referred  to. 

As  to  the  oviparous  habit  of  our  larger  Victorian  s|:)ecies 
(so  called  to  distinguish  it  from  the  smallei-  F.  in.sigihis),  I 
have  some  additional  evidence  to  offer  and  I  would  like  at 
the  same  time  to  recapitulate  the  main  arguments  in  favour 
of  my  view.  My  critics  have  entirely  ignored  all  that  is 
new  in  my  observations,  such  as  the  remarkable  sculptured 
egg-shell,  and  have  suggested  that  what  I  have  observed  is 
simply  a  case  of  abnormal  extrusion  of  eggs  such  as  takes 
place  sometimes  in  P.  novce-zealandice.  Professor  Hutton, 
however,  who  made  the  observation  on  the  New  Zealand 
species,  merely  states  that  the  eggs  are  often  extruded  before 
development  is  complete  and  then  always  die.  Professor 
Sedgwick  quotes  these  statements  in  his  monograph  of  the 
genus  and  j^et,  in  replying*  to  my  letter  in  Nature,  he  states 
that  "  no  one  knows  whether  the  eggs  so  extruded  undergo 
complete  development."  I  suppose  that  most  animals  some- 
times extrude  eggs  which  never  complete  their  development, 
but  this  has  really  little  to  do  with  the  question.  What  I 
have  been  endeavouring  to  prove  is  that  the  larger  Victorian 
species  of  Feripatus  is  normally  oviparous.  The  two 
principal  arguments  originally  brought  forward — both  of 
which  have  been  entirely  overlooked  by  my  critics — were 
(1)  that  female  specimens  dissected  at  various  times  of  the 
year  were  never  found  with  embryos  in  the  uterus,  as  has 
been  so  frequently  described  for  otlier  species,  but  generally 
with  large  undeveloped  eggs  of  definite  oval  shape  and  with 
a  thick  membrane  ;  (2)  that  the  shell  or  membrane  of  the 
eggs  after  (but  not  before)  being  laid,  is  very  definitely  and 
characteristically  sculptured    on  the  outer  surface,  in  such 

*  Nature,  September  21,  1891. 


32        Froeeediuf/s  of  the  Royal  ^Society  of  Victoria. 

a  manner  as  to  recall  the  eggs  of  some  insects.  This 
sculpturing  alone  a))pears  to  me  to  indicate  a  truly  ovipai'ous 
habit,  and,  inasnmch  as  it  affords  anotiier  character  common 
to  Ferlpatiin  and  tlie  Jnsecta,  to  deserve  special  attention. 
I  am  not  aware  that  a  sculptured  egg-shell  has  hitherto  been 
observed  in  Feripatus  and  1  should  be  glad  to  learn  from  Mr. 
Fletcher  whether  anything  of  the  kind  has  ever  been  found 
around  embryos  of  the  New  South  Wales  species  which  have, 
as  he  informs  us,*  been  extruded  in  the  process  of  drowning. 

The  additional  evidence  on  the  subject  which  I  now  wish 
to  bring  forward  consists  in  the  subsequent  history  of  the 
fourteen  eggs  which  were  laid  in  my  vivarium  between  the 
18th  May  and  the  31st  July  last  year  and  of  one  which, 
though  possibly  laid  about  the  same  time,  was  not  discovered 
until  September  16.  Before  going  any  further,  however,  I 
may  premise  that  the  fact  that  the  eggs  are  really  those  of 
Feripatu.s  has  been  absolutely  i)r()ved  by  their  development. 
It  may  also  be  as  well  to  relate  the  fate  of  the  parent  animals 
by  which  the  eggs  were  laid. 

It  may  be  remembered  that  on  the  3 1st  July,  1S!>1,  when 
the  eggs  were  lirst  found,  there  were  in  the  vivarium  three 
females  and  one  male,  all  apparently  in  good  health.  The 
male  specimen  died  shortly  afterwards  but  on  August  17th 
the  females  were  still  all  alive  and  apparently  healthy.  On 
August  31st,  as  mentioned  in  a  postcript  to  my  first 
communication  on  the  subject,  one  of  the  female  specimens 
was  found  dead.  On  being  dissected  the  reproductive 
organs  appeared  very  well  developed ;  but,  although  the 
ovary  and  oviducts  were  both  large  (the  former  containing  a 
great  many  ovarian  eggs),  there  was  not  a  single  egg  in 
either  of  the  oviducts,  all  having  been  doubtless  laid. 

On  September  IGth  the  two  i-emaining  females  wo'e  still 
alive.  I  killed  and  dissected  one.  The  organs  appeared 
healthy  and  well  developed.  In  the  lower  part  of  each 
oviduct  one  large  egg  was  found.  The  eggs  presented  the 
usual  characters,  having  a  very  thick  but  unsculptured 
envelope  filled  with  yolk.  Both  eggs  were  cut  open  and 
examined  microscopically,  but  I  did  not  succeed  in  recog- 
nising any  trace  of  an  embrj^o  in  either. 

On  completely  turning  out  the  vivarium  and  examining 
its   contents   carefully,   I   found   one    more   Feripatus  egg 

*  Proc.  Linn.  Soc.  N.S.W.,  Sej.tember  30,  1891. 


The  Oviparlty  of  the  lair/er   Victoricui  Pevipatas.    33 

amongst  the  rotten  wood  (September  IG).  It  looked  much 
healthier  than  those  which  had  previously  been  transferred 
from  the  vivarium,  many  of  the  latter  having  already  begun 
to  shrivel  up  and  acquire  a  dark  colour.  \\\  the  newly  tound 
egg  and  also  in  the  healthier-looking  of  those  previously 
obtained  there  now  appeared  to  be  a  dark  spot  in  the 
interior,  but  this  was  only  dimly  visible  through  the  thick 
sculptured  shell. 

On  September  25th  the  last  remaining  female  was  still 
apparently  in  good  health  but  on  Octobei-  1st  it  was  found 
dead— how  long  it  had  been  so  I  do  not  know.  On  dissection 
I  found  the  internal  organs  in  a  bad  condition.  I^eitlier 
eggs  nor  embryos  were  visible  in  the  oviducts.  The  ducts  of 
the  slime  glands  were  very  much  enlarged  and  swollen  out, 
while  the  bi-anching  portions  appeared  feebly  developed,  in 
fjict  not  distinctly  recognisable.  The  alimentary  canal  was 
almcst  empty  and  the  animal  seemed  to  have  died  of 
•starvation. 

On  October  3  I  dissected  one  of  the  eggs  from  the  hatch- 
ing box.  I  could  find  no  embryo  in  it  but  only  the  same 
semi-liquid,  yolk-like  contents  as  when  in  utero,  full  of  little 
oil  or  yolk  globules.  Inside  the  thick,  sculptured  "shell" 
there  was,  as  usual,  a  very  thin  and  delicate,  transparent  mem- 
brane. Probably  a  young  embryo  was  really  present  but 
was  broken  up  in  opening  the  egg  and  overlooked  ;  even  at 
a  much  later  period  the  embiyonic  tissues  are  extremel}- 
delicate. 

On  November  30  I  noted  that  several  of  the  eggs  were 
shewing  indications  of  an  embryo  appearing  coiled  up 
within  them,  but  the  shell  was  so  thick  and  opaque  that  it 
was  impossible  to  make  out  any  details.  I  dissected  the  vgg 
which  was  found  on  September  1(5  and  which  had  since  then 
been  kept  separate  from  the  rest.  I  found  in  it  a  beautiful 
embiyo  Peripatus  in  an  advanced  stage  of  development. 
The  embryo  was  surrounded  by  a  delicate,  transparent 
membrane,  which  fitted  closely  on  to  it  and  was  very 
difticnlt  to  remove  ;  outside  this  came  the  sculptured  shell. 
The  emliryo  possessed  a  distinct  head,  with  clearly  recogniz- 
able brain,  eyes  and  ringed  antenna^,  and  there  were  at 
least  seven  pairs  of  appendages  behind  the  antennae.  It  lay 
tightly  coiled  up,  with  the  posterior  extremity  resting 
against  the  side  of  the  neck,  in  such  a  position  as  to  make  it 
very  difiicult  to  count  the  appendages.  The  specimen  was 
stained  and  mounted  in  Canada  balsam.     This  embryo,  then, 

D 


:}4'        Proceed  1 1  Kjs  of  the  Royal  Society  of  Victoria. 

developed  Ibv  more  than  ten  weeks  after  the  egg  had  heen 
laid  and  did  not  show  the  least  sign  of  "  going  to  the  bad." 

I  need  hcirdly  say  that  during  the  heat  of  the  summer 
months  I  found  it  a  very  difficult  matter  to  keep  the  eggs  in 
a  suitable  condition,  of  moisture,  especially  as  I  had  no 
])i'evious  experience  to  guide  me.  Hence  it  is  not  to  be 
wondered  at  that  the  majority  of  the  eggs  perished,  shrivell- 
ing u])  and  being  attacked  by  a  mould.  As  I  was  away 
from  Melbourne  for  some  weeks  during  the  summer  I  entrus- 
ted the  eggs  to  the  care  of  the  Rev.  W.  Fielder,  who  most 
kindly  looked  after  them  for  me  in  my  absence.  Fiequent 
attention  was  necessary  in  renewing  the  supply  of  nioisture. 

On  April  14,  1892,  only  three  eggs  remained  in  the 
hatching  box,  the  others  having  been  removed  as  they 
showed  signs  of  going  bad.  One  of  the  remaining  three  had 
been  showing  dark  ])igment  inside  for  some  days  ]iast.  This 
egg  I  removed  and  carefully  dissected.  I  found  the  shell  of 
a  much  darker  (yellow)  colour  than  when  laid,  a  good  deal 
crumpled  on  the  surface,  and  very  soft,  as  though  beginning 
to  decay  away.  The  cfintained  embryo  was  removed  and 
found  to  be  in  excellent  condition,  although  outside  it  there 
appeared  under  the  raicrosco])e  a  great  many  very  fine 
threads,  which  I  take  to  be  the  hypha;  of  a  fungus.  Possibly 
this  fungus  might  have  ultimately  killed  the  embryo  but  the 
latter  was  so  far  advanced  that  it  seemed  to  be  on  the  verge  of 
hatching.  It  was  enclosed  within  the  usual  transparent 
delicate  membrane  lying  within  the  thick  shell.  I  could  not 
determine  whether  the  fungal  hyphse  had  penetrated  within 
this  inner  membrane  but  I  think  it  very  doubtful.  The 
embr3'o  was  tightl}^  coiled  up  as  in  the  previous  case.  When 
uncoiled  it  measured  about  5  mm.  in  length  (exclusive  of  the 
antennse)  and  1  mm.  in  breadth,  ^^i  the  appendages  were 
developed,  viz.,  antennte,  oral  papillae,  two  pairs  of  jaws  and 
fifteen  pairs  of  claw-bearing  legs.  The  eyes  were  conspicuous 
at  the  bases  of  the  antennae,  and  the  antennae  themselves 
showed  each  about  twenty  deeply  pigmented  annuli.  The 
remainder  of  the  body  was  nearly  white,  but  very  distinct, 
isolated  pigment  patches  (chiefiy  indigo  blue,  with  a  few 
specks  of  orange)  appeared  scatteied  pretty  abundantly  over 
the  legs  and  back.  The  mouth  was  surrounded  by  the  very 
characteristic,  thick,  transversely  furrowed  Ii]3.  The  dermal 
j)apill?e  were  very  obvious  and  exhibited  the  characteristic 
spines,  the  cuticle  being  very  strongly  developed.  The 
claws  on  the  feet  were  very  distinct.     The  alimentary  canal 


The  Oviparitii  of  the  kmjer   Victorian  Peripatus.    35 

was  full  of  granular  food  yolk.  The  specimen  was  stained 
witli  borax  carmine  and  mounted  in  Canada  balsam. 

This  embryo,  then,  developed  for  at  least  eight  months  and 
a  half  after  the  egg  was  laid  and  at  the  end  of  that  time  was 
a  perfect  young  Perlpcttus,  differing  externally  from  the 
adult  only  in  its  smaller  size  and  less  deeply  pigmented 
skin. 

There  are  still  two  eggs  left  in  the  hatching  box  but  they 
do  not  look  to  me  at  present  as  if  they  were  going  to  hatch. 
Whether  they  do  so  or  not,  however,  I  think  I  may  fairly 
claim  to  have  now  definitely  proved  that  the  larger 
Victorian  Peripatus  at  any  rate  sometimes  lays  eggs,  and 
that  these  eggs  are  capable  of  undergoing  development 
outside  the  body  until  pei-fect  young  animals  are  produced. 
The  great  length  of  time  required  for  the  development  of 
the  eggs  is  very  remarkable,  but  it  is  only  what  one  might 
expect  on  considering  the  unusual  length  of  time  required 
for  intra-uterine  development  in  other  species. 


D  2 


Art.  III. —  Nest  and  Egg  of  Queen    Victoria  s  Rifle  Bird 
{Ptilorhis    Victorice). 

(With  Plate  I.) 

By  D.  Le  Souef. 

[Read  March  9,  1892] 


The  nest  and  egg  of  the  Victoria  rifle  bird  here  depicted, 
was  taken  on  one  of  the  Barnard  I.slands  by  Mr.  H.  Barnard 
and  myself.  We  visited  tliese  islands  on  November  18,  1891. 
in  quest  of  the  egg  of  this  bird,  and  built  our  small  humpy 
about  eight  feet  from  a  screw  palm  (Pandanus  aquaticus), 
which  grew  just  above  high  water  mark.  We  saw  a  pair 
of  rifle  birds  in  some  light  scrub  close  by  our  camp,  and  they 
seemed  very  fearless,  the  hen  bird  especially  so.  Next 
morning  was  spent  in  searching  over  this  interesting  and 
densely  timbered  island,  but  without  success.  During  the 
afternoon,  however,  we  determined  to  watch  the  hen  bird, 
which  was  seen  on  a  tree  close  by,  and  so  posted  ourselves 
one  on  each  side  of  the  aforementioned  patch  of  sscrub. 
The  bird  had  some  moss  in  her  bill,  which  she  kept  dropping 
and  catching  again  before  it  reached  the  ground,  and  we 
naturally  thought  she  was  building,  but  presently  she  darted 
down  into  the  scrub  close  by  Mr.  Barnard.  In  a  few 
minutes  he  saw  her  fly  into  the  screw  palm  by  our  camp,  in 
which  we  found  she  had  her  nest.  The  nest  itself  which  was 
built  near  the  crown  of  the  tree  about  seven  feet  from  the 
ground,  not  being  visible,  and  all  we  could  see  was  the  head 
of  the  bird.  She  continued  sitting  most  of  the  next  day, 
having  ap{)arently  become  accustomed  to  our  presence. 

We  then  took  the  nest  and  found  it  contained  only  one 
iigg,  which  was  hard  set,  the  chick  being  about  seven  days 
old.  The  nest  was  built  principally  of  vine  tendrils  and 
leaves  rather  loosely  put  together  (Plate  I). 


Proc.RS.VictorLa  Plate  1.  1692. 


0^ 


RWendel  dell  Uth. 


Troedel&CoPrin, 


Nest  and  E(jg  of  Queen  Victorias  Rifle  Bird.       37 

Tlie  ecjor  has  a  little  more  ijloss  on  than  is  shown  in  the 
illustration. 

Another  egg  and  nest  which  was  said  to  belong  to  this 
species  was  previously  sent  to  Mr.  C.  French,  in  1886,  from 
the  Cardwell  Scrub,  but  the  egg  was  spotted  instead  of 
being  streaked,  as  in  the  present  specimen  ;  it  is  also  smaller, 
and  the  nest,  although  made  of  somewhat  similar  material, 
is  not  so  large,  and  is  much  more  compactly  built.  Eggs  of 
the  same  species  of  bird  often  vary  considerably  in  colour, 
markings,  and  size,  but  still  the  general  characteristics  are 
the  same,  except  in  a  few  instances,  notably  the  egg  of  the 
Gymnorhina  tibicen.  On  several  occasions  I  have  noticed 
that  the  eggs  laid  by  one  pair  of  birds  are  almost  identical 
with  those  laid  by  the  same  pair  in  the  previous  year,  espe- 
cially in  the  case  of  sea-birds,  and  it  would  be  of  interest  to 
ascertain  if  this  fact  has  been  noted  by  other  oologists. 


Art.  IV. —  Notes  on  the  Lilydale  Limestone. 

(With  Plates  VIII  and  IX.) 

By  Rev.  A.  W.  Cresswell,  M.A. 

[.Head  July  14,  1892.] 


The  limestone  formation  of  Cave  Hill  quany  at  Lilydale, 
the  subject  of  this  paper,  is,  for  the  most  part,  a  hard  semi- 
crystalline  marble  deposit,  wedged  in  between  hard  qiiartzite 
on  the  one  .side, and  soft  shales  and  mudstones  on  the  other; 
and  has  for  many  years  ])ast  been  recognised  by  Professor  Sii' 
F.  M'Coy  and  others  as  of  Upper  Silurian  age,  of  about  the 
horizon  of  the  English  Wenfock,  fi'om  an  inspection  of  its 
contained  fossils.  The  limestone  strata  dip  to  the  east  at 
varying  angles  of  from  35°  to  50°,  the  strike  being  nearly 
north  and  south  magnetic,  var3'ing,  however,  on  the  east 
side  of  the  quarry  to  as  much  as  18°  east  of  magnetic  north. 
The  exact  thickness  of  the  limestone  is  not  as  yet  known, 
for  it  does  not  naturally  crop  out  on  the  surface,  but  is  only 
artiticially  exposed  by  quany ing.  As  early  as  1850,  the 
late  director  of  ouj-  Victorian  Geological  Survey,  Mr.,  now 
Sir  A.  R.  C.  Selwyn,  speaks*  of  the  limestone  as  known  to 
exist,  but  as  only  discoverable  b}^  a  well-like  hole  on  the  side 
of  the  hill,  leading  into  a  cave  hollowed  out  in  the  rock,  and 
sloping  down  to  a  depth  of  120  feet,  with  stalactites,  &c.  (a 
specimen  of  which  is  shown).  But  about  fifteen  years  ago, 
a  quarry  was  excavated  in  the  limestone  on  the  side  of  the 
hill,  and  the  opening  to  the  cave  is  now  covered  up  with 
debris,  and  is  inaccessible. 

The  progress  of  quarrying  has  now  proved  the  limestone 
to  be  of  much  greater  thickness  than  was  at  first  reported.-f- 
The  measurement  across  the  outcrop  is  about  5  chains,  or 

*  Eeport  on  Geological  Structure  of  Colony  of  Victoria :  Basin  of 
Yarra,  &c.,  1856. 

t  "  Victorian  Naturalist,"  188o,  II,  No.  3,  p.  35. 


Notes  oil  the  Llhjdale  Limestone.  89 

830  feet,  and  allowing  for  the  average  dip  of  40°,  this  v.-ould 
mean  a  vertical  thickness  of  about  220  feet,  but  this  is  only 
so  far  as  it  is  at  present  exposed.  Its  eastern  limit  ma}'  be 
considered  to  be  about  already  reached,  for  almost  immedi- 
ately flanking  it  on  that  side  may  be  seen  an  extensive  series 
of  quartzite  and  conglomerate  strata,  running  conformably 


CORRIGENDA     ET     ADDENDA. 

Page  39.  -For  (See  Fig.  1),  read  (See  Fig.  9,  Plate  IXa). 
For  (See  Fig.  2),  read  (See  Fig.  10,  Plate  IXa). 
,,     41. — Read    "  Pleurorbynehus    costatus "    and     "  Pleurorhynchns 
bellulus"   under   Lamellibranchiata,    instead    of   under 
Gasteropoda. 
Plate  VIII.— 1.    Tremanotus  pritchardi. 

2.  Eunema  etiieridgei. 

3.  Stomatia  antiqua. 
IX. — 4.    Tryblidium  nycteis. 

5.  Pleurorhynchns  costatus. 

6.  Pleurorhynchus  bellulas. 

7.  Naticopsis  lilydalensis. 

8.  Ambonychia  tatei. 


appeals  in  uie  quarry,  wiiea  juokcu  ac  asa^wiioje,  aim  iiom 
a  distance,  is  cream,  or  almost  white,  especially  on  weatiieied 
exposures;  but  when  freshly  broken,  it  is  of  different  shades 
of  dark  or  light  bluish  grey,  ])inkish  brown,  or  grey  with 
pinkish  brown  patches,  it  is  for  the  most  part  semi-crystal- 
line, is  here  and  there  somewhat  l.)recciated  on  a  snifdl  scale, 
and  is  in  some  ])laces  roughly  oolitic  (a  slide  of  an  oolitic 
specimen  on  view). 


Art.  IV. —  Notes  on  the  Lilydale  Limestone. 


debris,  and  is  inaccessible. 

The  progress  of  quarrying  has  now  proved  the  limestone 
to  be  of  much  greater  thickness  tlian  was  at  first  reported.f 
The  measurement  across  the  outcrop  is  about  5  chains,  or 

*  Eeport  on  Geological  Structure  of  Colony  of  Victoria :  iJasin  of 
Yarra,  &c.,  1856. 

t  "  Victorian  Naturalist,"  1885,  II,  No.  3,  p.  35. 


Notes  0)1  the  L'dydale  Limestone.  S9 

330  feet,  and  allowing  for  the  average  dip  of  40°,  tiiis  would 
mean  a  vertical  thickness  of  about  220  feet,  but  this  is  only 
so  far  as  it  is  at  present  ex])osed.  Its  eastern  limit  may  be 
considered  to  be  about  alread}'-  reached,  for  ahnost  immedi- 
ately flanking  it  on  that  side  may  be  seen  an  extensive  series 
of  quartzite  and  conglomerate  strata,  running  conformably 
with  the  limestone ;  but  on  the  west  it  is  not  yet  deflned,  as  it 
is  still  partly  covered  over  with  basaltic  soil  and  decomj)osing 
basalt.     (See  Fig.  I.) 

About  half  a  mile  off,  or  rather  less,  to  the  west,  and  on 
the  other  side  of  the  valley,  are  a  series  of  sandstones,  shales, 
and  mudstones  exposed  beneath  the  basalt  in  the  jyielbourne 
Road  cutting,  and  seen  to  be  dipping  in  the  same  direction 
as  the  limestone. 

The  lateral  extension  of  this  Lilj^dale  limestone  is  a 
matter  of  much  uncertainty.  It  is  generally  believed  to  be 
like  most  other  Silurian  limestones,  and  especially  in  Vic- 
toria, a  mere  lenticular  or  cake-like  patch  that  ra])idly  thins 
out  in  all  directions,  but  as  the  country  in  the  neighbour- 
hood is  completely  covered  over  with  soil,  trees,  and  verdure, 
it  must  remain  more  or  less  a  matter  of  conjecture,  until 
someone  is  enterpiising  enough  to  prospect  the  country,  if 
not  with  a  diamond  drill,  at  least  with  a  geological  cheese- 
borer.  Its  uovtltem  extension  has  not  been  observed  at  all, 
but  what  looks  somewhat  like  a  southern  extension  of  it 
may  be  seen  in  the  shape  of  two  bosses  of  the  same  kind  of 
limestone  in  the  side  of  the  railway  cutting,  about  three- 
quarters  of  a  mile  towards  Melliourne,  and  somewhere  about 
on  the  line  of  the  strike  of  the  limestone  at  Cave  Hill  quai'ry. 

As  it  is  hardl}^  conceivable  that  the  great  thickness  of  the 
Cave  Hill  limestone,  however  lenticular  it  may  be,  can  thin 
out  so  rapidly  as  all  this,  these  two  bosses,  or  boulders,  in 
the  railway  cutting  are  probably  mere  outlying  boulcleis  of  a 
southerly  extension  of  the  limestone  that  is  mainly  concealed 
beneath  the  surface.     (See  Fig.  2.) 

The  general  colour  of  the  Cave  Hill  limestone,  as  it 
appears  in  the  quarry,  when  looked  at  as  a  whole,  and  from 
a  distance,  is  cream,  or  almost  white,  especially  on  weathered 
exposures;  but  when  freshly  broken,  it  is  of  different  shades 
of  dark  or  light  bluish  grey,  ])inkish  biown,  or  grey  with 
pinkish  brown  patches.  It  is  for  the  most  part  semi-crystal- 
line, is  here  and  there  somewhat  brecciated  on  a  small  scale, 
and  is  in  some  ])laces  roughly  oolitic  (a  slide  of  an  oolitic 
specimen  on  view). 


40        Proceedings  of  the  Royal  Society  of  Victoria. 

Tlu'  limestone  strata  are  separated  at  intervals  by  five 
dark  shaly  or  mudstone  partings,  averaging  Irom  18  inches 
to  4  feet  across ;  the  tliickest  one,  which  is  of  a  dark  brown 
or  claret  coloui-,  is  npwards  of  4  feet,  and  in  this  one  is  an 
almost  perj^endicnlar  shaft-like  cave,  91  feet  deep,  apparently 
caused  b}^  the  action  of  running  water. 

I  am  informed  by  the  owner,  Mr.  David  Mitchell,  of 
Burnley,  that  the  limestone  of  his  quarry  has  been  long  ago 
analysed,  and  runs  to  95  or  96  per  cent,  of  calcium  carbonate. 
The  limestone  is  in  high  esteem  as  the  best  and  purest 
source  of  lime  for  building  purposes  in  the  colony.  It  has 
also  been  lately  turned  to  profitable  account  in  the  manu- 
facture of  cement,  and  the  owner  informs  me  that  the  force 
required  to  ])ull  it  apart  is  985  lbs.  The  stone  has  not  been 
directly  used  for  building  purposes,  but  some  slabs,  which, 
when  polished  up,  have  an  ornamental  appearance,  have  been 
occasionally  used  for  marble  mantelpieces.  Upon  the  whole, 
I  think,  therefore,  we  may  congratulate  Mr.  David  Mitchell 
upon  having  a  more  payable  thing  in  his  possession  than 
many  a  gold  mine.  "When  the  quarry  has  been  in  full  work 
in  prosperous  times,  he  tells  me  that  he  has  had  as  many  as 
120  men  employed  on  it,  and  has  been  able  to  send  away 
the  almost  incredible  amount  of  70  tons  of  lime  a  day.  In 
these  days  of  depression,  however,  when  there  is  so  little 
demand  for  lime,  20  men  are  found  quite  sufficient  to  do  all 
the  work  of  the  quarry  that  is  required. 

The  limestone  of  Cave  Hill,  like  most  other  limestones, 
contains  occasional  patches  of  crystalline  calcite,  mostly  in 
rhombohedrons  or  in  modified  scalenohedron.s.  It  also  con- 
tains segregated  lumps  and  layers  of  chert,  in  which  corals 
and  other  small  fossils  are  sometimes  beautifully  preserved 
(specimens  of  both  calcite  and  chert  are  exhibited).  Associ- 
ated with  this  same  limestone,  the  following  minerals  have 
been  ibund,  but  not  in  sufficient  quantity  to  be  of  any 
commercial  importance : — Galena,  malachite,  azurite,  and 
copper  and  iron  pyrites  (specimens  on  the  table).  The  great 
interest,  however,  of  the  Lilydale  limestone  lies  in  its  fossils 
— of  which,  indeed,  the  limestone  itself  is  largely  composed — 
and  which  constantl}' attract  scientific  visitors  from  Melbourne 
and  other  places,  and  indeed  from  the  neighbouring  colonies. 

In  lact,  one  of  my  chief  reasons  in  writing  this  paper  is  to 
take  possession,  in  the  name  of  Victorian  geologists,  of  the 
priority  in  describing  some  of  the  fossils,  as  at  present  so 
many  of  them  have  been  described  by  geologists  outside  the 


Note,s  on  the  LUijdale  Limestone.  41. 

colony;  for  without  a  moment  wishing  to  do  away  with  that 
principle  of  free-trade  in  scientific  research  that  we  all  so 
much  rejoice  in,  or  desiring  to  make  any  undue  claim  for 
protection  to  native  industry  in  Victorian  geology,  1  think 
you  will  agree  with  me  that  it  is  but  right  that  we  should 
try  to  take  inventories  of  our  own  possessions  for  ourselves, 
and  not  leave  it  to  outsiders  to  do  it  for  us. 

The  following  is  a  list  of  the  Lilydale  fossils  that  have 
been  so  well  described  by  Mr.  Robert  Etheridge,  jun.,  Govern- 
ment Paheontologist  of  New  South  Wales,  in  Nos.  8  and  7, 
Vol.  I,  of  the  "  Records  of  the  Australian  iMuseum,"  there:— 
Favosites  grandipora,  Trochus  (Scalietrochus)  lindstromi, 
Niso  (Vetotuba)  brazieri,  Cyclonema  australis,  Cyclonema 
lilydalensis,  Plianerotrema  australis..  Oriostoma  northi,  Mur- 
chisonia  attenuata  (i),  Bellerophon  cresswelli,  Ambonychia 
poststriata.  In  addition  to  these,  Mr.  Etheridge  records, 
without  describing,  the  well-known  and  world-wide  Silurian 
brachiopod,  " Atrypa  reticularis,"  and  mentions  also  that  there 
are  three  species  of  the  well-known  Rhizopod  "  Strom atopora" 
yet  to  be  described.  The  fossils  which  I  myself  wish  to 
record,  as  also  occuiring  in  the  Lilydale  limestone,  and  as  a 
supplementary  list  to  that  supplied  by  Mr.  R.  Etheridge, 
iun.,  are  the  following  : — 

MOLLUSCA   AND    MOLLUSCOIDEA. 

Cephalopoda. — Orthoceratites,  sp;  ;  a«id  Discoceras?     sp. 

Bellerophontldw. — Tremanotus  pritchardi. 

Gasteropoda. — Eunema  etheridgei,  Stomatia  antiqua, 
Tryblidium  (Metoptoma)  nycteis,  Pleurorhynchus  (Oono- 
cardium)  costatus,  and  Pleurorhynchus  (Conocardiura) 
bellulus,  Naticopsis  lilydalensis. 

La7)iellibranchiata. — Ambonychia  tatei. 

Brachiopoda. — Strophomena  rugosa,  Leptcena  transver- 
salis,  Orthis  elegantula. 

CCELENTERATA. 

Actinozoa. — Heliolites,  sp. ;  Cyathophyllum,  sp. 

Some  of  these  names  will  at  once  be  recognised  as  being 
those  of  world-wide  Upper  Silurian  forms,  but  the  following 
.species  are  new,  as    far  as  my  knowledge   goes,  and  so  I 


42        Proceedings  of  the  Royal  Societi/  of  Victoria. 

will  venture  to  name  and  descrilje  tlieni  as  such,  at  least 
provisionally  : — 

The  first  and  most  important  to  be  described  is  a 
shell  belongino^  to  the  Bellerophontidtie,  a  group  of  extinct 
shells  of  generalised  form,  which  had  charactei's  that  are 
now  divided  between  the  Cephaloi)()da.  the  Heteropoda, 
and  groups  of  Gasteropoda,  of  which  Pleurotomaria  and 
Haliotis  are  respectively  the  t\qjes.  It  is  a  Trernaiiotiis 
v/hich  I  have  named  T.  pritchardi,  in  compliment  to  Mr. 
G.  B.  Pritchard,  a  well-known  ge(Jogical  friend,  who  has 
kindly  lent  me  the  best  specimen  that  I  have  with  me,  and 
which  he  found  in  the  Lilydale  quariy  some  time  ago. 
8p.  Char,  of  T.  pritchardi  shell  discoidal,  bi-concave, 
trumpet-shaped,  and  very  thick,  consisting  of  about  live 
rapidly  increasing  whorls,  forming  a  deep  umbilicus  on  both 
sides  ;  spire  elliptical  in  section,  and  back  symmetricall}^ 
convex.  Breadth  of  the  shell  about  two  inches,  length  from 
three  and  a  half  to  four  inches.  Aperture  very  much 
expanded  and  reflected  like  the  mouth  of  a  trumpet,  but 
more  so  anteriorly  than  laterally  ;  the  inner  surface  of 
expanded  outer  lip  quite  smooth.  No  slit  or  sinus  as  in 
Belleropbon,but  the  middle  dorsal  line  of  the  shell  is  pierced  by 
a  row  of  oval  siphonal  openings,  resembling  those  of  Haliotis, 
there  are  about  seven  of  them  to  an  inch  of  the  ])eriphery. 
The  outer  surface  of  the  shell  is  ornamented  with  spiiul 
fluctuating  lines  parallel  to  the  dorsal  keel,  and  becoming 
on  the  ex[)anded  outer  lip  more  flattened,  coarser,  and  more 
plait-like.  There  are  also  the  very  distinct  lines  of  growth 
in  a  transverse  and  backward  direction  to  the  dorsal  keel, 
that  are  so  characterestic  of  the  Bellei'ophontida3.  The  lines 
in  the  two  directions  condjining  in  this  shell  to  give  a  very 
distinct  fenestrated  appearance.  T.  pritchardi  has  in  general 
form  a  near  i-esemblance  to  "Tremanotus  maideni,"  described 
by  Mr.  Robert  Etheridge,*  from  tiie  Hawkesbur}^  (Trias) 
rocks  of  New  South  Wales,  and  which  he  legards  as  a 
curious  survival  from  Silurian  times,  but,  besides  other 
differences,  our  fossil  is  a  very  mucii  thicker  shell. 

The  next  fossil  to  be  briefly  described  as  far  as  may  be 
from  very  imperfect  specimens,  is  Eunema  etheridgei,  a 
gasteropod  shell  that  a[)pears  to  belong  to  the  Littorinida?, 


*  Departraeut  of  Mines. — Memoirs  of  Geological  Survey  of  New  South 
Wales.  PaliEontolouy  I.  Invertebrate  Fauna  of  Hawke-bury  ;  Wiauamatta 
Series,  by  llobett  Etheridge,  jun. 


J^otes  on  the  Lily  dale  Limestone.  43 

like  the  Cycloneinas  of  this  Ibi'iuatiou,  wliieh  it  very  much 
I'esembles,  only  that  the  s])ire  is  more  elevated.  In  typical 
Eiuiemas,  according  to  Nicholson,  "  the  whorls  are  more  or 
less  angular,  and  tlie  surface  is  often  adorned  with  elevated 
spiral  ribs."  In  our  Eunenia,  however,  the  whorls  are 
lounded  like  those  of  Cyclonema,  and  ai-e  traversed  with 
spiral  keels,  but  more  numerous  and  less  distinct  than  in 
C  australis,  and  C.  lilydalensis.  There  is  also  an  indistinct 
appearance  of  a  spiral  band  about  the  middle  of  the  whorls. 
C.  etheridgei  is  like  "E.  cirrhosa"  of  the  English  Wenlook, 
as  figured  in  Murchison's  Siiuria,  but  has  much  more 
numerous  keels.  I  have  taken  the  liberty  of  naming  this 
shell  after  the  celebrated  Palseontologist  of  New  South 
Wales,  who  has  taken  so  warm  an  hiterest  in  our  Victorian 
fossils,  and  which  I  hope  may  be  still  continued,  notwith- 
standing the  fact  that  he  is  an  outsider  and  lives  across  the 
border. 

The  few  other  shells  which  I  take  to  be  new,  shall  be  passed 
over  with  but  very  slight  notice,  as  time  hastens,  and  there 
are  three  other  papers  to  follow.  One  is  a  gasteropod 
shell,  a  Stomatia,  which  I  have  called  "  Stomatia  antiqua," 
because,  as  far  as  I  am  aware,  it  is  the  oldest  Stomatia  upon 
record.  The  whorls  are  somewhat  steeper  in  the  sides,  and 
more  flattened  than  Stomatias  usually  are,  and  though  the 
spn-e  is  bioken  off  in  the  only  specimen  I  have,  it  must 
have  been  higher  than  is  usually  found  in  that  genus,  but 
in  all  other  I'espects  the  appearance  of  the  shell  is  that  of  a 
"  Stomatia."  The  whorls  are  diagonally  crossed  with  very 
numerous  lamella;-like  lines  of  growth.  The  shell  is  one 
and  a  lialf  inches  long,  and  one  inch  wide.  Then  thei'e  are 
two  small  species  of  "  Pleurorhyncltus,  or  Conocardium  " — 
lamelli-branchiate  shells  belonging  to  the  Cardiidie,  one 
about  half-an-inch  long,  with  nine  simple  ribs  on  the 
anterior  part  of  each  valve,  and  about  seventeeL  on  the 
hinder  part,  and  which  I  have  called  ''  Pleurorltynchus 
costatus."  And  the  other  species  is  about  one-third  of  an 
inch  long,  with  the  body  of  the  shell  more  oblique  to  the 
hinge  line,  more  prettily  banded  and  ribbed  than  the  other 
species,  the  ribs  being  crossed  with  striaj,  and  the  valves 
having  a  distinctly  fenestrated  appearance  at  the  posterior 
end.  This  I  have  accoi-dinglj^  named  "  FleurorhyncJtus 
helluhis." 

There  are  two  other  shells  that  I  have  not  figni-ed  on 
that   diagram,   but  have  here  to    show    j'ou.     I    will    pass 


44        Proceedings  of  the  Royal  Society  of   Victoria. 

them  over  with  bare  mention,  not  having  yet  had  time  to 
examine  and  compare  tliem  with  other  shells.  One  is  a 
Gasteropod,  a  Aatico2%sis  appai-ently,  which  I  will  call 
N.  lilydalensis,  if  it  should  tnvn  out  to  be  new  ;  and  the 
other  is  a  lamelli-branchiate  shell,  an  Ambonychia,  differing 
from  A.  post-striata  of  Etheridge,  and  alluded  to  by  Professor 
Tate,  as  having  a  fenestrated  ornament  on  the  sides  of  the 
valves.  If  Professor  Tate  has  not  already  named  it,  and 
will  forgive  my  impudence,  I  will  take  possession  of  it  in 
the  name  of  Victorian  geologists  and  call  it  A.  tatei,  for  I 
am  pretty  sure  we  had  found  it  long  befoie  he  did. 

In  concluding,  I  desire  to-^ acknowledge  my  indebtedness 
to  Mt.  D.  Mitchell,  the  owner  of  the  quarry,  and  also 
to  his  foreman,  Mr.  J.  Fuller,  for  statistical  and  other 
information  about  the  quarry  ;  to  Mr.  G.  B.  Pritchard,  of 
the  Woi-king  Men's  College,  for  the  loan  of  fossils  ;  and  to 
Mr.  H.  J.  Stokes,  organist  of  St.  John's,  Camberwell,  for 
the  photogi-aphs  of  the  quarry  that  have  been  exhibited  in 
illustration  of  this  paper. 


Pnr.  P.  s   VK'.anr.    Plate  Vlll. 


. ,tis>»3«rS»V<;V->    ^. 


"^  • 


/ 


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r  Umitl .  lith  C  Tratdd  i  Cpml. 


^f^-. 


Proc.  RS  Vidma.    Plate  IX. 


A 


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u 


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a.iyendel  lua.  CTrocdd  iC" print. 


Proc    R    5.    Victoria    Plate  JX' 


4"    .'^'^^ 


I       I 


Art.  y.—Preliminari/  Account  of  the  Glacial  Deposits  of 
Bacchus  Marsh. 

(With  Plates  X,  XI  and  XII.) 

By  Graham  Officer,  B.Sc,  and  Lewis  Balfour, 

Melbourne  University. 
[Read  July  14,  1892.] 

The  subject  of  glaciation  is  one  that  is  always  of  the 
greatest  scientitic  interest.  Its  important  bearings  on  the 
questions  of  climate,  past  and  present,  and  on  the  problems 
connected  with  the  evolution  and  distribution  of  plants  and 
animals,  render  it  a  field  where  the  astronomer,  geologist 
and  biologist  may  meet  on  common  ground.  The  subject 
has  received  its  fair  share  of  attention  in  the  Northern 
Hemisphere,  in  Europe,  America,  and  Asia,  but  in  the 
Southern  Hemisphere,  where  the  evidence  of  past  glaciation 
is  not  so  conspicuous,  comparatively  little  has  been  done  in 
this  direction.  Any  evidence  of  past  ice-action  in  Aus- 
tralia that  may  be  discovered  is  of  peculiar  value,  on  account 
of  its  bearing  on  the  question  of  the  probable  cause  of  ice- 
ages. 

The  earliest  reference  to  glacial  action  in  Victoria  is  made 
by  Selwyn  in  his  work  on  the  Geology  of  this  colony.  In 
this,  a  conglomerate  is  mentioned  as  occuriing  near  Bacchus 
Marsh,  and  which  contained  boulders  which  he  and  Mr. 
(afterwards  Sir)  R.  Daintree  considered  could  only  have  been 
brought  there  through  the  agency  of  floating  ice. 

Mr.  James  Stirling,  F.G.S.,  and  Di".  Lendenfeldt,  have 
described  evidences  of  former  extensive  glaciation  in  the 
Australian  Alps.  To  these  discoveries  reference  will  be 
made  later  on. 

Mr.  E.  J.  Dunn,  F.G.S.,  has  contributed  two  papers  on  the 
Glacial  Conglomerates  of  Victoria — one  read  before  the 
Royal  Society  of  Victoria  ;  the  other,  in  which  the  first  is 
incorporated,  before  the   Australasian   Association   for   the 


4G         Pi'oceediiKj.^  of  the  Royal  Society  of  Victoria. 

Advancement  of  Science  at  the  1890  meeting.  Tliis  conglo- 
merate is  described  as  occurring,  among  other  places,  at 
Bacchus  Marsh,  and  is  said  to  consist  of  material,  the  great 
bulk  of  which  is  derived  from  schistose  and  other  ancient 
rocks  and  to  contain  pebbles,  boulders  and  masses  of  from  20  to 
30  tons  M^eight.  Granites,  gneiss,  schist,  quartz-rock,  sand- 
stone, lydianice,  agate,  shale,  porpliyry,  and  jasper,  &c.,  are 
said  to  occur  in  it.  Many  of  the  included  stones  are 
striated,  and  often  flattened  on  one  or  more  sides  ;  others  are 
well  rounded,  and  others  again  are  rough  angular  fragments. 
Erratic  blocks  of  granite  occur  on  the  surface  at  Wild  Duck 
Creek,  near  Heathcote. 

Mr.  Dunn  is  of  the  opinion  that  "  no  other  conclusion  can 
be  arrived  at  than  that  floating  ice  has  been  the  agent  by 
which  the  material  has  been  brought  into  its  present  position." 
"Much  of  the  material,"  he  says,  "is  foreign,  and  many  of 
the  rocks  are  not  known  to  occur  at  present  in  this  Conti- 
nent anywhere  near  Victoria."  He  also  considers  the 
conglomerate  at  Wild  Duck  Creek  to  be  very  similar  to  the 
Dwyka  glacial  conglomerates  of  Soutli  Africa.  Mr.  Dunn's 
description  is  very  general,  and  the  evidence  on  which  his 
conclusions  are  based  is  somewhat  vague  and  wanting  in 
specitic  detail. 

Mr.  T.  S.  Hall,  M.A.,  has  also  given  a  short  account  of 
these  deposits  at  Heathcote  in  the  "  Victorian  Naturalist," 
(Vol.  VIII,  No.  2).  He  also  considers  the  beds  to  be  of 
iceberg  origin. 

Victoria  is  divided  into  two  main  areas  by  a  continuation 
of  the  Australian  Cordillera,  known  as  the  Main  Divide,  or 
Great  Dividing  Range.  This  extends  from  Forest  Hill  on 
the  east  to  the  Grampians  on  the  west.  In  the  eastern  part 
of  the  south  division  we  have  the  South  Gippsland  and 
Westernport  Ranges,  of  which  the  Southern  Spur  forms  a 
leading  feature.  West  of  Port  Phillip  we  have  the  isolated 
Otway  Ranges.  The  Main  Divide  reaches  its  highest 
elevation  in  its  eastern  portions.  Mount  Kosciusoo  being 
over  7000  feet  and  several  other  mountains  over  (iOOO  feet 
above  sea  level.  Passing  westwards  the  elevation  gradually 
diminishes.  The  average  elevation  is  about  8000  feet, 
but  in  some  places  it  sinks  to  1000  feet  above  sea  level. 
The  average  distance  from  the  coast  is  about  seventy  miles. 
There  are  three  main  drainage  systems — (1)  The  Murray 
System,  north  of  the  Divide ;  (2)  The  South  Eastern  or 
Gippsland  System,  south  of  the  Main  Divide  and  east  of  the 


Account  of  Glacial  DcpO'^iit^  of  Bacchii.^  AJarslt.      47 

southern  spur;  (3)  The  South  Western  System,  south  of  the 
Main  Divide  and  west  of  the  Southern  Spur.  The  Main 
Divide,  according  to  Murray,  is  a  "  longitudinal  area  of 
Lower  Palaeozoic  rocks,  with  granite  and  trappean  intrusions." 
These  may  be  overlaid  by,  or  flanked  by  IJ])per  Palaeozoic, 
Mesozoic,  and  Tertiary  deposits. 

The  Bacchus  Marsh  district  is  situated  about  midway 
between  Melbourne  and  Ballarat,  and  thus  belongs  to  the 
South  West  Drainage  System.  The  principal  streams  in 
the  locality  are  the  Werribee  and  its  tributaries,  the 
Myrniong  and  Korkui)ei'rimul  Creeks  and  the  Lerderderg 
River.  The  town  of  Bacchus  Marsh  is  picturesquely 
situated  in  a  broad  extensive  valley  343  feet  above  the 
sea,  and  which  has  pi'obably  formed  the  basin  of  an  ancient 
lake.  On  the  one  side  runs  the  Werribee,  and  on  the  other 
the  Lerderderg,  the  junction  of  the  two  streams  taking  place 
about  a  mile  below  the  town.  From  Bacchus  Marsh  the 
country  gradually  rises  to  the  Dividing  Range,  only  a  few 
miles  distant  to  the  north.  The  rising  ground  between 
Bacchus  Marsh  and  the  Dividing  Range  is  known  as  the 
Pentland  Hills. 

The  so-called  glacial  conglomerates  are  well  developed  in 
this  district  and  numerous  sections  exposed  to  view  by 
the  Creeks  and  various  cuttings  provide  very  favourable 
conditions  for  their  study. 

The  first  section  we  examined  is  situated  on  the  Ballarat 
Road,  about  three  miles  on  the  Ballarat  side  of  Bacchus 
Marsh,  and  is  at  a  height  of  about  750  feet  above  the  sea. 
The  deposit  exposed  consists  of  a  matrix  of  clay  of  a  quite 
unstratified  appearance,  and  of  a  somewhat  variable  con- 
sistency. It  is  tough  and  hard  in  places,  while  in  others 
it  is  softer  and  less  tenacious.  The  colour  is  an  indefinite 
patchwork  of  white,  yellow  and  purple.  Through  this 
matrix  are  scattered  irregularly  numerous  stones  of  various 
sizes  and  sorts,  rounded  and  sub-angular.  These  stones  do 
not  show  the  slightest  trace  of  arrangement  either  in  size  or 
in  position.  Some  of  the  stones  are  waterworn,  but  many 
present  quite  another  and  distinct  appearance.  These  often 
show  one  or  more  flattened  sides,  while  the  edges  and  ends 
are  frequently  rough  or  sub-angular.  Besides  these  peculiari- 
ties many  show  striated  surfaces,  the  striae  generally  running 
in  the  direction  of  the  longest  axis,  but  several  sets  of  striae 
can  often  be  distinguished.  Certain  kinds  of  stones  show 
strifb  much  better  than  others. 


48       Froceeduup  of  t/ie  Royal  yiocietij  of  Victoria. 

A  hard  fine  grained  argillaceous  sandstone  varying  in 
colour  from  a  very  light  to  a  darker  green  is  the  predomina- 
ting rock-material,  and  this  kind  usually  exhibits  the  most 
marked  striations.  Another  common  variety  is  a  hlue-black 
very  hard  quartzite.  These,  though  commonly  exhibiting 
flattened  or  facetted  sides  and  angles,  seldom  show  stria', 
their  surfaces  being  more  often  polished. 

Granite  often  occurs  though  not  so  frequentl}'  as  the  other 
kinds  of  rock.  The  largest  boulders  are  of  this  material. 
One  taken  from  the  cutting  can  be  seen  13'ing  at  the  side  of 
the  road,  which  is  well  flattened  on  one  side.  The  granite 
is  generally  considerably  decomposed.  At  the  top  of  the 
cutting  a  large  angular  fragment  of  sandstone  occurs,  while 
another  piece  can  be  seen  at  the  base  ;  whether  the  latter  is 
in  situ  or  not  we  have  not  yet  determined. 

The  unstratifled  nature  of  this  deposit,  together  with  the 
peculiar  nature  and  want  of  arrangement  of  the  included 
stones,  at  once  stamps  it  as  of  glacial  origin. 

A  few  feet  back  from  the  top  of  the  cutting  on  the  south 
side,  an  outcrop  of  white  silicious  sandstone  occurs.  We 
are  inclined  to  think  that  the  glacial  deposit  is  banked  uj) 
against  this,  really  overlying  it. 

A  short  distance  below  the  cutting  a  small  lateral  road 
joins  the  main  one,  and  on  this  road,  a  hundred  yards  or  so 
from  the  junction,  another  cutting  exposes  a  good  section. 
The  material  exposed  is  similar  to  that  just  described,  but 
is  of  a  more  uniform  purple  colour.  It  is  also  somewhat 
harder.  It  is  quite  unstratifled  and  contains  numerous  well 
striated  stones.  On  the  surface,  on  both  sides  of  this  cutting, 
glaciated  stones  aj-e  scattered  about  in  great  profusion  and 
variety.  This  deposit  apparently  overlies  sandstones  and  is 
continuous  with  that  exposed  on  the  main  road. 

Before  making  our  next  visit  to  the  locality,  we  wrote  to 
Mr.  Chai'les  Brittlebank,  of  Dunbar  farm,  near  Myrniong, 
who,  we  were  led  to  believe,  could  give  us  information  in 
our  researches.  Mr.  Brittlebank  readil}'  responded,  and 
during  our  subsequent  visits  has  rendered  us  much  valuable 
aid.  He  has  accompanied  us  on  most  of  our  expeditions  and 
shown  us  much  hospitality,  while  his  intimate  knowledge 
of  the  locality,  as  well  as  his  keen  powers  of  observation, 
have  been  of  the  greatest  assistance  to  us.  Mr.  Brittlebank 
informs  us  that  he  found  glaciated  stones  in  this  district 
four  years  ago.  He  thus  appears  to  have  been  the  first  to 
actually  prove  the  glacial  origin  of  the  deposits  in  question. 


Account  of  Glacial  J)eposits  of  Bacchus  Mavsli.      49 

The  valley  of  the  Myrniong  Creek  for  some  little  distance 
above  its  junction  with  the  Werribee  is  cut  through  basalt 
and  sandstones  and  conglomerates  to  a  depth  of  over  (iOO  feet. 
Good  sections  are  exposed  along  this  valley. 

On  the  south  side,  about  half  a  mile  above  the  confluence 
of  the  two  streams,  a  depth  of  over  100  feet  of  a  material 
similar  to  that  described  on  the  Ballarat  Koad  is  exposed. 
It  consists  of  a  mass  oi  yellowish  white  clay,  quite  unstrati- 
fied,  and  in  texture  somewhat  soft  on  the  weathered 
surface,  but  much  harder  on  being  penetrated.  Numerous 
stones  of  all  sorts  and  sizes,  from  mere  grit  to  boulders 
several  feet  in  diameter,  are  scattered  irregularly,  and 
without  any  trace  of  arrangement  thioughout  this  clay. 
Among  these  stones,  the  principal  varieties  are  those  . 
occuri'ing  in  the  cutting  on  the  road  already  described. 
Chiastolite  and  other  varieties  of  slate  were  found,  together 
with  quartz,  bits  of  jasper,  and  a  hard,  red  quartzitic  sand- 
stone. Most  of  these  stones  are  sub-angular,  often  showing 
one  or  more  smoothed  and  flattened  surfaces,  while  the  edges 
and  ends  are  roughly  angular  ;  many  are  will  striated  and 
grooved  in  a  characteristic  manner.  On  some  large  1)0ulders 
lying  at  the  base  of  the  clift',  the  strisB  and  grooves  are 
exceptionally  well  developed.  This  deposit  can  be  traced 
up  the  valley  for  about  a  quarter  of  a  mile  above  this  point, 
when  it  thins  out,  and  is  seen  to  overlie  and  flank  the 
sandstones  through  wliich  the  valley  has  been  worn.  It  is 
overlaid  by  basalt  known  as  the  newer  volcanic,  and  assigned 
to  Pliocene  age  (Fig.  1). 

On  the  other  side  (north)  of  the  Myrniong  Creek,  but 
nearer  its  junction  with  the  Weriibee,  the  glacial  deposit  is 
again  well  shown  to  a  depth  of  about  ]  50  feet.  It  is  much 
the  same  as  that  on  the  opposite  side  of  the  valley,  and 
striated  stones  are  numerous.  This  extends  to  within  "100 
yards  or  so  from  the  junction  of  the  two  streams.  It  can  be 
traced  over  the  brow  of  the  valley  up  to  about  the  level  of 
Mr.  Brittlebank's  house,  about  350  feet  above  the  Creek,  and 
about  1100  feet  above  the  sea.  It  then  spreads  out  over  the 
surface. 

It  would  seem  evident  then,  that  the  valley  now  occupied 
by  the  Myi-niong  Creek  at  this  point  at  any  rate  is  a  very 
ancient  one,  and  was  at  one  time  probably  almost  filled  up 
by  this  glacial  conglomerate.  The  sandstones  and  conglom- 
erates through  which  the  valley  is  worn,  were  set  down  as 
Upper  Palaeozoic  by  the  Geological  Survey  ;  then,  on  the  dis- 


50       Proceedings  of  the  Royal  Society  of  Victoria. 

covery  of  three  species  of  Gangamopteiis,  Professor  McCoy 
assigned  tliem  to  Triassic  times.  Last  year,  more  fossils 
were  obtained.  These  were  somewhat  fragmentary,  but 
Sir  Frederick  McCoy  thinks  he  can  identify  Schizoneura 
and  Zeugopliyllites,  indicating  a  lower  Triassic  age  for  the 
rocks  in  question. 

After  the  glacial  material  had  been  deposited  in  this 
ancient  valle}',  it  was  overflowed  by  basaltic  lavas  of 
Pliocene  age.  Whether  the  older  basalt  of  Miocene  times 
also  overflowed  this  valley  ])reviously  to  the  foi-mer,  we 
cannot  say  with  certainty.  We  have  seen  no  evidence 
of  it  at  any  rate.  Since  Pliocene  times  the  valley  has 
been  again  denuded  to  its  present  condition. 

From  the  general  characters  presented  by  the  so-called 
glacial  conglomerates,  we  were  much  inclined  to  the  opinion 
that  they  would  turn  out  to  be,  not  an  iceberg-drift,  but 
in  reality  till,  or  boulder-clay—  in  fact  the  ground  moraine 
of  ancient  glaciers.  These  characters  may  be  summed  up 
as  follows: — (I)  The  unstratifled  nature  of  the  clayey 
matrix.  (2)  The  number  and  variety  of  the  included  stones. 
(3)  The  striated  and  glaciated  aspect  of  many  of  these 
stones,  (4)  Their  total  want  of  arrangement.  In  fact, 
these  deposits  bear  such  a  striking  resemblance  in  every 
wa}'  to  the  till  of  Scotland  and  elsewhere  in  the  Northern 
Hemisphei-e,  that  it  can  haixlly  be  doubted  that  they  are  of 
similar  origin.  Corroboration  was  therefore  to  be  sought 
for  in  the  shape  of  roches  moutonnees,  or  shattered  rock 
surfaces  beneath  this  deposit.* 

In  the  valley  of  the  Myrniong  Creek,  opposite  the  section 
described  as  occuri'ing  on  the  south  side,  can  be  seen  rounded, 
hummocky-looking  masses  of  sandstone,  the  appearance  of 
which  is  very  suggestive  of  glacier  action.  It  is  very 
probable  that  the  glacial  conglomerate  not  long  since  covered 
these  rocks,  and  thus  protected  them  during  a  long  period 
from  the  effects  of  weathering.  It  must  also  be  ren)embered 
that  the  glacial  conglomerate  itself  must  have  been  piotec- 
ted  for  a  considerable  time  by  the  basalt.  The  sandstone  is 
hard  and  massive,  and  is  just  the  kind  of  rock  on  which  the 
abrading  and  rounding  effect  of  glacier  ice  would  be  well 
represented.     Certainly,  stride  and  grooves  are  absent,  but 


*  Having  had  opportunities  of  observing  the  till  and  other  phenomena  of 
glaciation  in  Scotland,  Ireland,  and  Switzerland,  I  can  vouch  for  the  striking 
resemblance  of  our  glacial  deposits  to  the  boulder-clay  of  the  Northern 
Hemisphere. — Graham  Officer. 


Account  of  Gldcial  Deposits  of  Biwchiis  Marsh.      51 

these  may  liave  weathered  away.  In  many  parts  of  the 
Scottish  Highlands,  where  the  whole  country  shows  the 
rounded  and  flowing  contour  chai-acteristic  of  ice-action,  it 
is  often  very  difficult  to  find  actual  scorings  and  grooves. 

Some  little  distance  further  up  the  Creek  a  section  has 
been  exposed  by  the  stream,  showing  some  feet  of  a  hard 
unstratitied  material  containing  striated  stones.  This  was 
much  hardei-  than  any  we  had  previousl}',  examined,  and  was 
traversed  by  joints.  It  was  seen  to  be  clearly  overlaid  by 
sandstones,  the  junction  between  the  two  being  very 
distinct,  there  being  apparently  an  unconformit3^  Here  a 
fault  occurs  through  tl^e  sandstones  and  the  underlying 
material,  the  displacement  being  about  seven  feet,  and  the 
hade  at  a  high  angle.  There  would  seem  to  be  no  doubt 
that  the  overlying  sandstones  are  continuous  Avith  the 
surrounding  ones,  which,  as  we  have  seen,  are  probably 
Triassic.  So  now  it  seemed  probable  that  we  had  to  deal 
with  two  glacial  deposits. 

At  the  junction  of  the  Myrniongand  Werribee,  the  latter 
stream  is  seen  to  be  flowing  over  the  highly  inclined  and 
sorely  denuded  edges  ot  Lower  Silurian  rocks,  here  consisting 
of  very  hard,  flne-grained,  well  stratified  sandstones.  On 
the  weathered  surface  the  colour  of  these  is  of  a  patchy 
yellow  rusty  colour,  but  on  the  fractured  fresh  smface  they 
are  of  a  light  greenish  white,  or  light  slaty  white  colour. 
On  proceeding  up  the  Werribee  from  the  junction,  we  found 
ourselves  walking  over  another  kind  of  inaterial,  which  was 
seen  to  rest  unconformably  on  the  Silurian  rocks,  which  it 
closely  resembles  in  colour.  The  Creek  has  cut  its  way 
through  this  to  the  Silurian,  so  that  on  the  floor  of  the  river 
course  one  walks  now  on  a  few  feet  of  Silurian,  and  now  on 
this  other  deposit,  while  sections  are  exposed  on  both  sides 
of  the  stream.  This  deposit  consists  of  an  exceedingly  hard 
clayey  material,  through  "which  are  scattered  stones  and 
boulders  of  considerable  size,  of  granite,  quartzite,  flne- 
grained  hard  sandstones  (very  similar  to  the  underl3dng 
Silurian),  and  quartz.  Nearly  all  these  present  the  flattened 
sides,  and  striated  and  grooved  surfaces  characteristic  of  ice 
action.  The  stones  and  boulders  at  this  point  are  very 
numerous,  and  the  scorings  and  scratchings  exceptionally 
well  developed.  This  conglomeiate  resembles  those  already 
described,  in  the  absence  of  any  appearance  of  stratification, 
the  character  of  the  included  stones,  and  the  total  want  of 
arrangement  of  the  latter.     In  fact,   it  cannot  be  distin- 

E  2 


52        Proceedings  of  the  Royal  Society  of  Victoria. 

giiislied  fiom  boiildei--clay  or  till.  However,  it  differed  from 
tliose  we  had  yet  examined,  except  the  last  described,  in 
being  so  excessively  hard  and  tough,  and  in  being  traversed 
by  numerous  joints.  Till  one  has  actually  tried,  it  is 
inijiossible  to  give  an  idea  of  the  difficulty  of  extracting  a 
stone  from  this  material,  which  will  ordy  come  awaj'  in  small 
angular  fragments,  in  a  manner  that  is  peculia)-!}' exaspera- 
ting. On  the  north  side  of  the  Creek,  a  short  distance  from 
the  junction,  a  section  of  a  similar  deposit  is  exposed,  which 
pi'esents  a  somewhat  stratified  a])pearance  ;  striated  stoiies 
occur  irregularly  through  this,  but  they  ai-e  not  so  numerous 
as  on  the  opposite  side  of  the  Creek.  The  appearance  of 
stratification  presented  may  possibly  be  due  to  pressuie.  It 
is  overlaid  by  basalt. 

On  proceeding  up  the  Weiribee  a  few  yards  furtlser  on  the 
south  side  we  found  a  small  cliff,  where  the  junction  of  the 
conglomerate  with  the  underlying  Silurian  could  be  well 
seen  in  section.  Here  was  a  ])lace  wliere,  if  the  conglomerate 
were  a  true  till,  we  might  expect  to  find  the  underlying- 
rock  smoothed  and  striated,  or  else  shattere<l.  The  section 
exposed  showed  the  Silurian  rocks  rising  in  a  hummocky 
way,  and  closely  overlaid  by  the  conglomerate.  A  aloser 
irispection  revealed  a  certain  rounded  and  facetted  appear- 
ance, that  was  very  suggestive  of  ice  action.  Ha\ing  found 
a  place  where  the  overlying  deposit  was  thinner  than  usual, 
we  resolved  to  clear  aw^ay  a  portion,  and  after  some  difficulty 
and  hard  work  succeeded  in  laying  bare  a  portion  of  the 
rock  below.  We  were  amply  le warded  for  our  trouble. 
The  Silurian  rock  presented  in  a  beautiful  manner  a  well 
sm(.)othed  and  striated  surface,  with  deeper  parallel  grooves, 
all  running  in  a  north  and  south  direction,  and  of  the  glacier 
origin  of  which  there  could  be  no  doubt  whatever.  The 
Silurian  strata  here  dip  west,  at  angles  of  from  50°  to  (iO°. 
So  it  will  be  seen  that  the  sti'ata  are  cut  across  at  right 
angles  to  the  dip,  in  fact  in  the  direction  of  the  strike.  It 
is  quite  impossible  that  this  can  be  due  to  the  action  of  the 
Creek,  or  indeed  to  the  action  of  water  at  all.  The  strife 
and  grooves  point  right  across  the  Creek.  Tlie  contiguous 
portions  of  the  overlying  deposit,  when  removed,  were 
found  to  retain  perfect  mouldings  of  the  grooves  and  strife 
beneath. 

This  striated  and  grooved  rock  surface,  taken  in  connection 
with  the  nature  of  the  overlying  deposit,  leaves  no  room  for 
doubt  as  to  the  glacier  origin  of  the  latter,  and  that  it  is  a 


AccoLLiit  of  Glacial  Deposits  of  Bacchus  Marsh.      53 

true  till,  or  moraine  profoiide.  This  till  can  be  traced  down 
the  Werribee  to  its  junction  with  the  Myrniong  Creek,  and 
a  little  way  beyond  on  the  latter  Creek.  It  liere  is  appa- 
rently overlaid  by  the  Tricvssic  rocks.  With  heavier  tools 
than  we  had  at  our  disposal,  and  a  little  more  time,  it  would 
not  be  diiiicult  to  remove  more  of  the  till  from  the  under- 
lying Silurian,  and  thus  lay  bare  more  of  the  moutonnee 
surface. 

A  few  days  after  this  discovery,  we  received  a  letter  from 
Mr.  Brittlebank,  stating  tliat  he  had  found  a  further  example 
of  roche  moutonnee  at  the  lower  end  of  the  Werribee  Gorge, 
nearly  two  miles  below  its  junction  with  the  Myrniong. 
On  our  next  visit,  we  accordingly  proceeded  to  the  spot,  and 
examined  the  rocks  in  question. 

The  Gorge  has  been  cut  to  a  depth  of  over  GOO  ft.  through 
a  mass  of  Silurian  rocks,  flanked  by  the  Triassic  sandstones 
and  conglomerates,  the  former  having  formed  a  ridge  or 
island  in  the  Triassic  sea  or  lake  (Fig.  1).  The  Silurian 
rocks  heie  consist  of  slates, finely  laminated  shales,  and  hard 
quartzitic  sandstones  ;  quartz  veins  are  frequent,  and  a  dyke 
of  porphyry  also  occurs.  The  strata  are  inclined  at  the 
usual  high  angles,  being  often  almost  vertical. 

At  the  place  indicated  b}^  Mr.  Brittlebank  we  found  the 
till  again  overlying  the  Silurian.  Here,  it  presents  much  the 
same  ajipearance  as  that  last  described,  glaciated  stones  and 
pebbles  being  frequent.  At  this  point,  at  a  spot  where  the 
till  was  only  about  a  foot  thick,  Mr.  Brittlebank  had  laid 
bare  a  portion  of  the  underlying  rock.  An  exoniple  of 
roche  moutonnee  was  thus  exposed  to  view,  which  was  even 
better  than  the  one  h'lst  discovered.  More  of  the  overlj'ing 
deposit  was  now  removed,  and  a  greater  surface  of  the 
nnderl3'ing  rock  uncovered,  this  being  an  operation  of  some 
difficulty.  The  surface  exposed  ])resented  the  apjjearance  of 
three  smooth  parallel  ridges,  well  scored  and  striated,  with 
well  rounded  grooves  six  or  more  inches  deep  between. 
Here,  as  before,  the  stride  and  grooves  run  noi-th  and  south, 
in  the  direction  of  the  strike,  and  right  across  the  river 
(PI.  XI).  In  several  places,  the  rock  has  been  fractured  at 
right  angles  to  the  groovings.  Photographs  of  these  were 
obtained.  This  was  by  no  means  the  only  spot  in  this 
locality  where  roches  moutonnees  were  found.  A  short 
distance  further  up  the  Creek  can  be  seen  a  niunded 
huunnock  of  Silurian  rock,  which  has  been  denuded  of  the 
overlying  till.     The  effects  of  weathering  have   obliterated 


54        Proceedings  o/  the  Royal  Society  of  Victoria. 

all  stiia^  and  grooves,  bvit  the  rounded  contour  still  remains. 
In  several  other  places  small  portions  of  the  till  weie 
removed,  and  a  striated  and  grooved  surj'ace  invariably 
exposed,  the  direction  of  the  strife  being  still  constant. 
The  till  here  is  about  ten  or  twelve  feet  in  thickness,  and  is 
distinctly  overlaid  by  the  Triassic  rocks.  On  the  opposite 
side  of  the  river  (south  side),  a  good  section  is  ex])osed. 
The  till  is  again  seen  resting  on  the  Silurian  rocks,  which 
here  also,  as  seen  in  section,  appear  to  have  been  subjected 
to  the  action  of  ice  (PI.  XII).  The  strata  are  nearly  vertical. 
The  till  here  is  seen  to  thin  out,  forming  a  wedge-shaped 
mass.  It  is  overlaid  by  the  Triassic  rocks  which,  below  the 
lower  end  of  the  wedge,  rest  directly  on  the  Silurian.  The 
till  and  overlj'ing  formation  extend  a  short  distance  up  the 
Creek  from  this  point,  when  they  terminate  against  the 
U})risiiig  ridge  of  Silurian  strata. 

There  would  seem  to  be  little  doubt  that  the  Triassic 
rocks  overl}"  the  till  unconformably.  It  will  now  be  seen 
that  there  are  two  distinct  glacial  deposits.  Of  these,  one 
is  overlaid  by  the  Triassic  sandstones  and  conglomerates, 
and  is  undoubtedly  an  ancient  till,  or  moraine  profonde  ; 
the  other  overlies  the  Triassic  rocks  and  is  similar  to  the 
lower  till,  except  that  it  is  not  so  hard  nor  so  traversed  by 
joints,  which  is  hardly  a  matter  for  surprise. 

Numerous  well  striated  stones  and  boulders  are  scattered 
ovei'  a  great  part  of  the  suifoce  between  the  Ballarat 
Road  and  the  Myrniong  and  Werribee  streams,  up 
to  an  elevation  of  over  11 00  feet  above  the  sea.  These 
stones  can  be  traced  flanking  the  ridges  that  overlook  the 
Wenibee.  At  a  point  opposite  the  Gorge,  at  the  lower  end, 
the  stones  are  especially  numerous  and  very  well  sti'iated. 
In  addition  to  the  commoner  varieties,  a  hard  semi- 
crystalline  sandstone,  of  a  dark  pink  colour,  occurs. 
The  stones  here  overly  the  Triassic  .sandstones,  and  can  be 
traced  along  a  small  lateral  gully  right  down  to  the 
Werribee.  The  de[iosit  from  which  they  come  is  exposed  at 
various  points  along  this  gully,  and  is  quite  similai'  in  its 
unstratitied  nature,  and  in  the  irregular  arrangement  of  the 
included  stones  to  that  described  before.  In  places  it  presents 
a  ver}-  hard  texture,  sometimes  somewhat  resembling  the  till 
below  the  Triassic  rocks,  in  other  places  it  is  softer,  but  in 
seveial  places  where  its  junction  with  the  underlying  sand- 
stones could  be  seen,  it  was  so  invariably  hard  and  thick 
that  we  could  not  clear  any  away  so  as  to  expose  the  under- 


Account  of  Glacial  Deposits  of  Bacckus  Marsh.     55 

lying  rock.  However  in  places,  as  seen  in  section,  the  latter 
presented  a  roundel  appearance  that  was  very  suggestive  of 
ice  action. 

At  the  intake  of  the  Bacchus  Marsh  water  supply  on  the 
VVerribee,  about  ;»  mile  below  the  Gorge,  where  the  valley  is 
very  broad,  a  splendid  section  of  a  till-like  deposit  is  exposed  ; 
there  being  over  70  feet.  The  matrix  is  a  yellowish-white 
clay,  very  tough  and  hard,  and  stones  and  boulders  of  the 
usual  kind  are  scattered  through  it  in  a  pell-mell  fashion, 
with  no  trace  of  arrangement.  There  is  no  stratification, 
but  irregular  bands  occur  here  and  there,  sometimes 
lenticular  in  form.  These  bands  are  in  some  cases  of  a  line 
sandy  material ;  others  consist  of  minute  angular  fragments 
of  much  the  same  nature  as  the  rest  of  the  deposit,  but 
coarser.  These  bands  are  only  about  eighteen  inches  or  two 
feet  in  thickness,  and  seem  to  have  been  formed  by  the 
intermittent  action  of  running  water.  Similar  bands  and 
lenticular  patches  of  sand  and  other  material  occur 
fre([uently  in  the  till  of  the  Northern  Hemisphere,  having 
been  formed  by  the  action  of  sub-glacial  streams.  We  have 
not  yet  been  able  to  determine  definitely  the  relations  of 
this  deposit,  but  froan  its  nature  and  position,  as  well  as  its 
great  thickness,  we  incline  to  the  opinion  that  it  belongs  to 
the  upper  glacial  deposit.  It  occurs  again  about  half  a  mile 
fuither  down  the  river,  where  good  sections  of  it  are  exposed. 
It  here  does  not  contain  nearly  so  many  stones,  wdiile  those 
that  do  occur  a'e  generally  small,  otherwise  it  is  similar  to 
that  last  described. '  We  have  not  found  the  deposit  between 
this  point  and  Bacchus  Maish  along  the  Werribee. 

About  four  miles  up  the  Korkupeirimul  from  the  bridge 
on  the  Ballarat  Road,  a  glacial  conglomerate  is  again  met 
with  containing  numerous  typical  glaciated  stones.  The 
matrix  is  exceedingly  hard  and  devoid  of  stratification.  In 
places,  when  looked  at  from  one  point  of  view,  an  appeai-ance 
of  a  somewhat  irregular  stratification  can  be  seen.  However, 
a  more  careful  examination  reveals  the  fact  that  what  ai'e 
apparently  lines  of  stratification,  are  in  reality  curved 
divnsion-planes,  which  are  probably  due  to  shearing  stresses. 
At  one  place  in  this  section  a  departure  from  the  usual 
irregular  disposition  of  the  stones  may  be  observed.  The 
stones  ai-e  arranged  in  a  sloping  fashion,  along  an  irclined 
plane.  This  arrangement  is  sometimes  met  with  in  the  till 
of  the  Northern  Hemisphere.  At  this  place  also  a  bouldei-, 
about  eighteen  inches  long  and  somewhat  pear-shaped,  can 


;)(j        Proceedings  of  the  Royal  Society  of  Victoria. 

he  seen  resting-  in  the  matrix  in  a  vei'tical  position.  Now, 
if  such  a  boulder  were  dropped  from  an  iceberg,  we  might 
expect  it  to  remain  in  an  upriglit  position  in  the  soft  clay, 
but  if  so,  we  should  certainly  expect  t(^  find  the  clay  indented 
beneath  it.     Of  this,  there  is  not  the  sliglitest  indication. 

A  little  fnrthei-  up  the  Creek  another  section  is  exposed. 
Here  our  till-like  deposit  )-ests  on  massive  sandstone,  but  we 
were  unable  to  remove  sufficient  of  the  former  in  order  to 
expose  the  surface  beneath.  At  one  point,  however,  a 
somewhat  remarkable  feature  occurs.  In  the  sandstone  is 
an  oblique  gap  about  four  oi-  five  feet  deep,  as  if  a  block  had 
been  torn  out.  This  cavity  is  filled  with  the  overlying 
material,  and  two  or  three  flattened  and  striated  stones  rest 
on  its  lo\ver  side  (not  bottom).  It  is  difficult  to  conceive 
how  icebergs  could  have  deposited  stones  in  this  manner, 
while  on  the  other  hand  it  is  readily  explained  on  the  glacier 
theor^^ 

The  locality  between  this  ])lace  and  the  large  quai-ry, 
about  two  miles  further  down  the  Creek,  we  have  not  yet 
examined.  Between  this  quarry,  situated  on  the  north  side 
of  the  Koi'kuperrimul,  and  the  bridge  on  the  Ballarat  Road, 
the  valley  in  which  the  Creek  flows  follows  approximately 
the  axis  of  what  has  once  been  an  anticlinal  fold  of  the 
Triassic  sandstones.  Opposite  the  lai-ge  quarry,  the  valley 
is  a  little  to  the  light  of  this  axis.  Between  this  large 
quarry  and  the  Creek,  striated  stones  are  numerous.  A 
small  lateral  gully  exposes  sections.  One  of  these  shows  a 
somewhat  loamy  clay,  in  which  ar-e  irregularly  imbedded 
large  angular  fragments  of  sandstone,  in  ap[)earance  very 
like  the  underl3'ing  rock.  Large  granite  boulders,  quartzite, 
slate,  quartz,  and  fragments  of  jasper  also  occur,  many 
showing  flattened  and  striated  surfaces. 

On  the  Creek  opposite  the  quarry,  a  clifl"  of  about  (iO  feet 
of  the  glacial  dep  )sit  is  exposed.  It  is  very  similar  to  that 
desci-ibed  on  the  Ballarat  Road.  It  rests  on  sandstones,  the 
broken  ends  of  vv^hich  can  be  seen  protruding  from  the  base 
of  the  deposit,  which  towai-ds  the  top,  presents  a  somewhat 
stratified  appearance.  On  the  opposite  side  of  the  Creek, 
high  clifls  of  basalt  (newer)  occur.  This  has  evidently  filled 
up  the  valley  at  this  place,  probably  covering  the  glacial 
deposit  and  having  since  been  denuded  away  to  its  ])resent 
state. 

Several  hundred  yards  further  down  tlie  Creek,  on  the 
right  hand  side,  a   section  exposed  shows  a  few  feet  of  an 


A  ceo  a  lit  of  Glacial  Deposits  of  Bacchus  Marslt.     57 

unstratitied  Diaterial  bearing  striated  stones,  and  overlaid  by 
very  irregularly  stratified  tumultuous-looking  sandstones. 
These  sandstones  are  very  probably  simply  beds  associated 
with  the  glacial  deposit.  This  is  indicated  by  their 
tumultuous  ajipearance,  and  by  the  fact  that  we  found 
several  well  scored  stones  in  them.  Moreover,  a  small  patch 
of  a  material  sin.ilar  to  that  beneath  occurs  intercalated 
\vith  them.  'J'he  basalt  is  banked  right  u[)  against  this,  the 
line  of  junction  being  almost  \ertical.  The  whole  mass 
probably  formed  a  ridge  in  the  valley  at  the  time  the  basalt 
overflowed  it.  Striated  stones  can  be  traced  foi'  about  a 
third  of  a  mile  further  down  this  valle}',  on  the  right  hand 
side,  being  overlaid  by  basalt  (Fig.  2)  The  characteristic 
stones  of  the  glacial  deposit  can  be  traced  along  the  hills 
flanking  the  valley  on  the  left.  At  one  spot,  between  the 
big  quairy  and  another  smaller  one  further  down  the  valley, 
a  conglomerate  occurs,  which  consists  of  a  loam}-  matrix,  in 
which  are  scattered  angular  fragments,  in  all  |)i)sitions,  of 
soft  sandstone.  This  rests  on  the  denuded  edges  of  well 
stratifled  Triassic  saridstone,  from  which  the  fragments  have 
apparently  been  derived. 

Some  distance  further  on,  a  small  quarry  occurs  in  the 
Triassic  sandstones,  whicli  here  dip  E.S.E.  about  r35°.  The 
glacial  conglomerate  can  be  traced  to  about  200  feet  above 
the  Ci-eek.  and  in  the  quari-y  can  be  seen  in  section  lesting  on 
the  sandstf)nes  to  a  depth  of  about  Ave  feet.  On  the  left 
hand  side  of  this  section,  the  junction  is  very  marked,  wh.ile 
ti-acing  it  to  the  right,  it  becomes  very  indeflnite  by  the 
disintegration  of  the  sandstone.  This  section  is  at  right 
angles  to  the  dip.  At  the  same  quarr}^  another  section  is 
exposed  at  right  angles  to  the  former.  This  exhibits 
remarkable  and  important  features.  Beginning  at  the  lower 
en;l  of  the  section,  a  pell-mell  accumulation  of  rough  angular 
and  rounded  blocks,  up  to  eighteen  inches  and  two  feet 
in  diameter,  embedded  in  a  loamy  matrix,  is  seen  overlying 
soft  purplish  stratifled  clays  or  shales.  The  lattei  are  much 
broken  up  and  disintegrated  at  their  junction  with  the  over- 
lying deposit.  Anoular  blocks  of  sandstone  in  every 
conceivable  position  are  mixed  up  in  the  ruin,  and  in  fact 
a  deflnite  junction  it  is  almost  impossible  to  distinguish. 
Further  aloiig  the  section,  this  mixed  matei-ial  merges  into  a 
purplish  mass  of  clay,  ovei'lying  broken  and  shattered  sand- 
stones. (The  shales  and  sandstones  are  of  the  same 
formation.)      This  purplish  clay,  which  is  evidently  derived 


58        Proceedings  of  the  Royal  Society  of  Victoria. 

from  the  shales,  presents  the  appearance  of  liaving  been 
pushed  over  the  sandstones,  anguhir  blocks  of  which  are 
scattered  through  it.  A  little  further  along,  a  large  irregular 
fracture  in  the  sandstone  occurs,  being  seven  or  eight  feet 
deep.  This  is  litei-ally  stuffed  with  stones  and  boulders 
of  the  various  kinds  met  with  in  the  glacial  conglomerate. 
Many  of  these  show  flattened  and  striated  surfaces.  A 
granite  boulder,  over  a  yard  in  dian)etei-,  is  jammed  into  the 
bottom  of  this  fracture,  wjiile.  broken  and  angular  fragments 
of  the  sandstone  are  also  scattered  through  it,  the  whole 
being  imbedded  in  a  loamy  clay-like  material,  which  seems  to 
have  been  squeezed  into  the  fracture  (Fig.  S).  At  several 
other  sections  exposed  in  this  quarry,  similar  appearances  can 
be  noted.  The  sandstone  has  been  fractured,  and  the  glacial 
materia]  literally  injected  into  the  ci'acks  and  fissures. 
Several  striated  stones  were  picked  out  from  one  of  these 
fissures. 

It  will  be  seen  that,  as  in  the  case  of  the  Myrniong  Creek, 
the  glacial  deposit  lies  in  an  ancient  valley  of  denudation. 
It  was  probably  overflowed  by  Pliocene  basalt,  which  would 
thus  be  the  means  of  protecting  the  underlying  formations 
during  a  considerable  period.  We  could  not  find  any  more 
traces  of  the  glacial  material  between  this  place  and  the 
Werribee. 

This  concludes  the  evidence  we  have  so  far  collected,  and 
it  all  points  irresistibly  to  the  conclusion,  that  glacier-ice 
has  been  the  agent  b}^  which  the  effects  described  have  been 
accomplished.  No  iceberg  theory  will  account  for  the  facts 
presented  at  the  quairy.  How  will  such  a  theory  account  for 
the  fracLin-ing  of  the  underlying  rocks,  and  the  ramming  of  the 
fractures  with  laige  erratic  boiddersand  the  material  in  which 
these  boulders  ai-e  imbedded  ?  On  the' other  hand,  these  are 
facts  which  are  readily  explained  on  the  glacier  hypothesis. 
In  the  Northern  Hemisphere  shattered  surfaces  are  frequently 
met  with  below  till.  In  his  "Great  Ice  Age,"  p.  1(5,  Prof 
James  Geikie  says  : — "  Soft  sandstones  and  highly  jointed 
rocks  .  .  .  often  show  a  broken  and  shattered  surface 
below  till  ;  sometimes,  indeed,  thick  sandstones  appear 
'broken  up'  to  a  depth  of  many  feet  below  boulder-clay, 
the  ccmi-se  angular  debris  shading  gi-adually  into  till  of  the 
normal  type."  This  corresponds  exactly  with  the  features 
presented  at  the  quarry,  where  the  sandstones  are  soft  and 
easily  disiiitegrated.  Cases  in  Scotland  and  elsewhere  in 
the    Northern    Hemisphere  are  not  uncommon,  where  the 


Account  of  Glacial  Deposits  of  Bacchus  Marsh.     59 

shattered  surfjice  of  the  underlying  rock  is  "  stuffed  "  with 
erratic  stones  and  boulders. 

The  conckision,  then,  to  which  we  are  led  is,  that  the 
deposits  we  have  been  considering  constitute  a  true  till,  or 
moraine  protbnde.  This  is  borne  out  by  further  considei'a- 
tions.  It  is  worthy  of  note,  that  the  stones  occurring  in 
tliis  till,  at  the  quarr}'  we  have  heen  speaking  about,  are  not 
nearly  so  well  striated  as  those  occurring  in  the  region  of 
the  Werribee  Gorge.  In  the  former  case,  we  have  seen  that 
the  underlying  sandstone  is  very  soft,  and  would  not  striate 
stones  well ;  on  the  other  hand,  the  underlying  rocks  in  the 
i-egion  of  the  Gorge  are  much  harder,  consisting  to  a  great 
extent  of  conglomei'ates,  just  the  kind  of  rocks  that  would 
produce  marked  scorings  on  the  stones  of  the  till. 

These  are  specific  evidence  against  the  iceberg  theory. 
There  are  also  more  general  arguments.  These  arguments 
have  been  used  l>efore  to  refute  the  iceberg  hypothesis  of  the 
origin  of  the  boulder-clay  in  the  Northern  Hemisphere,  and 
they  ap|)ly  equally  well  hei-e. 

Mr.   Dunn    describes    the    so-called    glacial   conglomerate, 

besides  being  found  at  Bacchus  Marsh,  as  occurring  on  both 

sides  of   the   Dividing  Range,  at  Wahgunj^ah,    Rutherglen, 

The   Springs,   El   Dorado,   Wooragee,    Tarrawingee,    Badda- 

ginnie,  at  various  points  on  the  road  between  Wangaratta  and 

Kilmore,  north   east  of  Costeriield,  Wild  Duck  Creek  (west 

of  Heathcote),  underlying  the  auriferous  deposits  at  Caris- 

l))-ook  and  Oreswick.     South  of  the  Dividing  Range,   it  is 

met  with  about  four  miles  east  of  Gordons,  Barrabool   Hills, 

and  near  Foster  in  South  Gi]ipsland.     Thus  it  will  be  seen 

that  the  deposit  is  widely  distributed,  and  it  appears  to  be 

of  ccjnsiderable  thickness,  beino-  over  100  ft.  in  several 
1  •  'ft 

known  instances. 

It  has  been  shown  (CroU,  "  Climate  and  Time ;" 
Geikie,  "  Great  Ice  Age,"  etc.),  that  the  amoui.t  of  material 
carried  by  icebergs  is  quite  inconsiderable,  and  what  is 
carried  genei-ally  consists  of  rubbish  and  angular  blocks 
that  have  fallen  on  the  surface  of  the  parent  glacier. 
In  the  case  of  the  ice-sheet  that  is  at  present  desolating 
Greenland,  the  surface  of  the  ice  is  ver}'  free  from  debiis  of 
any  kind,  and  so  it  is  quite  a  rare  thing  to  find  an  iceberg 
shed  from  one  of  the  vast  glaciers  of  that  countiy  bearing 
any  material  at  all.  Yet  a  tremendous  amount  of  erosion 
must  be  going  on,  and  the  eroded  material  is  being  accumu- 
lated   beneath    the    ice    as   a    moraine    profonde,    although 


00       Pfoceedinf/s  of  the  Royal  Society  of  Victoria. 

])rodigious  quantities  must  be  carried  away  by  sub-glacial 
streams.  Dr.  Wright  ("  Ice  Age  in  North  America")  calcu- 
lates that  from  the  great  Muir  glacier  in  Alaska  ovei-  33^ 
million  cubic  yards  of  sediment  is  annually  cairied  away  by 
sub-glacial  streams.  Little,  if  any,  of  the  sub-glacial  material 
can  be  carried  away  by  icebergs — a  few  stones,  })erh;ips,  frozen 
into  the  bottom  of  the  bergs.  The  finer  material  carried 
away  b}^  streams  from  beneath  these  great  glaciers  must 
inevitabh-  be  stratitied,  and  well  stratified,  as  the  quantity 
of  material  brought  down  must  vary  considerably  from 
time  to  time.  Even  if  much  line  matter  weie  carried  by 
icebergs,  it  would  inevitabl}^  be  re-assorted  by  the  water  ; 
the  stones,  too,  would  assuredly  show  some  trace  of 
arrangement. 

In  the  deposits  we  have  been  considering,  the  absence 
of  stratification  and  the  total  want  of  arrangement  of  the 
included  stones,  are  their  chief  and  most  striking  cliaiacter- 
istics.  Then  again,  in  the  great  mass  of  the  sections  we 
have  examined  angular  fragments  are  comparatively  rare, 
except  as  we  have  seen,  where  the  till  rests  on  the  under- 
Ivinsf  rock.  So  here  aoain,  we  have  a  strono-  armament  in 
favour  (jf  the  glacier  theory. 

Further,  these  deposits  are  found  up  to  a  height  of  1400  ft. 
at  Bailan  ;  so,  to  account  for  them  en  the  iceberg  theory,  we 
would  require  a  submergence  of  at  least  2000  ft.  to  allow 
icebergs  to  float,  and  as  icebergs  can  only  transport  material 
from  higher  to  lower  levels,  it  is  quite  impossible  to  account 
for  the  mingling  of  fragments  of  the  underlying  rock  in  the 
overlying  till,  at  an  elevation  not  exceeding  800  ft.  above 
the  sea.  Besides,  such  a  submergence  would  considerably 
diminisli  the  area  frcnn  which  the  deposits  could  be  derived, 
and  their  extent  indicates  a  large  surface.  Again,  such  a 
submergence  would  tend  to  produce  climatic  conditions 
which  would  be  quite  o[)posed  to  the  production  of  glaciers, 
even  were  the  astronomical  conditions  favoural)le.  It  must 
also  be  observed  that,  so  far  as  we  have  seen,  these  deposits 
are  quite  unfossiliferous. 

Mr.  Dunn  states  that  much  of  the  rock  material  occurring 
in  the  till  is  not  known  at  ]n-esent  to  occur  in  situ  on  this 
Continent  anywhere  near  Victoria.  Daintree  remarks  that 
a  granite  occurs  in  the  formation  at  Bacchus  Marsh,  which 
\\t>  had  not  observed  south  of  Queensland.  However,  as  he 
has  not  described  this  granite,  it  would  be  difficult  now  ti> 
identify  it.      We  would  reply  to  this  that  further  search  will 


Account  of  Glacial  Deposits  of  Bacchus  Marsh.     Gl 

probably  reveal  the  sources  of  this  material.  The  .i!;eoloi;y 
of  Victoria  has  not  been  so  lully  worked  out  as  to  warrant 
us  asserting  that  a  certain  kind  oi'  rock  does  not  occur  in 
situ.  Then  again,  it  must  be  remembered  that  these 
deposits  are  anterior — as  we  shall  sliow — to  the  Miocene 
and  Pliocene  lava  flows,  and  probably  to  the  Miocene  leaf- 
beds,  so  that,  not  to  speak  of  the  effects  of  denudation,  a 
a  great  deal  of  the  then  rock  surface  is  now  concealed. 

Of  the  various  kinds  of  rock  met  with  in  the  till  in  the 
Bacchus  Marsh  district,  the  gieat  majority  are  derived  from 
Silurian  rocks,  which  form  the  main  part  of  the  Dividing- 
Range.  In  the  Werribee  Gorge  several  kinds  of  slate  occur, 
which  are  identical  with  slates  found  in  the  till.  Quartz 
veins  are  als(^  numerous  in  the  Silurian  rocks.  We  also 
noted  a  quartzitic  sandstone  in  the  Gorge,  which  is  very 
similar  to  fragments  found  in  the  till.  Several  varieties  of 
quartzite  occur  in  the  till  which  we  have  not  yet  seen  in 
situ,  but  we  have  not  yet  examined  the  Ranges  to  the  north, 
and  it  is  very  probable  they  will  be  found  there,  as  quartzites 
frequently  occur  in  the  Silurian.  Fragments  of  schist(;se 
rocks  have  also  been  observed  in  the  till,  and  these  occur 
in  situ  to  the  north. 

Several  kinds  of  granite  occur  in  the  till.  Granite  is 
i'onnd  in  situ  in  the  localit}^  and  among  the  granite  boulders 
some  occur  that  seem  identical  with  this  gianite.  A  very 
coarsely  cr^'stalline  variet}^  is  also  met  with,  the  crystals  of 
felspar  being  sometimes  over  an  inch  in  length.  Though  we 
ourselves  have  not  seen  this  in  place,  3'et  the  Geological 
Surve}'  report  a  granite  with  very  large  crystals  of  felspar 
as  occurring  in  this  locality.  Pegmatite  and  aplite  also  are 
found  in  the  till.  As  both  of  these  may  occur  as  veins  in 
other  granite,  it  would  not  be  surprising  if  they  have  been 
overlooked.  It  is  not  unlikely  even  that  they  may  be  now 
concealed  beneath  the  basalt  that  is  well  developed  in  this 
distiict. 

Summing  up,  then,  the  results  of  our  investigations,  it 
would  appear  that  two  main  points  are  clearly  brought  out. 
The  first  of  these  is,  that  there  are  two  distinct  glacial 
deposits  ;  and  the  second,  that  both  of  these  deposits  are 
due  to  glacier  ice,  and  not  to  icebergs — in  fact,  both  being 
moraines  profondes.  Both  are  of  similar  character,  except 
that  the  lower  one  is  more  indurated  and  jointed.  Of  these, 
the  latter  has  been  seen  to  closely  enwrap  the  smoothed, 
grooved,  and  furrowed  surftices  of  Silurian   rocks,  of  the 


G'2       Proceedinfiii  of  the  Royal  Society  of  Victoria. 

glacier  origin  of  whieli  there  can  be  no  doubt.  It  is  useless 
at  this  stage  of  geological  inquir_y  to  maintain  that  icebergs 
can  produce  roches  moutonnees.  A  full  discussion  of  this 
point  may  be  read  in  Dr.  Croll's  "  Climate  and  Time," 
Geikie's  "  Great  Ice  Age,"  and  in  "The  Labradoi-  Coast,"  by 
Dr.  Packard.  In  connection  with  the  upper  till,  though  no 
undoubted  roches  moutonnees  have  yet  been  met  with,  yet, 
as  we  have  seen,  shattered  rock  surfaces  below  the  till  are 
found,  which  may  be  said  to  be  quite  as  characteristic  of 
the  action  of  glacier-ice  as  a  smoothed  and  moutonnee 
surface. 

It  now  becomes  a  most  important  and  interesting  question 
to  determine  the  respective  ages  of  the  two  tills.  It  seems 
certain  that  we  must  look  to  astronomy  for  the  explanation 
of  ice  ages.  Dr.  Croll's  celebrated  theory  has,  until  now, 
notwithstanding  considerable  adverse  criticism,  been  the 
most  satisfactory  ex])lanation  offeied.  Recently,  however, 
Sir  Robert  Ball  in  his  little  work  "  The  Cause  of  an  Ice  Age," 
has  re-stated  the  astronomical  theory,  pointing  out  an  error 
made  by  Croll.  It  would  be  beyond  the  scope  of  this 
present  paper  to  enter  into  a  discussion  on  the  cause  of  ice 
ages,  it  will  suffice  to  say  that  Sir  Robert  Ball  has  stated 
the  case  with  great  force  and  clearness.  The  theory  as  it 
now  stands  shows  that  when  the  astronomical  conditions 
for  the  production  of  extensive  glaciation  arise,  then  we  have 
a  period  during  which  several  glacial  epochs  alternate  with 
genial  epochs  between  the  two  hemispheres,  the  length  of 
each  epoch  being  10,500  years.  The  conditions  for  this  state 
of  things  then  graduall}^  disappear,  and  do  not  occur  again 
till  after  the  lapse  of  long  ages.  Sir  Robert  Ball  says  he 
makes  no  attempt  to  state  the  date  of  the  last  glacial  period, 
nor  to  say  when  the  next  is  to  take  place.  So,  according  to 
this  theory,  using  the  term  "period "to  embrace  several 
glacial  and  genial  "  ejwchs,"  we  should  expect  to  find 
evidence  of  glaciation  in  both  hemispheres  during  the  same 
period,  though  not  necessarily  to  the  same  extent,  for  of 
course  the  astronomical  conditions  for  glaciation  are  liable 
to  considerable  modification  by  the  existing  distribution  of 
land  and  sea,  and  the  elevation  of  mountain  chains. 

Now,  taking  the  case  of  our  lower  till  first,  we  have  seen 
that  it  is  overlaid  (apparently  unconformably)  by  rocks 
which  have  been  assigned  to  Lower  Triassic  age.  In  the 
Permian  Period  in  the  Northern  Hemisphere,  there  are  clear 
indications  of  a  glacial  epoch  or  epochs.     In  England,  Dr. 


Account  of  Glacial  Deposits  of  Bacchus  Marsh.     63 

Ramsay  describes  "  brecciated  conglomerates,"  consisting  of 
"  pebbles  and  large  blocks  of  stone,  generally  angular, 
imbedded  in  a  marly  paste."  Man}-  of  these  stones  are  as 
well  scratched  as  those  found  in  modern  moraines,  or  in 
boulder-clay.  Similar  boulder-beds  occur  in  Scotland, 
Ireland,  and  Gei'many.  Mr.  Wallace  ("  Island  Life  ")  states 
that  these  phj'sical  indications  are  corroboi'ated  by  a 
consideration  of  the  life  of  the  period,  which  is  characterised 
by  its  poveity.  In  India,  similar  Permian  boulder-beds 
occur,  in  which  large  striated  stones  and  boulde)-s  are  found. 
In  one  instance,  the  rock  surface  beneath  this  deposit  was 
glacially  scored  and  striated.  These  beds  have  beeti  corre- 
lated with  similar  ones  in  South  Afiica,  also  of  Permian  age. 
Mr.  G.  W.  Stow  has,  according  to  Dr.  Ramsay,  given 
elaborate  accounts  of  these  South  African  boulder-beds.  He 
says  that  in  Natal  the  great  masses  of  "moraine  matter"  not 
only  contain  ice-scratched  stones,  but  the  underlying  rocks 
are  well  rounded  and  mammilated,  and  covered  by  "  deeply 
incised  glacier  giooves  "  in  a  direction  that  at  last  leads  one 
to  the  p)'e-Permian  mountains,  whence  the  stones  forming 
the  moraines  have  been  derived.  In  Natal,  the  striated 
rocky  floor  is  only  30°  south,  and  in  India,  only  20°  north  of 
the  equator. 

That  evidence  of  severe  glaciation  should  be  found  in  the 
same  period  in  both  hemis))heres,  and  so  near  the  equator — 
being  actually  within  the  tropics  in  one  case — is  a  strong 
argument  in  favour  of  the  astronomical  theory,  betokening  a 
much  wider  cause  than  mere  local  elevation.  This  being  the 
case,  we  might  expect  to  find  traces  of  a  glacial  period  during 
Permian  times  here  in  the  more  southern  parts  of  Australia. 
The  position  of  our  lower  glacial  conglomerate,  or  till,  is 
quite  compatible  with  its  being  of  Permian  age,  and  when  to 
this  we  add  the  considerations  just  noted,  this  conclusion  is 
much  strengthened.  There  is  a  strong  break  in  the  flora  at 
the  close  of  the  Permo-carboniferous  series  in  New  South 
Wales  (Prof  David,  Address  A.A.A.S.,  1890).  It  is  possible 
that  this  break  may  correspond  with  a  Permian  glacial 
period. 

Now,  as  regards  our  upper  till.  We  have  not  as  yet  been 
able  to  arrive  at  any  very  definite  conclusion  as  to  the  age 
of  this  deposit.  As  we  have  seen,  it  lies  on  the  denuded 
surface  of  the  Triassic  rocks,  and  is  certainly  overlaid  by 
the  Pliocene  basalt.  That  it  is  also  overlaid  by  the  older 
basalt  admits  of  little  doubt,  for  although  this  basalt  occurs 


6-i       Proceedings  of  the  Royal  Sociefi/  of  Vicfoi-ia. 

in  the  locality,  yet  we  have  never  found  a.  trace  of  any 
volcanic  material  in  the  till.  The  same  reasoning  applies  as 
to  its  rehxtion  to  the  Miocene  leaf-beds  that  are  well 
developed  in  the  district.  These  beds  consist  for  the  most 
part  of  hard  clay-ironstone,  in  which  leaf  and  plant  impres- 
sions are  very  numerous,  and  as  a  rule  exceedingly  well 
preserved.  As  we  have  not  found  any  fragments  in  the 
till  that  in  any  way  resemble  the  material  of  these  beds,  it 
seems  highly  probable  that  the  upper  till  is  pre-Miocene. 

In  Europe,  we  have  evidence  of  glaciation  in  Eocene 
times.  In  the  "  Flysch"  of  Switzerland,  huge  erratics 
occur.  One  of  these  measured  105  ft.  in  length,  90  ft.  in 
breadth,  and  45  ft.  in  height  (Croll,  "  Climate  and  Time," 
p.  305).  Although  the  Eocene  fossils,  both  in  Europe  and 
Australia,  indicate  a  mild  climate,  yet,  as  has  been  pointed 
out  by  Croll  and  other  eminent  authoiities,  the  life  of  a 
glacial  epoch  would  be  characterised  b}-  negative  conditions. 
As  it  is  of  the  very  essence  of  the  astronomical  theory  of 
ice-ages  that  glacial  alternate  with  genial  epochs,  it  is  only 
to  be  expected  that  the  life  of  the  genial  epochs  would  be 
the  more  likely  to  be  preserved.  So  it  is  possible  that  out 
upper  till  is  Eocene  ;  this,  however,  we  merely  throw  our 
as  a  suggestion,  in  the  absence  of  any  furtiier  evidence  a.t 
present.  Considering  the  great  amount  of  erosion  that  took 
place  in  Upper  Mesozoic  and  early  Tertiary  times,  it  seems 
improbable  that  this  deposit  is  earlier  than  Eocene. 

Mr.  Stirling  and  Dr.  Lendenfeldt  have  described  undoubted 
evidences  of  glaciation  in  the  Australian  A1])S.  These 
gentlemen  found  glaciated  surfaces  on  Mt.  Cobberas  at 
elevations  between  6000  ft.  and  4000  ft.  above  the  sea,  on 
Mt.  Pilot,  and  on  Mt.  Kosciusco.  Erratics  of  huge  basaltic 
boulders  occur  in  "  linear  extension  for  miles  "  in  the  Reewa 
River  and  Snowy  Creek  valle3^s,  the  nearest  basaltic  outliers 
being  twenty  miles  away.  Perched  blocks  of  hornblende 
porphyrite  occur  on  "  crests  of  spurs  and  sidelings"  in  a 
regulai-  descending  series  from  near  the  summit  of  Mt. 
3ogong  towards  the  Reewa  valley,  many  of  them  resting 
on  smoothed  surfaces  of  pegmatite.  Moraines  occur  at  the 
base  of  Mt.  Bogong,  at  1000  ft.  above  sea  level.  Similar 
evidences  of  former  glaciation  have  also  been  described 
by  Mr.  Stirling  as  occurring  in  the  Livingstone  valley, 
Parslow's  Plains,  and  elsewhere  in  our  Alpine  regions. 

There  would  seem  to  be  no  doubt  that  the  glaciation 
iudicated   by  these  evidences  in  the  Australian  Alps  is  of 


Account  of  Glacial  Deposits  of  Bacchus  Marsh.      65 

much  more  recent  age  than  that  represented  by  the  upj3er 
till  at  Bacchus  Marsh.  The  piesence  oferractics  of  basalt, 
in  "linear  extension  "  along  the  valleys  and  on  the  slopes  of 
the  Alps  is  sufficient  to  show  this.  Dr.  Lendenfeldt 
considered  that  this  period  of  glaciation  only  terminated 
between  2000  and  3000  years  ago,  but,  as  Professor  Hutton 
has  shown,  there  is  no  evidence  to  sustain  this.  Professor 
Hutton  has  expressed  the  opinion  that  there  was  no  evidence 
to  indicate  that  the  Southern  Hemisphere  had  ever  had  a 
glacial  period.  That  glaciers  had  formerly  existed  in  the 
Australian  Alps,  he  has  explained  on  the  hypothesis  of  a 
local  elevation  of  the  Alps,  to  about  3000  feet  above  their 
present  level.  Now  this  glaciation  took  place  since  Miocene 
times,  as  is  shown  by  the  basaltic  ei'ratics.  Mr.  Stirling  has 
assigned  it  to  the  Pleistocene  Period.  It  is  impossible  that 
it  can  be  earlier,  for  if  it  were,  the  erractics  would  have  long 
ago  disappeared  from  their  positions  on  mountain  sides  and 
spurs. 

During  the  Pliocene  Period  we  have  evidence,  in  the 
distribution  of  marine  gravels,  of  a  submergence  of  nearly 
1000  feet  below  the  present  level,  and  since  then  the  land 
has  graduallj^  risen  to  its  present  condition  (Muri-ay).  In 
his  address  to  the  Biological  Section  of  the  A.A.A.S.,  at 
Hobart,  Professor  Spencer  says  : — "  We  must  conclude  from 
the  mammalian  fauna,  that  there  has  been  no  absolute  land 
connection  between  South-east  Australia  and  Tasmania 
since  practically  the  end  of  the  Tertiary  Period  or  early  in 
Pleistocene  times,  as  otherwise  it  would  be  impossible  to 
account  for  the  absence,  not  only  of  the  dingo,  but  of  the 
large  and  specialised  diprotodont  fauna,  of  which  the 
Pleistocene  Period  saw  the  rise  and  fall  upon  the  mainland." 
From  the  evidence  supplied  by  raised  beaches,  and  by  the 
great  depth  to  which  many  of  our  river  channels  have  been 
cut,  it  is  apparent  that  the  land  has  been  gradually  rising 
for  a  considerable  period.  It  is  thus  pretty  certain  that, 
since  the  beginning  of  Pleistocene  times,  the  land  surface  has 
never  stood  higher,  relatively  to  the  sea,  than  it  does  now, 
and  in  Pliocene  times,  as  we  have  seen,  there  was  a 
submergence  of  nearly  1000  feet  below  the  present  level. 

If  denudation  has  been  the  means  of  i^educing  the  height 
of  our  Alpine  regions  by  about  3000  feet  since  the  last 
glaciation  took  place,  then  it  would  be  quite  impossible  for 
lines  of  erratic  boulders  and  perched  blocks  on  mountain 
spurs  to   be   preserved.     Many  of  these,  according  to    Mr. 

F 


()<3         Proceedings  of  the  Royal  Society  of  Victoria. 

Stirling,  even  yet  show  striated  and  grooved  surfaces.  If 
the  mountains  had  suffered  much  denudation,  the  stri?e  and 
grooves  would  certainly  have  been  removed  and  roches 
moutonnees  would  have  vanished — except  where  protected 
by  overlying  deposits — long  ago. 

80  then,  seeing  that  the  theory  of  greater  elevation  cannot 
be  sustained,  we  must  look  in  another  direction  for  the 
explanation,  and  we  have  the  astronomical  theory  at  hand. 
According  to  this  theory,  we  should  expect  to  find  evidence 
of  a  Pleistocene  glacial  period  here,  corresponding  with  that 
of  the  Northern  Hemisphere.  As  we  have  seen,  this  is  the 
period  to  which  Mr.  Stirling  has  referred  this  latest  glaciation 
of  the  Australian  Alps.  As  eminent  authorities  have  already 
observed,  in  trying  to  realize  the  probable  effect  of  astro- 
nomical conditions  favourable  to  glaciation  in  the  Northern 
and  Southern  Hemispheres  iCspectively,  the  great  proportion 
of  sea  to  land  that  now  obtains  in  the  south  must  always  be 
borne  in  mind.  The  effect  of  this,  in  the  present  distribution 
of  land  and  sea,  would  undoubtedly  be  to  mitigate  these 
conditions.  In  Pleistocene  times,  there  is  no  evidence  to 
show  that  oui-  mountains  were  appreciably  higher  than  now  ; 
it  seems  more  probable  that  our  land  surface  stood  actually 
lower.  So  that  the  astronomical  conditions  which,  during 
this  period,  resulted  in  producing  such  a  severe  glaciation 
in  the  Northern  Hemisphere,  weie  probably  so  mitigated  in 
the  Southern  Hemisphere  that  glaciers  only  appeared  in  the 
higher  mountains. 

Mr.  G.  S.  Griffiths,  in  a  papei'  on  the  "Evidences  for  a 
post-Miocene  Glacial  Period  in  Victoria,"  describes  heavy 
boulder  washes,  distributed  in  many  parts  of  the  Colony. 
These  "  washes  "  are  ascribed  to  glacial  action.  Though  the 
evidence  for  this  is  not  conclusive,  yet  it  is  by  no  means 
improbable  that  these  heavy  deposits  of  bo\ilders — many  of 
them  basaltic — were  formed  at  the  period  of  the  last  glacia- 
tion of  the  Alps,  when  the  precipitation  was  much  greater 
than  now.  The  Dividing  Range,  except  in  its  eastern  parts, 
not  being  high  enougli  for  the  production  of  glaciers,  in  the 
short  hot  summers  of  the  epoch  vast  floods  from  melting 
snow  swept  down  from  the  mountains,  swelling  the  rivers 
and  depositing  these  boulder  bed.s. 

At  the  two  earlier  periods  of  glaciation  we  have  indicated, 
it  is  not  improbable  that  there  was  a  greater  southward 
extension  of  land  by  way  of  Tasmania  than  now  obtains. 
In  Upper  Palaeozoic  times,  tlie  Main  Divide  must  have  stood 


Accoiuit  of  Glacial  Deposits  of  Bacclais  Mar«h.      07 

many  thousand  feet  higher  tluiii  it  does  now.  In  Eocene 
times,  though  enormous  denudation  had  then  taken  ])lace, 
this  mountain  chain  must  have  been  veiy  much  higher  than 
now.  Under  these  conditions,  the  glaciation  during  an  ice 
age  might  be  of  considerable  severity. 

We  would  thus  appear  to  ha\e  evidence  of  three  periods 
of  glaciation  in  Australia,  which  may  be  provisionally 
assigned  as  follows  : — (1)  One  in  Permian  times,  of  consider- 
able severity  ;  (2)  one  in  Eocene  times,  also  severe  ;  (3) 
one  in  Pleistocene  times,  mild,  being  represented  only  by 
glaciers  in  the  higher  mountains.  At  these  periods  then,  it 
would  appear  that  the  Dividing  Range  nourished  great 
glaciers  which  radiated  outwards,  and,  in  the  two  earlier 
periods  at  least,  spread  to  some  distance  over  the  lower 
ground.  Beneath  these  glaciers  the  till,  or  glacial  conglomer- 
ate, was  accumulated  as  a  ground  moraine. 

Undoubted  evidence  of  glaciation  has  been  adduced  by 
Professor  Tate  and  Mr.  G.  B.  Pritchard  from  South  Austi-alia. 
and  traces  have  also  been  noted  in  Tasmania,  although 
Mr.  Johnston  remarks  ("  Geology  of  Tasmania  ")  that  there 
is  no  evidence  there  to  show  that  a  glacial  period  has  ever 
taken  place.  However,  it  will  seem  strange  if  further 
evidence  from  Tasmania  be  not  forthcoming. 

In  concluding  this  pajjer,  we  would  urge  the  careful 
examination  and  mapping  of  our  glacial  deposits,  and  the 
collection  of  all  evidence  bearing  upon  them.  In  the  words 
of  Sir  Robert  Ball—"  A  strict  search  for  glacial  indications 
among  all  deposits,  primary,  secondary,  and  tei-tiary,  would 
be  one  of  the  most  valuable  pieces  of  scientific  work 
possible  at  the  present  time." — ("  Cause  of  an  Ice  Age," 
p.  149). 

Literature. 

Prof  David— ^' A  Correlation  of  the  Coal-fields  of  N.S.W." 

A.A.A.S.,  1890. 
E.  J.  Dunn,  F.G.S. — "  Notes  on  the  Occurrence  of  Glaciated 

Pebbles  and    Boulders    in  the    so-called    Mesozoic 

Conglomerate  of    Victoria,"   Proc.   Roy.  Soc.   Vic, 

Vol.  XXIV,  Pt.  1.     "  The  Glacial  Conglomerates  of 

Victoria,"  A. A.  AS.,  1890. 
O.  S.  Griffiths,  F.G.S.—"  Evidences  of  a  Glacial  Period  in 

Victoria  during  post-Miocene  Times,"  Ti-ans.  Rov. 

Soc.  Vic,  Vol.  XXI. 

F   2 


C8        Proceedings  of  the  Ro>jal  Sooietij  of  Victoria. 

T.  S.  Hall,  M.A.— "  Victorian  Naturalist,"  Vol.  VIII,  Mo.  2. 

Plot".  Hutton,  F.G.S. — "  The  Supposed  Glacial  Epoch  in 
Australia,"  Proc.  Lin.  Soc.  N.S.W.,  Vol.  X,  Pt.  2. 

P.  M.  Johnston,  F.G.S. — "  The  Geology  of  Tasmania." 

R.  Von  Lendenfeldt,  Ph.  D. — "  The  Glacial  Period  in  Aus- 
tralia," Proc.  Lin.  Soc.  N.S.W.,  Vol.  X. 

Sir  Frederick  McCoy,  K.C.M.G.,  F.R.S.,  &c.— "  Prodromus 
of  the  Palaeontology  of  Victoria,"  Decades  II  and 
IV. 

R.  A.  Murray. — "  Geology  and  Physical  Geography  ot 
Victoria." 

Prof.  W.  Baldwin  Spencer,  M.A. — "  The  Fauna  and  Zoological 
Relationships  of  Tasmania,"  A.A.A.S.,  1892. 

Jas.  Stirlhig,  F.L.S.,  F.G.S. — "  Notes  on  Some  Evidences  of 
Glaciation  in  the  Australian  Alps,"  Trans.  Roy.  Soc. 
Vic,  Vol.  XXII.  "  On  Some  Further  Evidences 
of  Glaciation  in  the  Australian  Alps,"  Proc.  Lin. 
Soc.  N.S.W.,  188G. 

Annual  Report  of  the  Mining  Department  of  Victoria  for 
1891,  p.  30. 

Also, 

Sir  Robert  Ball. — "  The  Cause  of  an  Ice  Age." 

Dr.  Croll. — "  Climate  and  Time." 

Prof  Jas.  Geikie. — "  The  Great  Ice  Age." 

Dr.  Packard. — "  The  Labrador  Coast." 

Dr.  Ramsay.—"  Nature,"  Vol.  XXII,  p.  389. 

Dr.  Wriofht. — "  The  Ice  Affe  in  North  America." 


Proc.    R    S      Victoria    Plate  X 


R.    S.    VICTORIA,    PLATE 


PHOTOLINE 


516    GEORGE    ST.,    SYDNEY 


PROC,    R.    S.    VICTORIA,    PLATE 


PHOTOLINE    PRINTING    CO., 


515    GEORGE    ST.,    SYDNEY 


Art.  VI. — Sij}iopsi.'^  of  the  Australian  Calcarea  Hcterocada  ; 
with  a  proposed  Classification  of  the  Group  and 
Descriptions  of  some  Neio  Genera  and  Species. 

By  Arthur  Dendy,  D.Sc,  F.L.S. 

Demonstrator  and  Assistant  Lecturer  in  Biology  in  the  University  of 
Melbourne. 

[Read  September  8,  1892.] 


1.   Introductory   Remarks. 

•  Owing  to  the  reduction  of  the  Government  grant  to  the 
Royal  Society,  it  has  been  found  impossible  at  present  to 
continue  the  publication  of  the  volume  of  the  Transactions 
which  the  Council  of  the  Society  generously  set  apart  for 
"A  Monograph  of  the  Victorian  S[)onges."  Under  these 
circumstances  the  Council  has  kindly  agi-eed  to  m}^  proposal 
to  divide  the  work  by  publishing  in  the  Proceedings  of  the 
Societ}'  only  the  necessary  systematic  portion.  Avithout 
illustrations,  and  publishing  the  anatomical  portions,  for 
which  numerous  illustrations  are  indispensable,  elsewhere. 
It  is  hoped,  however,  that  the  continuation  of  the  Mono- 
graph in  its  original  form  may  be  carried  out  at  some  future 
date. 

I  have  in  my  possession,  in  addition  to  the  very  fine  series 
of  Victorian  Heterocoela  collected  by  Mr.  Bi-acebridge  Wilson, 
a  number  of  very  valuable  specimens,  including  sevei'al 
remarkable  new  species,  from  otiier  parts  of  Australia,  and 
it  seemed  desirable  to  incorporate  descrij)tions  of  these  in  the 
present  communication.  I  have  also  a  number  of  fragments 
of  type  specimens,  generously  forwarded  to  me  by  the 
authorities  of  the  British  Museum,  which  brings  the  total 
number  of  species  of  Australian  Calcarea  Heterocoela  in  m}^ 
collection  up  to  forty-seven. 


70        Proceedings  of  the  Royal  Society  of  Victoria. 

Up  to  the  present  time  sixtj'-tvvo  species  of  thi.s  group  of 
sponges  have  been  described  from  Australian  seas,  chiefly  by 
HfBckel,  von  Lendenfeld,  Polejaeff  and  Carter.  Sixteen 
new  species  are  described  in  the  present  paper,  which  brings 
the  total  number  of  Australian  species  of  Calcarea  Heterocoela 
u[)  to  seventy-eight.  In  order  to  make  the  work  as  complete 
as  possible,  I  have  decided  to  include,  not  onl}'  descri[)tions 
or  notes  of  the  forty-seven  species  which  I  have  been  able  to 
study  for  myself,  but  also  references  to  those  species  which 
1  have  not  seen,  and  thus  to  provide  a  complete  Synopsis  of 
the  Australian  Calcarea  Heterocoela. 

In  or(ier  to  arrange  the  sjiecies  satisfactorily,  I  have  been 
obliged  to  adopt  a  classification  which  has  many  new 
features.  This  classification  has  not  been  arrived  at  hastily, 
nor  without  careful  consideration  of  the  work  of  previous 
writers.  It  is  impossible  to  justify  it  fully  in  this  place, 
because  it  is  based  upon  a  minute  study  of  anatomical 
details  and  a  careful  consideration  of  historical  questions  of 
y)riority  in  nomenclature  into  which  I  have  not  space  to 
enter.  I  intend,  however,  to  publish  a  paper  on  the  minute 
anatomy  and  classification  of  the  group  in  another  place,  in 
which  these  questions  will  be  fully  discussed  ;  and,  in  the 
meantime,  I  may  perhaps  point  out  that  the  classification 
proposed  is  based  upon  tlie  careful  anatomical  examination  of 
a  very  large  number  of  species. 

It  will  be  seen  that  more  stress  is  laid  upon  the  arrange- 
ment of  tiie  skeleton  than  is  usual  at  the  present  day,  and 
less  upon  the  form  and  arrangement  of  the  fiagellated 
chambers,  which  I  find  to  vary  considerably,  even  within 
the  limits  of  a  single  species.  This  change  certainly  facili- 
tates the  identification  of  specimens,  and  will  probably  be 
welcomed  by  those  workers  who  have  not  at  their  disposal 
the  elaborate  appliances  required  for  the  preparation  of 
stained  microscopical  sections. 

Polejaefi'*  clearly  showed  that  no  hard  and  fast  line  could 
be  drawn  between  the  Sycons  and  Leucons  of  Hfeckel.  'J'his 
idea  was  followed  up  by  von  Lendenfeld,  who  has  created  a 
special  group,  the  iSylleibidai,-\-  to  include  the  intermediate 
forms.  This  group,  however,  seems  to  me  to  be  very  arti- 
ficial, as,  judging  from  my  own  observations,  it  appears  that 

*  Keport  ou  the  Calcarea  of  the  Challenger  Expedition. 

t  See  especially  "  Die  Si^ougieu  der  Adria.  I.  Die  Kalkscliwamme." 
Zeitsclirift  fiir  WissenschaftlicheZoologie,  Vol.  5H,  1891. 


Synopsis  of  the  Australian  Calcarea  HetevocMa.    71 

the  transition  from  the  Sycon  to  the  Leucon  type  of  canal- 
system  has  not  taken  place  along  a  single  line  of  descent, 
hut  along  several.  Here,  as  in  other  cases,  we  must  classify 
by  an  assemblage  of  characters.  The  canal-system  alone  is 
l)y  no  means  sufficient,  though,  when  taken  in  conjunction 
with  the  skeleton,  it  is  often  of  great  value. 

In  enumerating  the  various  genera  and  species  I  have  not 
attempted  to  give  a  complete  list  of  synonyms  and  references, 
as  this  would  have  taken  up  a  large  amount  of  space.  In  the 
case  of  the  species,  however,  I  have  given  those  synonyms 
and  references  which  are  most  important. 

My  warmest  thanks  are  due  to  Mr.  J.  Bracebridge  Wilson 
for  the  bulk  of  the  specimens  here  described,  to  Mr.  Thos. 
Whitelegge  for  a  very  valuable  collection  from  Port  Jackson,' 
to  the  Adelaide  Museum  for  some  very  interesting  specimens 
from  St.  Vincent's  Gulf,  to  Professor  Spencer  foi-  a  number 
of  specimens  from  Port  Jackson,  to  Sir  Frederick  M'Coy  for 
permission  to  examine  the  collection  in  the  Melbourne 
National  Museum,  and  to  Dr.  Glinther  for  fragments  of  type 
sjiecimens  in  the  British  Museum. 


2.     Proposed   Classification^   of   the   Calcarea 
Heteroccela. 

Order    HETEROCCELA,    Polejaeff. 

Calcareous  sponges  in  which  the  collared  cells  are  confined 
to  more  or  less  well-defined  flagellated  chambers. 


Family  1. — Lelx'ascid.e. 

Flagellated  chambers  very  long  and  nai'row,  copiousl}^ 
branched  ;  communicating  at  their  proximal  ends  with 
exhalant  canals  which  converge  towards  the  oscula  ;  their 
blind  distal  ends  covered  over  by  a  dermal  membrane 
pierced  by  true  dermal  pores  which  lead  into  the  irregular 
spaces  between  the  chambers.  Skeleton  consisting  princi- 
pally of  small  radiates  irregularly  scattered  in  the  walls  of 
the  chambers  and  exhalant  canals  and  in  the  dermal 
membrane. 


72        Proceedin(/s  of  the  Royal  SGciety  of  Victoria. 

Genus  I. — Leiica^cun,  nov.  g-en. 
Diagnosis. — The  same  as  that  of  the  family. 
(For  species  see  Part  3  of  the  present  paper.) 


Family  -1. — Sycettid^. 

Flagellated  chambers  elongated,  ai-ranged  radially  around 
a  central  gastral  cavity,  their  distal  ends  pn^jecting  more  or 
less  on  the  dermal  surface  and  not  covered  over  by  a  con- 
tinuous cortex.     Skeleton  radially  S3aiimetrical. 

Genus  II. — Sijcdta  (Hieckel,  emend.) 

Diagnosis. — Radial  chambers  not  inter-commuuicating. 
Articulate  tubar  skeleton.  No  tufts  of  oxea  on  the  distal 
ends  of  the  chambers. 

(For  example  see  Part  3  of  the  present  ])aper.) 

Genus  III. — Sycon  (Risso,  emend.) 

Diagnosis. — Radial  chambers  not  inter-communicating. 
Articulate  tubar  skeleton.  The  distal  ends  of  the  chambers 
provided  each  with  a  tuft  of  oxeote  spicules. 

(For  examples  see  Part  3  of  the  present  paper.) 

Genus  IV. — Sycaiifha,  von  Lendenfeld. 

Radial  chambers  long,  united  in  groups  ;  those  of  each 
group  inter-communicating  by  openings  in  their  walls  and 
each  group  with  a  single  common  opening  into  the  gastral 
cavity.  The  radial  chambers  have  freel}'  projecting  distal 
cones  surmounted  by  oxeote  spicule.s.  Tubar  skeleton 
articulate. 

No  Australian  species  of  this  genus  has  yet  l)een  found. 
The  t3qoe  is  von  Lendenfeld's  SycantJiu  tenella.* 


Family  3. — Grantid.e. 

There  is  a  distinct  and  continuous  dermal  cortex  covering 
over   the    chamber   layer   and    pierced    by  inhalant    pores. 

*  "  Die  Spougien  cler  Aclria.     I.    Die  Kalliscliwiimme,"  p.  51. 


St/uopsis  of  the  Australian  Calcarea  Heterocoda.    73 

There  are  no  subderraal  sagittal  triradiafces  or  quadriradiates.* 
The  flagellated  chambers  vary  from  elongated  and  radially 
arranged  to  spherical  and  irregularly  scattered,  while  the 
skeleton  of  the  chamber  layer  varies  from  regularly  articulate 
to  irregularly  scattered. 

Genus  V. —  Grant ia  (Fleming,  eraend.) 

Diagnosis. — The  elongated  flagellated  chambers  are  ar- 
ranged radially  around  the  central  gastral  cavity  ;  they  are 
not  provided  with  tutts  of  oxea  at  their  distal  ends,  but  are 
covered  over  by  a  dermal  cortex  composed  ])rincipally  of 
triradiate  spicules  and  without  longitudinally  disposed  oxea. 
An  articulate  tubar  skeleton  is  presents 

(For  examples  see  Pai-t  3  of  the  present  paper.) 

S  lib-Gen  us. — Gra  nt  iops  is,  no  v. 

Diagnosis. — The  sponge  has  the  form  of  a  greatly  elongated, 
hollow  tube,  whose  wall  is  composed  of  two  distinct  layers 
of  about  equal  thickness.  The  outer  (cortical)  layer  is 
provided  with  a  very  strongly  developed  skeleton  of  radiate 
spicules  and  contains  the  narrow,  ramif}dng  inhalant  canals. 
The  inner  (chamber)  layer  is  formed  b}^  elongated  I'adial 
chamljers  arranged  very  regularly  side  by  side.  The 
skeleton  of  the  chamber  layer  is  very  feebly  developed ;  the 
normal  subgastral  triradiates  ai-e  replaced  by  quadiiradiates  ; 
the  tubar  skeleton  is  articulate,  and  composed  of  very 
abnormal  sagittal  triradiates  whose  ])aired  rays  are  greatl}^ 
reduced. 

(For  species  see  Part  3  of  tlie  present  paper.) 

Genus  VI. —  Ute  (Schmidt,  emend.) 

Diagnosis. — The  ends  of  the  elongated  radial  chambers 
are  covered  over  by  a  well  developed  cortex,  consisting  in 
great  part  of  large  oxeote  spicules  arranged  parallel  to  the 
long  axis  of  the  sponge.  Tlie  tubar  skeleton  is  articulate 
or  else  composed  entirely  of  the  basal  rays  of  subgastral 
triradiates. 

(For  examples  see  Part  3  of  the  present  paper.) 

*  I  propose  these  names  for  spicules  lying  beneath  the  dermal  surface  and 
with  inwardly  directed  basal  or  apical  rays  as  the  case  may  be.  Such 
spicules  are  of  great  importance  for  purposes  of  classification. 


7-i        Procee(/inf/s  of  the  Royal  Society  of  Victoria. 

Sub-Genus. — Synute,  Dendy. 

Dhifj'nos'o-;.  —  Sponge  compound,  consisting  of  many 
Ute-lika  individuals  completely  fused  together  and  invested 
in  a  common  cortex  composed  largely  of  huge  oxeote 
spicules. 

(B'or  species  see  Part  .'^  of  tlie  ]_)resent  ]mper.) 

Genus  VII. —  Utella,  nov.  gen. 

Diagnosis. — Flagellated  chambers  elongated,  arranged 
I'aclially  around  the  central  gastral  cavity.  ThfU'e  are  no 
longitudinally  arranged  oxea  in  the  dermal  cortex,  but  a 
layer  of  oxeote  spicules  lies  beneath  and  parallel  to  the 
gastral  surface.     The  tubar  skeleton  is  articulate. 

I  propose  this  genus  for  H^ckel's  Sycandra  hystrix* 
Schmidt's  Ute  utriculus  (the  Sycandra  utriculus  of 
Hgeckel-f-)  may  perhaps  also  be  included  therein. 

No  Australian  species  are  as  yet  known. 

Genus  VIII. — Anamixilla   (PolejaeH"  emend.) 

D'nujnosls. — Flagellated  chamber's  elongated  and  radially 
arranged.  There  is  no  special  tubar  skeleton,  the  skeleton 
of  the  chamber  layer  consisting  of  large  radiate  spicules, 
arranged  without  regard  to  the  direction  of  the  chambers, 
and  of  the  outwardly  directed  basal  rays  of  subgastral 
tri  radiates. 

(For  species  see  Part  3  of  tl)e  present  paper.) 

Genus  IX. — Sycyssa,  Hpeckel. 

Diagiiosis. — The  tiagellated  chambers  are  elongated  and 
arranged  radially  around  the  central  gastral  cavity.  The 
skeleton  consists  exclusively  of  oxeote  spicules. 

The  only  known  species  of  this  genus  is  Hyeckel's  Sycyssa 
huxleyi,\  from  the  Adriatic. 

Genus  X. — Limeandra  (Hreckei,  emend.) 

Diagnosis. — The  flagellated  chambers  are  spherical  or  sac- 
shaped,  irregularly  ari-anged.  and   communicating  with  the 


Die  Kalk.scliwiiinme.     Vol.  II,  p.  375.  t  Loc  cit.,  p.  370. 

I  Loc.  cit.,  p.  260. 


Synopsis  of  the  Au.sfralian  Calcarea  Heterocada.    75 

gastral  cavity,  or  main  exhalant  canals,  1)}^  a  more  or  less 
complicated  canal-system.  The  skeleton  of  the  chamber 
layer  is  irregular. 

(For  examples  see  Part  3  of  the  present  paper.) 

Genus  XI.  — Lelapia  (Gray,  emend.) 

Diagnosis. — Canal  ,S3^stem  unknown.  Skeleton  of  gastral 
surface  composed  of  ordinary  radiates.  Skeleton  of  dermal 
surface  composed  of  triradiates,  quadriradiates  and  minute 
oxea.  Skeleton  of  the  chamber  layer  composed  of  large, 
longitudinally  arranged  oxea,  crossed  at  right  angles  by 
bundles  of  tuning-fork-shaped  triradiates  whose  oral  rays 
are  directed  towards  the  gastral  cavit}^  while  the  basals 
point  towards  the  dermal  surface. 

(For  species  see  Part  8  of  the  present  paper). 

Genus  ^ll.—Leucyssa,  Hreckel. 

Diagnosis. — Flagellated  chambers  (presumably)  spherical 
or  ovoid,  irregularly  arranged.  Skeleton  composed  solely  of 
oxeote  spicules. 

No  species  of  this  remarkable  genus  ai'e  as  yet  recorded 
from  Australian  seas,  the  only  examples  being  Hseckel's 
Leucyssa  spongilla,  L.  crdacsa  and  L.  incrustans.^ 


Family  4. — HETEROPiD/t:. 

There  is  a  distinct  and  continuous  dermal  cortex  covering 
over  the  chamber  laj'er  and  pierced  by  inhalant  pores. 
Subdermal  sagittal  triradiates  are  present.  The  flagellated 
chambers  vaiy  from  elongated  and  radially  arranged  to 
spherical  and  irregularly^  scatteied.  An  articulate  tubar 
skeleton  may  or  may  not  be  present. 

Genus  XIII. — Grantessa  (von  Lendenfeld,  emend.) 

Diagnosis. — The  flagellated  chambers  are  elongated  and 
arranged  radially  around  the  central  gastral  cavity.  The 
dermal  cortex  consists  principally  of  triradiates  and  does 
not  contain  longitudinally  disposed  oxea. 

(For  examples  see  Part  3  of  the  present  paper.) 

*  Loc.  cit.,  pp.  137,  138,  139. 


76       Proceedings  of  the  Royal  Society  of  Victoria. 

Genus  XIV. — Heteropia  (Carter,  emend.) 

Diagnosis. — The  distal  ends  of  the  elongated  radial 
chambers  are  covered  over  by  a  well-developed  dermal 
cortex,  consisting  principally  of  large  oxea  arranged  parallel 
to  the  long  axis  of  the  sponge. 

I  propose  to  retain  this  generic  name  for  Carter's  Apltvo- 
ceras  raniosa,*  which  he  observes  belongs  to  his  genus 
Heteroina.     No  species  are  yet  known  from  Australia. 

Genus  XV. —  Vosmaerop.sis,  nov.  gen. 

Diagnosis. — Flagellated  chambers  spherical  or  sac-shaped, 
never  truly  radial.  Dermal  cortex  composed  principally  of 
triradiates,  without  longitudinally  disposed  oxea. 

(For  species  see  Part  3  of  the  present  paper.) 


Family  5. — Amphoriscid^. 

There  is  a  distinct  and  continuous  dermal  cortex  covering- 
over  the  chamber  layer.  Subdermal  quadrii-adiates  are  pre- 
sent. The  flagellated  chambers  var}^  from  elongated  and 
radially  arranged  to  spherical  and  irregularly  scattered. 

Genus  XVI. — Heteropegma  (PolejaefF,  emend.) 

Diagnosis. — The  flagellated  chambers  are  elongated  and 
arranged  radially  around  the  cential  gastral  cavity.  There 
is  a  vestigial  tubar  skeleton  of  minute  radiates.  The  dermal 
coi'tex  is  very  thick,  composed  principall}'  of  large  triradiate 
spicules. 

(For  species  see  Part  3  of  the  present  paper.) 

Genus  XVII. — Amphoriscus  (Hreckel,  emend.) 

Diagnosis. — The  flagellated  chambers  are  elongated  and 
arranged  radially  around  the  central  gastral  cavity.  The 
skeleton  of  the  chamber  layer  is  composed  exclusively  of  the 
apical  rays  of  subdermal  and  subgastral  quadriradiates. 

(For  examples  see  Part  3  of  the  present  paper.) 

Genus  XVIII. — Sycidmis  (H?eckel,  emend) 

Diagnosis. — The  flagellated  chambers  are  elongated  and 
ari-anged  radially   around  the   central  gastral  cavity.     The 

*  Proc.  Lit.  PhiL  Soc.  Liverpool,  VoL  XL,  Appeudi-N:,  1886,  p.  92. 


synopsis  of  the  Ausfrdllaii  Calcarea  Heterocwla.    77 

skeleton  of  the  chamber  layer  is  composed  of  the  apical  rays 
of  subdermal  and  subgastral  quadriradiates.  There  is  a 
root-tuft  of  oxea  and  anchoring  quadriradiates. 

The  only  known  species  of  this  remarkable  genus  is 
Hiieckcl's  Sijculmis  suYKipla,^'  from  Bahia. 

Genus  XIX. — LcuciUa  (Ha^ckel,  emend.) 

Diagnosis. — Flagellated  chambers  spherical  or  sac-shaped, 
never  truly  radial. 

(For  examples  see  Part  o  of  the  present  paper.) 

Genus  XX. — Faraleucilla,  nov.  gen. 

Diagnosis. — Chambers  spherical  or  sac-shaped  (?).  Be- 
neath the  dermal  cortex  a  series  of  subdermal  cavities 
supported  by  an  outer  and  inner  layer  of  quadriradiates 
whose  apical  xixya  cross  each  other  in  opposite  directions. 
Beneath  these  comes  the  chamber  layer,  whose  skeleton 
consists  ot  irregularly  arranged  quadriradiates.  Large, 
longitudinally  arranged  oxea  occur  between  the  triradiates 
of  the  dermal  cortex. 

(For  species  see  Part  3  of  the  present  papei'.) 


o.   Synopsis  of  the   Australian   Species  of   Calcarea 
Heteroccela. 

1.   Leucascus  simplex,  n.  sp. 

Sponge  usually  more  or  less  flattened,  cushion-shaped, 
spreading,  with  convex  upper  surface  ;  sometimes  becoming 
almost  globular.  Oscula  irregularly  scattered  on  the  upper 
surface,  one  or  several,  varying  in  size,  naked.  Surface 
smooth.  The  largest  specimen  is  a  rather  thin,  ovoid,^ 
flattened  ci'ust,  which,  from  its  shape,  appears  to  have 
grown  on  a  crab's  back ;  it  is  about  3o  mm.  long,  20  mm. 
broad,  and  only  about  2  mm.  thick  in  most  parts  ;  the  other 
specimens,  though  smaller,  are  much  thicker,  one  being 
nearly  spherical.  The  surface  is  covered  by  a  thin,  pore- 
bearing  dermal  membrane. 

*  Die  Kallvschwamme,  VoL  11,  p,  288. 


7«S       Froceedings  of  the  Royal  Society  of  Victoria. 

The  flagellated  chambers  (if  one  may  use  the  term)  are 
greatly  elongated,  tubular  and  copiously  branched  ;  their 
terminal  branches  end  blindly  beneath  the  dermal  mem- 
biane.  I'heir  walls  are  thin  and  pierced  by  numerous 
})rosopyles.  There  is  no  central  gastral  cavity  but  the 
chambers  open  into  exhalant  canals,  devoid  of  collared  cells, 
which  convei'ge  towards  the  oscula.  The  scattered  dermal 
pores  lead  into  wide,  irregular  sjjaces  between  the  tubular 
chambers. 

The  skeleton  is  extremely  simple,  consisting  of  small, 
regular  triradiates,  irregularly  scattered  in  the  walls  of  the 
chambers  and  exhalant  canals,  and  in  the  dermal  membrane. 
All  the  spicules  are  alike,  except  that  some  exhibit  an 
incipient  apical  ray.  The  rays  aie  stiaight,  conical,  fairly 
sharply  pointed  ;  measuring  about  ()"1  by  001  mm. 

Localities. — Near  Port  Phillip  Heads  (Stations  1  a)id  5, 
coll.  J.  B.  Wilson)  ;  Watson's  Bay,  Port  Jackson  (coll.  T. 
Whitelegge). 


2.    Leucasciis  clavatus,  n.  sj). 

The  t3'pe  specimen  of  this  species  is  a  sub-globular  sponge 
about  14  mm.  in  maximum  diameter,  with  a  single  rather 
wide  osculum.  It  very  closely  resembles  L.  simplex  in 
general  appearance,  canal  system  and  skeleton,  and  the  only 
point  of  distinction  of  any  importance  which  I  have  been 
able  to  find  consists  in  the  presence  in  L.  clavatus  of  large 
club-shaped  oxea  lying  at  right  angles  to  and  with  the 
club-shaped  extremity  projecting  slightl}'  beyond  the  dermal 
surface.  These  spicules  attain  a  length  of  about  07,  and  a 
diameter,  in  the  thickest  part,  of  about  Ol  mm.  The  outer 
end  of  the  spicule  is  usually  more  or  less  rounded  off  and 
slightly  curved,  while  the  inner  portion  is  straight  and 
tapers  gradually  to  a  sharp  point.  The  triradiates  are  like 
those  of  L.  simplex,  and  of  nearly  the  same  size,  perhaps  a 
little  larger  on  an  average, 

A  second  specimen  is  ii-regularly  lobate,  and  differs  from 
the  type  in  its  much  denser  texture,  which  is  due  to  the 
stronger  development  of  the  mesoderm.  It  contains  very 
numerous  etnbryos,  which  fact  may  be  associated  with  the 
strong  development  of  the  mesoderm. 

Locality.— 'Nenv  Port  Phillip  Heads  (coll.  J.  B.  Wilsoii). 


Synopsis  of  the  Australian  Calcarea  Heierocoila.    79 

3.    Sycetta  primUiva,  Ha?ckel. 

Si/cetta  prlmitiva,  Hseckel,  Die  Kalkscliwanime,  Vol.  II, 
p,  237. 

Locality. — Bass  Straits,  Gulf  of  St.  Vincent  (Hasckel). 

4.    Sycon  coroiiatum,  Ellis  and  Solaiider,  sp. 

Spongia  coronata,  Ellis  and  Solander.    Zoophytes,  p.  190. 

Grantia  ciliata,  auctoruni. 

Sycandra  coronata,  Hseckel.  Die  Kalkschwamme,  Vol.  II, 
p.  804.. 

Locality. — East  coast  of  Austi-alia  (Hc>3ckel.  Also  recorded 
from  the  Mediterranean,  Atlantic  and  Pacific). 

5.    Sycon  carteri,  n.  sp. 

Colonial;  consisting  of  very  many  small  Sycon  individuals 
united  in  a  copiously  branching,  bushy  mass ;  branching 
irregular.  Sycon  individuals  about  5  mm.  in  length  b}' 
15  mm.  in  diameter;  cylindrical;  with  minutely  conulose 
surface  and  naked,  terminal  oscula. 

Canal-system  typical ;  chambers  thimble-shaped,  rather 
short,  with  freely  projecting  distal  cones. 

Skeleton  ai-ranged  in  typical  manner-.  Spicules  as 
follows: — (1)  Gastral  quadriradiates  ;  sagittal;  oral  rays 
shorter  and  stouter  than  basal,  slightl^y  recurved,  gradually 
sharp-pointed,  measuring  Oil  x  0()07  mm.;  basal  ray 
rather  more  slender,  straight,  very  gradually  sharp-pointed, 
about  0"2  mm.  long  ;  apical  ray  variable,  stout,  more  or  less 
curved,  often  angulate,  gradually  sharp-pointed,  about 
0'077  nim.  long.  (2)  Gastral  triradiates  ;  like  the  quadri- 
radiates  but  without  the  apical  ray.  (3)  Subgastral  sagittal 
triradiates  ;  oral  rays  extended  almost  in  a  line,  gradually 
sharp-pointed,  measuring  about  0*06  x  0007  mm. ;  basal 
ray  veiy  long  (0175  mm.),  straight,  gradually  sharp-pointed, 
extending  for  more  than  half  the  length  of  the  chamber  and 
forming  by  itself  about  half  of  the  articulate  tubar  skeleton. 
(4)  Ordinary  tubar  triradiates  ;  like  the  last  but  with 
shorter  basal  ray  and  oral  angle  diminishing  towards  the 
distal  cone.  (o)  Oxea  of  the  distal  cones  ;  rather  shoi't 
and  stout,  angulate,  with  shorter  and  stouter  outer,  and 
longer  and  slenderer  inner  segments  ;  fairly  sharp-pointed  at 
both  ends  ;  measuring  about  0  15  x  001  mm. 

Locality. — St.  Vincent's  Gulf,  S.  Australia,  (coll.  Adelaide 
Museum). 


80        Proceediiif/s  of  the  Royal  Society  of   Victoria. 

6.    Sycon  minutum,  ii.  sp. 

Solitary;  sessile,  or  with  very  short  stalk;  sub-cylindrical 
or  sausage-shaped,  with  naked  terminal  osculum  surrounded 
by  a  membianous  extension  cxf  the  wall  of  the  gastral  cavity. 
Texture  characteristically  soft  and  s])ongy;  surface  minutely 
conulose.  Usually  only  abotit  5  or  6  mm.  in  height  by 
2  mm.  in  breadth. 

Canal  system  typical  ;  chambers  rather  short,  thimble- 
shaped,  often  octagonal  in  transverse  section,  with  low 
rounded  distal  cones ;  inhalant  canals  usuall}'  square  in. 
transverse  section. 

Skeleton  arranged  as  usual.  Spicules  as  follows : — 
(1)  Gastral  quadriradiates  ;  facial  rays  straight,  subequal  in 
length,  very  long,  slender  and  gradually  sharp-pointed, 
measuring  about  012  by  00035  mm.  ;  oral  angle  somewhat 
smaller  than  the  paired  angles;  apical  ray  short,  relatively 
stout,  slightly  curved,  sharp-pomted,  about  0-03  mm.  long. 
Towards  the  osculum  these  spicules  become  much  more 
markedly  sagittal.  (2)  Gastral  triradiates ;  like  the  foi'e- 
going,  but  without  apical  ray.  (3)  Subgastral  triradiates, 
not  distinguishable  in  form  from  the  ordinary  tubar  spicules. 

(4)  Tubar  triradiates  ;  varying  from  sagittal,  with  very 
widely  e::tended,  slightly  curved,  oral  rays,  to  sub-regular; 
rays  long  and  slendei',  gradually  sharp-pointed,  the  basal 
not  much  longer  than  the  orals,  measuring  about  O'l  by 
0006  mm.  ;    these  spicules  are  rather  irregularly  arranged. 

(5)  Oxea  of  the  distal  cones ;  not  very  numerous  ;  long, 
slender,  straight  or  very  slightly  curved ;  fusiform  and 
gradually  sharp-pointed  at  each  end  ;  measuring  about  0'28 
by  0007  mm. ;  arranged  in  loose  tufts  which  project 
obliquely  upwards  from  the  distal  cones  towards  the 
osculum. 

Locality. — Watson's  Bay,  Ft.  Jackson  (coll.  T.  Whitelegge). 

7.    Sycon  raphaiius,  0.  Schmidt. 

Sycon  raphanus,  0.  Schmidt.     Spong.  Adriat.  Meer.,  p.  14. 

Abundant  in  the  collection.  Solitary,  usually  about  half 
an  inch  in  height,  with  well  developed  stalk,  markedly 
conulose  surface  and  small  oscular  fringe.  In  spiculation  I 
can  find  no  tangible  distinction  between  this  common 
Victorian  species  and  the  European  S.  raphanus  as  described 
by  Hajckel  in  "  Die  Kalkschwamme." 


Synopsis  of  the  Australian  Galcarea  Heterocosla.    81 

Localities. — Near  Port  Phillip  Heads  (Stations  1,  8,  14, 
and  outside  the  Heads,  coll.  J.  B.  Wilson)  ;  King  Island 
(coll.  Profe.ssor  Si)encer).  Ha:jckel  also  records  the  species 
from  Bass  Straits  and  the  Gulf  of  St.  Vincent. 

8.    Sf/con  setosum,  0.  Schmidt. 

Sycon  setosum,  0.  Schmidt.     Spong.  Adriat.  Meer.,  p.  15. 

I  identify  two  specimens  in  the  collection  with  this 
species.  They  differ  from  the  typical  8.  rwphanus  in  the 
more  hairy  surface,  due  to  the  greatei-  length  of  the  dermal 
oxea,  and  also  in  the  elongation  of  the  apical  rays  of  the 
gastral  quadriradiates.  Probably  it  is  merely  a.  slight 
variety  of  S.  raphanus.  The  species  has  hitherto  only  been 
recorded  from  the  Mediterranean. 

Locality. —  Near  Port  Phillip  Heads  (Stations  (J,  9,  coll. 
J.  B.  Wilson). 

9.   Sycon  ensiferum,  n.  sp. 

Solitary,  stipitate,  closely  resembling  S.  rapkanus ;  with 
very  markedly  conulose  surface  and  little  or  no  oscular  fringe. 
The  two  specimens  are  rather  larger  and  especially  stouter 
than  most  Australian  specimens  of  >S'.  raphanus  which  I 
have  seen. 

Canal-system  typical ;  chambers  of  good  length,  termi- 
nating in  low,  rounded  distal  cones. 

Skeleton  arranged  in  typical  manner.  The  species  is 
distinguished  by  the  following  characters  in  its  spiculation, 
which  in  general  characters  resembles  that  of  8.  ra/phanus 
closely  : — (1)  The  apical  rays  of  the  gastral  quadriradiates 
are  very  strongly  developed,  swelling  out  into  long  club- 
shaped  foi-m  (sword-shaped  in  longitudinal  section),  but 
fairly  sharply  pointed  and  only  very  slightly  curved,  very 
much  broader  in  the  distal  than  in  the  proximal  half. 
(2)  The  ordinary  tubar  radiates  very  frequently  have  a 
small  apical  lay  developed.  (8)  The  basal  rays  of  many  of 
the  most  distally  situated  tubar  triradiates  are  very  strongly 
bent  outwards  from  the  wall  of  the  chamber,  so  as  to  curve 
over  and  protect  the  entrances  to  the  inhalant  canals. 
(4)  The  oxea  at  the  distal  ends  of  the  chambers  are  of 
moderate  length  and  thickness,  straight  or  nearly  so, 
symmetrical  and  fairly  gradually  sharp-pointed  at  each  end. 

Locality. — Near  Port  Phillip  Heads  (Station  9,  coll. 
J.  B.  Wilson.) 


82        Proceedings  of  tJte  Ro/jal  Sucieiy  of  Victoria. 

10.    Sy con  .suhkispidum,  Cavtev,  s[). 

Grantia  siibldnpida,  Carter.  Annals  and  Magazine  of 
National  History,  July  1886,  p.  36. 

This  species,  described  by  Mr.  Carter  from  Mr.  Wilson's 
collection,  evidently  belongs  to  the  genus  Sycon,  but  I  have 
not  been  able  to  identify  any  of  my  specimens  therewith. 

Locality. — Near  Port  Phillip  Heads  (Carter). 

1 1 .   Sycon  rcmisayi,  von  Lendenfeld,  sp. 

Sycandra  ramsayi,  von  Lendenfeld.  Proc.  Linn.  Soc, 
N.S.W.,  Vol.  IX,  p.  1097. 

I  have  only  seen  specimens  of  this  species  from  Port 
Jackson.  Mr.  Carter,  however,  records  it  amongst  Mr. 
Wilson's  sponges  from  Port  Phillip.  The  gastral  cavity 
is,  according  to  my  experience,  widely  dilated,  and  not 
comparatively  small,  as  stated  in  the  original  desci-iption. 

Localities. — Port  Jackson  (von  Lendenfeld,  &c.)  ;  near 
Port  Phillip  Heads  (Carter). 

12.   Sycon  boomerang,'^  n.  sp. 

Solitary,  stipitate  ;  of  slightl}^  compressed,  ovoid  shape, 
tapering  gradually  below  to  form  the  narrow  stalk,  which  is 
about  equal  in  length  to  the  main  body  of  the  sponge  ;  with 
a  rathei-  small,  terminal,  naked  osculum.  Total  height  of 
the  specimen  about  37  mm.,  greatest  breadth  12-5  mm.  The 
dermal  surface  is  smooth  and  even,  but  with  a  characteristic 
porous  appearance.  The  wall  of  the  sponge  is  very  thick 
and  the  gastral  cavity  correspondingly  narrow. 

The  radial  chambers  are  very  long  and  nari-ow  and  branch 
re])eatedly,  the  branches  running  parallel  and  becoming- 
much  narrower  as  they  approach  the  dermal  surface.  The 
itdmlant  canals  are  correspondingly  long  and  narrow,  and 
their  outer  ends  are  covered  over  by  a  delicate  pore-bearing 
membrane  which  stretches  between  the  rounded  distal 
ends  of  the  chambers.  The  gastral  cortex  is  thin.  The 
skeleton  is  arranged  in  typical  manner,  the  spiculation 
being  as  follows  : — (1)  Gastral  quadriradiates  ;  sagittal  ; 
with  very  long,  slender,  hastate  basal  ray,  measuring 
about  0'2  X  0007  mm.,  sometimes  longer ;  paired  rays 
about  one-third  to  one-half  the  length  of  the  basal  ray 
and   somewhat  stouter,    straight,   conical,   gradually  sharp- 

*  So  called  from  the  shape  of  the  apical  raj's  of  the  gastral  quadriradiates. 


Si/noim6  of  the  Australian  Galcarei,  Heterocoela.    (So 

pointed ;  rti)ical  ray  very  strongly  developed,  gradually 
thickening  tor  about  two-thirds  of  its  length,  wliere 
it  is  extraordinarily  stout,  then  bending  shaiply  and 
tapering  more  suddenly  to  a  sharp  ])oint,  length  about 
O'loo  mm.,  greatest  tliickness  up  to  ()()28  mm.  though 
generally  less.  (2)  Gastral  triradiates ;  sagittal  or  suli- 
regular,  with  long,  slender,  gradually  sharj)-pointed  rays, 
the  oi-al  rays  often  somewLat  curved.  (3)  Subgastral 
sagittiil  triradiates,  not  clearh^  differentiated  from  tlie 
ordinaiy  tubar  spicules.  (4)  Tubar  triradiates  ;  with  very 
long,  straight,  slender,  conical  basal  ray  and  shorter,  widely 
■extended,  often  slightly  curved  oiul  rays.  In  spicules 
taken  from  about  the  middle  of  the  length  of  the  chamber 
the  basal  ray  measures  about  0"17  by  ()-07  mm.,  and  the 
-orals  about  ()-l  by  0'07  mm. ;  but  there  is  a  good  deal  of 
variation.  In  some  of  the  tubar  spicules  a  fairly  well 
developed  apical  ray  is  found.  In  some  of  the  most  distal 
triradiates  the  basal  ray,  now  much  shortened,  is  curved 
outAvards  so  as  to  lie  in  tlie  pore-bearing  membrane, 
which  is  also  supported  b\-  small,  scattered  tiiradiates  and 
oxeotes  like  those  found  at  the  distal  ends  of  the  chambeis. 
{5)  Oxea  of  the  distal  ccmes  ;  short  but  i'airly  stout,  more 
or  less  club-shaped,  usually  with  the  thick  distal  portion 
bent  at  an  angle  to  the  remainder;  measuiing  about  0()<S 
by  0-008  mm. ;  these  characteiistie  little  spicules  are 
arranged  in  dense  tufts  at  the  distal  ends  of  the  chambers. 
Locality.— ^e-Av  Port  Phillip  Heads  (coll.  J.  B.  Wilson). 

13.    Si/con  (jcldinosum,  Blainville,  sp. 

Alcijojicelluriif/elatino.sinn,  Blainville.  Actinologie,  p.  52!.). 

Sycandra  alcyoncellum,  Hjeckel.  Die  Kalkschwamme. 
Vol.  2,  p.  333. 

Sycandra  arhorca,  Ha^ckel.  Die  Kalkschv/anime,  Vol.  2. 
p.  331. 

This  common  Australian  species  is  veiy  variable  in  foi'uj, 
being  eithei-  colonial  (generally  richly  blanched)  or  solitary, 
with  the  oscula  sometiiues  naked  and  sometimes  provided 
with  a  fringe  of  .spicules.  The  shape  of  the  dermal  oxea  also 
varies  greatly,  from  club-  or  nail-shaped  to  sharply-pointe'd 
at  each  end.  The  extensive  series  of  specimens  in  my  col- 
lection, from  various  parts  of  Australia,  shows  that  it  is  quite 
impossible  to  separate  Hiieckel's  two  species,  arborea  and 
alcyoncellwni,  from  one  another,  and  I  revert  to  Blainvilie's 
■original  name,  gelatinosuni,  for  both. 

C4    2 


84-        Proceedimjs  of  lite  Royai  Society  of  Vicforia. 

Localities. — Port  Jackson ;  Port  Phillip ;  Bass  Straits;  St. 
Vincent's  Gulf;  west  coast  of  Australia  (various  authors  and 
collections  ;  Hseckel  also  records  the  species  from  Java). 

K3a.    Sycon  [jelatinosum  var.  whiteleggil,  nov. 

I  propose  to  distinguish  by  the  above  name  a  very 
beautiful  variety  of  the  foregoing  species  found  by  Mr.  T. 
Whitelegge  at  Watson's  Bay,  Port  Jackson.  There  are  nine 
specimens,  all  solitary  and  with  a  well-developed  oscular 
fringe  of  long  silky  spicules.  In  addition  to  this  oscular 
fiinge,  however,  all  have  a  beautiful  frill  or  collar  of 
long,  silky  spicules,  projecting  like  a  halo  from  the  base 
of  the  oscular  fringe  and  at  right  angles  to  the  long- 
axis  of  the  sponge.  In  external  form  this  variety  closely 
i'(>.sembles  Hpeckel's  beautiful  figure  of  S.  (SycaviuTYi) 
clegaDS.  The  dermal  oxea  are  long  and  slender,  and  gradu- 
ally sharp-pointed  at  each  end,  and  the  more  distal  tubar 
triradiates  are  greatly  enlarged,  with  long  and  stout,  but 
still  straight  basal  rays.  These  peculiarities  in  spiculation 
are,  however,  found  in  some  specimens  of  ;S^.  gelatiiwsum, 
horn  which  the  present  variety  cannot  be  sharply 
distinguished. 

Locality. — Watson's  Bay,  Port  Jackson  (coll.  T.  White- 
legge). 

14.    Sycon  glganteum,  n.  sp. 

Solitary,  with  very  short  stalk  or  none  at  all.  Tubular, 
greatly  elongated,  in  parts  more  or  less  compressed,  but  not 
varying  greatly  in  diameter  throughout  ;  with  a  single, 
wide,  naked  osculum.  Both  specimens  are  curved  or  bent. 
The  largest  is  nearly  100  mm.  in  length  by  14  mm.  in 
breadth ;  the  other  is  only  a  little  shorter.  The  wall  of  the 
sponge  is  about  3  mm.  in  thickness.  The  dermal  surftice  is 
in  part  quite  smooth  and  in  part  tessellated. 

The  radial  chambers  are  narrow  and  gi-eatly  elongated,  they 
branch  repeatedly  and  the  branches  run  parallel  with  one 
another  to  the  dermal  surface.  They  communicate  with  the 
gastral  cavity  by  long  exhalant  canals,  from  which  they  are 
separated  by  diaphragms.  These  exhalant  canals  appear 
like  continuations  of  the  chambers  only  without  collared 
cells,  they  may  unite  together  before  opening  on  the  gastral 
surfiice.  The  chambers  are  approximately  circular  in  trans- 
verse section.     The  inhalant  canals  are  irregular  and  very 


Syno2Jsis  of  the  Australian   Calcarea  Heteroc<ela.    85 

narrow,  o})ei\ing  on  the  dermal  surface  tlir(3Ugli  narrow, 
irregular  chinks  between  the  tufts  of  oxea. 

The  skeleton  is  arranged  as  usual,  the  spiculatioii  heing 
as  follows: — (1)  Gastral  quadriradiates;  small,  very  irregu- 
hirly  and  confusedly  arranged,  so  as  to  form  a  dense  though 
not  very  thick  cortex  ;  usually  more  or  less  strongly  sagittal, 
but  very  vai-iable  in  the  proportions  of  the  rays.  The  basal 
ray  varies  from  two  or  three  times  the  length  of  the  orals 
to  about  the  same  length  or  even  shorter  ;  it  is  straiglit  and 
conical.  The  oral  i-a^'s  are  usuallj'  slightly  curved  towards 
one  another,  conical  and  sharp-pointed,  averaging  about 
O'Oi  by  0005  mm.  at  the  base  ;  apical  I'ay  conical,  very 
slightly  curved,  sharply  pointed,  about  O'Oo  mm.  long. 
(2)  Subgastral  sagittal  spicules,  indistinguishable.  (3)  Tubar 
triradiates,  with  rather  short  and  stout,  conical,  sharp- 
pointed  rays  ;  the  oral  rays  very  widely  extended,  often 
nearly  at  right  angles  to  the  basal,  nearly  straight,  averag- 
iiig  about  0  084  mm.  by  0-0U9  mm.  at  the  base  ;  basal  ray 
varying  from  about  the  same  length  to  considerably  longer 
than  the  orals,  the  disproportion  being  greatest  at  the 
distal  ends  of  the  chambers.  (4)  Tubar  quadriradiates, 
differing  from  the  foregoing  in  tlie  development  of  a  sliort, 
stout,  curved  and  sharply  pointed  apical  ray  ;  abundant, 
especially  towards  the  gastral  surface,  where  the  tubar 
skeleton  becomes  very  irregular.  (.5)  Oxea,  short,  straight 
oi-  rather  crooked,  slender,  tapeiing  to  a  sharp  point  at  the 
proximal  end  and  with  the  di.stal  end  swollen  out  into  an 
ovoid  head,  like  that  of  a  spermatozoon,  length  about  Oiv, 
thickness  below  the  head  0007  mm.,  head  nearh^  twice  as 
tliick.  These  spicules  are  arranged  in  XQvy  dense  tufts  at 
the  dittal  ends  of  the  chambers  and  their  thickened  ends 
form  an  almost  continuous  crust  over  the  dermal  surface  of 
the  sponge.  The  whole  skeleton  is  \evy  dense  and  compact, 
so  that  the  texture  of  the  sponge  is  very  firm,  as  in 
S.  (jelatlnosuon,  which  it  closely  appi'oaches  in  spieulation. 

Locality. — St.  Vincent's  Gulf  (coll.  Adelaide  Museum). 

15.    Sycon  compres^iim,  auctorum. 

Graiitia  compvessa,  various  authors  {exj.,  Bowerbank). 

Sijcandra  compressa,  Hiieckel.  Die  Kalkschwamme,  Vol. 
11,  p.  3G0. 

This  common  European  species  is  recorded  from  Australia 
both    by   von    Lendenfeld    and    Carter,  but    1    have    never 


«0       P roceedlngs  of  the  Royal  Society  of  Victoria. 

myself  seen  specimens  from  Australian  seas.  Yon  Lenden- 
feld*  states  that  all  the  specimens  in  Australian  waters  are 
cylindrical  and  must  be  referred  to  Hseckel's  variety  lohata, 
which  he  j)i-oposes  to  erect  into  a  species  under  the  name 
(iraittki  lohata.  Carter-|- simply  records  Grantia  conprcfisa 
amongst  Mr.  Wilson's  sponges,  and  also  a  tubular  variety 
which  he  terms  fistulata,  and  which  is  probably  identical 
^^  ith  von  Lendenfeld's  lohata. 

T  include  the  species  in  the  genus  Sycon  on  account  of  the 
tufts  of  oxea  which  crown  the  radial  tubes.  The  dt-rmal 
cortex  is  ver}^  thin. 

Localities. — Port  Jackson  (von  Lendenfeld)  ;  near  Port 
Piiillip  Heads  (Carter). 

16.    Grantia  lahyrinthica,  Carter. 

TeiclioneUa  lahyrinthica,  Carter.  Annals  and  Magazine  of 
Natural  History,  July  1878,  p.  37. 

Gra)itia  lahyrinthica.  Carter.  Annals  and  Magazine  of 
Natural  History,  July  1880,  p.  38. 

I  have  already  given  a  detailed  account  of  the  history  and 
anatomy  of  this  remarkable  species  in  my  memoii'  "  C)n  the 
Anatomv  of  Grantia  lahijrintltica.  Carter,  and  tlie  so-called 
Family  Teichonidte,"  published  in  Vol.  XXXII,  N.S.,  of  the 
Quarttn-ly  Journal  of  Microscopical  Science.  The  species 
appears  to  l)e  fairly  common  near  Port  Phillip  Heads,  the 
largest  specimens  which  I  have  seen  measure  no  less  than 
live  inches  across  the  top,  a  truh'  gigantic  size  for  a  single 
Sycon  individual. 

Localities. — Fremantle,  W.A.  (Carter)  ;  near  Port  Philli}) 
Heads  (Station  5  and  outside  the  Heads,  Carter  and  coll. 
J.  B.  Wilson). 

17.    Grantia  extusarticulata,  Carter,  sp. 

Hypof/i'antia  extusarticulata.  Carter.  Annals  and  Maga- 
zine of  IS'atural  History,  July  1880,  p.  43. 

Solitary,  sessile,  sac-shaped,  broadest  below  and  tapei'ing- 
gradually  to  the  terminal  o.sculum,  which  is  naked.  The 
single  specimen  is  markedly  compressed,  and  measures  25 
nnn.  in  height  and  11  mm.  in  greatest  width.  The  wall  of 
t!ie  sac  is  not  much  more  than  1  mm.  in  thickness  and  the 
gastr.il    (^•lvity    is   correspondingly    spacious.      The    dermal 

*  Proc.  Linn.  Soc,  N.S.W.     Vol.  IX,  p.  1106. 
I  Ann.  and  Mag.  Nat.  Hist.     July  1886,  p.  37. 


Synopsis  of  the  Australian  Calcarea  Heievocada.    87 

SLirfoce  is  very  smooth.  The  anatc^uiy  is  very  typical.  The 
radial  chambers  are  straight,  cylindrical  and  only  slightly 
branched,  and  extend  from  gastral  to  deimal  coitex.  The 
inhalnnt  pores  are  irregularly  scattered  through  the  dermal 
cortex,  which  is  well  developed  and  about  007  mm.  thick. 
The  gastral  cortex  is  of  about  the  same  thickness  and  is 
perforated  by  the  short,  wide  exhalant  canals,  one  coming 
from  each  chamber  and  separated  from  it  by  a  constricted 
diaphragm. 

The  skeleton  is  arranged  in  typical  manner,  the  spicnlation 
being  as  follows  : — (1)  Gastral  quadriradiates  ;  sagittal,  oral 
angle  a  little  wider  than  the  laterals  ;  facial  rays  straight, 
conical,  graduall}'  sharp-pointed;  basal  ray  about  0-5  by  O'Ol 
mm.  ;  oral  rays  012  by  001  mm.  ;  apical  ray  short,  fairly 
stout,  only  moderately  sharply-pointed,  slightly  curved, 
about  0-06  mm.  long.  (2)  Gastral  triradiates  ;  similar  to  the 
foregoing,  luit  with  no  apical  ray.  (3)  Subgastral  sagittal 
trii-adiates ;  stronyly  developed,  with  widely  extended, 
slightly  recurved,  gradually  sharp-pointed  oral  rays  and  very 
long,  straight  basal  ray  gradually  tapering  to  a  sharp  point; 
oral  rays  about  Oi5  by  0-014  mm.  ;  basal  ray  about  0-35  by 
0-0 1 4"  mm.  (4)  Tubar  triradiates ;  somewhat  smaller  than  the 
foregoing  but  well  developed,  with  straight  or  nearly  straight 
rays,  gradually  sharp-pointed,  the  basal  considerably  longer 
than  the  other  two.  (5)  Dermal  tri'-adiates  ;  sagittal,  very 
similar  to  the  tubar  tru-adiates  but  perhaps  a  little  longer 
and  placed  horizontally  in  the  dermal  cortex.  (6)  Oxea  of 
the  dermal  cortex  ;  very  small,  straight,  graduall^y  sharp- 
pointed  at  the  inner  end  and  beautitully  hastate  or  lance- 
pointed  at  the  outer  ;  about  0-04.3  by  O'OOS  mm. ;  arranged 
at  right  angles  to  the  dermal  surface.  Occasionally  a  large 
oxeote  s[)icule  is  found  around  the  margin  of  the  osculum, 
but  these  are  extremely  rare. 

Mr.  Carter's  specimen,  described  from  Mr.  Wilson's  collec- 
tion, was  "  agglomerated."  I  have  little  doubt  as  to  the 
specific  identity  of  the  two,  but  there  are  sufficient  points  of 
distinction  between  my  specimen  and  Mr.  Carter's  original 
description  to  render  a  fresh  description  desij-able. 

Locality. — Near  Ft.  Phillip  Heads  (Carter;  and  Station  9, 
coll.  J.  B.  Wilson). 

1 8.    Granfia  gracilis,  von  Lendenfeld,  sp. 
Vosniaeria  gracilis,  von  Lendenfeld.     Proceedings  of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  ]>.  1111. 


88       Froceediiuj^  of  the  Royal  Society  of  Victoria. 

In  canal  system,  so  far  as  we  can  judge  tVom  the  author's 
description,  this  species  appears  to  ieseml)le  my  Grantia 
vosmaeri,  the  radial  chambers  communicating  with  the 
gastral  cavity  by  elongated  exhalant  canals. 

Locality. — Port  Jackson  (von  Lendenfeld). 

19.    Grantia  votsmaerl,  n.  sp. 

Specimen  solitary,  sessile  (?),  sac-shaped,  tapering  gradually 
above  to  the  naked,  terminal  osculum  ;    15  mm.  high  and 

7  mm.  in  greatest  transverse  diameter.  Texture  hard, 
dermal  surface  echinated  by  the  large,  projecting  oxea. 
Wall  of  sac  only  about  1  mm.  thick. 

The    dermal    cortex   is    very   strongly    developed,    about 

008  mm.  thick  ;  the  gastral  cortex  is  two  or  three  times 
as  thick,  but  less  dense  and  not  so  well-defined.  The  radial 
chambers  are  rather  short  and  moie  or  less  branched.  Their 
distal  ends  abut  against  the  dermal  cortex,  while  proximally 
they  communicate  with  the  gastral  cavity  by  means  of  long, 
wide,  exhalant  canals,  which  penetrate  the  gastral  cortex 
and  may  unite  together  before  opening  onto  the  gastral 
surface.  The  chambers  are  sepai-ated  trom  the  exhalant 
canals  by  constricted  diaphragms.  The  inhalant  pores  take 
the  form  of  irregular  canals  penetrating  the  dermal  cortex. 

Except  for  the  unusual  thickness  of  the  cortex  and  the 
great  size  of  the  oxeote  spicules  the  skeleton  is  ari'anged 
in  the  ordinary  manner.     The  spiculation  is  as  follows  : — 

(1)  Gastral  quadriradiates  ;  usually  more  or  less  sagittal, 
but  very  variable ;  rays  straight  or  paired  raj's  slightly 
curved,  stout,  conical  and  gradually  sharp-pointed ;  size 
very  variable  ;  paired  rays  averaging,  say,  about  0  2  by 
0-02.5  mm.  ;  basal  ray  about  the  same  thickness  and  usually 
somewhat  longer  ;  a})ical  ray  thorn-like,  short,  stout,  conical, 
u.sually  slightly  curved,  finely  pointed,  about  0"07  mm.  long, 
projecting    into    the    gastral    cavity    and    exhalant    canals. 

(2)  Gastral  triradiates ;  usually  like  tlie  foregoing  but 
without  the  apical  ray ;  towards  the  osculum,  however, 
they  become  much  more  strongly  and  regularly'  sagittal, 
the  widespread,  slightly  recurved  oral  rays  being  much 
longer  and  stouter  than  the  basal  ray ;  amongst  these 
suboscular  spicules  quadriradiates  are  very  rare.  (3)  8ub- 
gastral  sagittal  trii-adiates ;  indistinguishable  from  the 
remainder  of  the  tubar  skeleton,  which  merges  somewhat 
gradually  into  the  gastral  cortex.      (4)    Tubar  trii-adiates  ; 


Syyiopsis  of  the  AudraJUui  Calcarea  Heterocmla.    89 

strongly  sagittal ;  with  widely  extended,  short,  straight, 
conical  and  gradually  sharp-pointed  oral  rays,  and  much 
longer,  straight,  gradually  sharp-pointed  basal  ray;  oral 
rays  about  Ol  by  0-()]5  mm.;  basal  ray  about  0-28  by 
0-0 15  mm.  (5)  Triradiates  of  the  dermal  cortex;  usually 
slightly  sagittal  but  uearly  regular  ;  rays  stout,  straight 
or  slightly  curved,  conical,  gradually  sharp-pointed  ;  about 
0"24?  by  0  028  mm.  ;  arranged  in  several  layers  parallel 
with  the  dermal  surface.  (6)  Oxea ;  very  lai-ge,  stout, 
fusiform,  usually  gently  curved,  gradually  sharp-pointed  at 
each  end;  varying  in  size  up  to  about  1'8  by  007  mm. 
The  outer  ends  of  these  spicules  project  far  teyond  the 
dermal  surface,  while  their  inner  ends  extend  through  the 
chamber  layer  into  the  gastral  cortex. 

Locality. — Watson's  Bay, Port  Jackson  (coll.  T.  Whitelegge). 

20.  Grantia  (.^)  infreque^ni.  Carter,  sp. 

Ilypogmntia  infrequens.  Carter.  Annals  and  Magazine 
of  Natural  History,  July  ]886,  p.  :39. 

The  chief  peculiarities  of  this  species  appear  to  be  the 
presence  of  a  very  strong  dermal  cortex  and  tlie  fact  that 
the  tubar  skeleton  is  made  up  entirely  of  the  V)asal  itxya  of 
subgastral  sagittal  triradiates.  I  have  not  seen  the  species 
myself,  and  Mi'.  Carter  apparently  had  only  a  single  small 
specimen,  collected  by  Mr.  Wilson.  I  include  it  provisioiially 
in  the  genus  Grantia. 

Locality. — Near  Port  Phillip  Heads  (Cartel-). 

21.  Grantia  (})  IcBvigata,  Hteckel,  sp. 

Sycortii^  kevigata,  Hseckel.  Die  Kalkschwamme,  Vol.  II, 
p.  285. 

In  his  description  Hseckel  distinctly  states  that  in  the 
case  of  the  tabar  trii-adiates  the  basal  ray  is  always  directed 
centrifugally  outwards,  while  in  the  dermal  trii-adiates  it 
lies  parallel  to  the  long  axis  of  the  sponge  and  points  to  the 
aboral  pole.  No  mention  is  made  in  the  text  of  any 
subdermal  sagittal  triradiates  with  inwardly  directed  basal 
ray,  but  in  the  figure  (Plate  49,  Fig.  3)  such  appear  to  be 
present.  Hence  if  the  figure  be  correct  we  should  have  to 
place  the  species  in  the  genus  Grayitessa,  but  the  evidence  is 
hardly  strong  enough  to  justify  us  in  so  doing  at  present. 

Locality. — Gulf  of  St.  Vincent  (Hseckel). 


90       ProceediiKj.s  of  the  Royal  Society  of  Victoria. 

22.    GrayitiopHis  cyliiuirica,  n.  sp. 

Sponge  forming  long,  cylindrical  tubes,  which  may  branch, 
with  single,  terminal,  slightly  constricted,  almost  naked 
oscula.  Sui'face  not  hispid  but  slightly  roughened  by  the 
large  triradiate  spicules  of  the  cortex.  Tlie  largest  tube  in 
the  collection  is  unbranched  and  slightly  crooked,  57  mm. 
long  and  with  a  nearly  uniform  diameter  of  5  mm. 

'i'he  wall  of  the  tube  is  about  1  mm.  in  thickness,  and  is 
divided  into  two  sharply  defined  concentric  laj^ers  of  about 
equal  thickness.  The  outer  of  these  layers  forms  a  firm 
cortex  with  a  very  strongly  developed  skeleton.  ,  The  inner 
la3'er  is  soft  and  spongy,  consisting  almost  entirely  of  the 
radial  chambers,  which  have  but  a  feebly  developed  tubar 
skeleton. 

The  iidialant  pores,  scattered  in  groups  over  the  dermal 
surface,  lead  into  veiy  sharply  defined,  wide  inhalant  canals, 
which  penetrate  the  cortex,  uniting  into  larger  trunks  which 
conduct  the  water  to  the  ordinary'  "  intercanals"  between 
the  radial  chambers. 

The  radial  chambers  ai'e  ari-anged  side  by  side  with  great 
regulaiity.  Each  is  a  straight,  wide,  unbranched  (or  very 
sliglitly  blanched)  tube,  extending  completely  through  the 
chain] )er  layer.  In  cross  section  they  vary  from  nearly 
square  to  neai-l_y  circular.  Each  o])ens  directly  and  separately 
into  the  gastral  cavity,  the  gastral  cortex  being  so  tliin  tliat 
no  special  exhalant  canals  are  required.  Kacli  is  provided 
at  its  i)roximal  end  with  a  membranous  diaphragm,  which, 
in  the  s]nrit  specimen,  almost  closes  the  Oj)eniiig. 

The  arraiigement  of  the  skeleton  is  a  slight  modilication 
of  t!)e  Gravtia  type,  but  the  spiculation  is  xevy  peculiar,  as 
follows: — (1)  Gastral  quadriradiates  ;  sliiilitly  sagittal,  with 
small  facial  and  enormous  apical  rays  ;  the  oral  angle  is  rather 
wider  than  die  lateral  angles,  but  tliere  is  not  much  difference 
in  the  length  of  the  facial  ra_ys,  which  are  nearly  straight 
(the  orals  may  be  sliuhtl}'  curved),  fairly  stout,  conical  and 
graduallj'  sharp-pointed,  about  0-056  by  O'OOT  mm.  The 
apical  ray  is  slightly  curved,  very  .stout,  sword-shaped  in 
optical  section,  thickest  hi  about  the  middle,  gradually 
sharp-pointed,  about  014  by  001 4  mm.  These  apical 
rays  thickly  echinate  the  gastral  surface.  (2)  Subgastral 
quadriradiates;  strongly  sagittal;  the  oral  rays  very  widely 
extended  and  parallel  to  the  gastral  surface  ;  the  basal  ray 
extending  centrifugally  towards  the  dermal  surface  through 


Synojisi'^  of  the  Australian  Galcarea  Heterocoela.    91 

about  half  the  thickness  of  the  chamber  layer  ;  the  apical  ra\' 
projecting  into  the  gastral  cavity,  almost  in  a  line  with  the 
basal  ray.  The  basal  ray  is  long,  straight,  and  gradually 
sharp-pointed,  about  ()"28  by  O'Ol  mm.  ;  the  oral  rays  are 
short,  straight,  conical  and  gradually  sharp-pointed,  about 
0*056  by  0008  mm. ;  the  apical  ray  is  slender,  conical, 
elongated,  slightly  curved  and  finely  pointed,  about  0O9  by 
0-007  mm.  (3)  Tubar  triradiates  ;  consisting  almost  entirely 
of  the  strongly  developed,  centrifugaliy  directed  basal  ray^ 
which  is  straight,  fusiform,  gradually  sharp-pointed  at  the 
distal  end,  and  at  the  proximal  end  provided  with  a  pair  of 
minute,  widely  divergent,  conical  teeth,  which  represent  the 
extremely  reduced  oral  rays.  The  basal  ray  measures  about 
0-3  by  O'OOS  mm.,  while  the  teeth  representing  the  oral  rays 
are  only  about  0  003  mm.  in  length.  The  entire  tubar 
skeleton  is  made  up  of  these  spicules  and  the  basal  rays  of 
the  subgastral  quadriradiates,  arranged  usually  in  single 
series  but  with  overlapping  ends,  each  series  comprising  only 
about  three  spicules.  (4)  Triradiates  of  the  dermal  cortex  ; 
very  lai'ge  and  stout  and  regularly  sagittal,  the  oral  angle 
being  very  considei-ably  wider  than  the  paired  angles  ;  all 
the  rays  are  straight,  conical  and  gradually  sharp-pointed, 
the  oral  rays  measuring  about  O'o-t  by  0-07  mm.,  and  the 
basal  ray  somewhat  shorter  and  slenderer.  These  spicules 
are  arranged  in  many  layers,  parallel  to  the  dermal,  surface 
and  extending  through  the  entire  thickness  of  the  cortex. 

(5)  Dermal  oxea ;  short,  slender,  slightly  crooked,  sharply 
pointed  at  each  end,  about  01  by  O'OOO  mm.  These  spicules 
occur  in  large  numbers  on  the  dermal  surface,  projecting  at 
right   angles    from    the    outermost    portion    of    the    cortex. 

(6)  Oxea  of  the  oscular  fringe  ;  long,  straight  and  slender, 
gradually  sharp-pointed  at  each  end  though  hastate  at  the 
outer  end.  These  spicules  form  a  kind  of  vertical  palisade 
inside  the  margin  of  the  osculuin,  their  outer  ends  project- 
ing to  form  a  feebly  developed  oscular  fringe.  (7)  Oscular 
triradiates  :  closely  resembling  the  remarkable  tubar  tri- 
radiates, but  A\-ith  the  oral  rays  not  quite  so  much  reduced. 
These  spicules  occur  mixed  with  the  inserted  portions  of  the 
oscular  oxea,  and  assist  the  latter  in  forming  a  dense 
]:»alisade  ;  the  paired  raj'S  are  directed  towards  the  osculum, 
and  no  part  of  the  spicule  projects  freely  like  the  oxea  do. 

This  sponge  is  decidedl}^  the  gem  of  Mr.  Wilson's  collection. 
The  exquisite  symmetry  of  the  skeleton  and  canal-system, 
combined  with  the  remai-kable  spiculation,  render  it  one  of 


02       Proceedings  of  the  Royal  Society  of  Victoria. 

the  most  beautiful  and  interesting  sponges  which  I  have 
ever  seen,  although  the  external  t'oim  is  not  particularly 
attractive. 

Locality.— ^eav  Port  Phillip  Heads  (coll.  J.  B.  Wilson). 

23.    Ute  syconoides,  Carter,  sp. 

Aphroceras  syconoides,  Carter.  Annals  and  Magazine  of 
Natural  History,  August  1886,  p.  135. 

I  identify  with  this  species  a  single  specimen  collected  by 
Mr.  Wilson  and  a  number  of  very  beautiful  examples  sent 
to  me  from  Port  Jackson  by  Mr.  T.  Whiteiegge.  As  pointed 
out  by  Mr.  Carter,  the  species  closely  resembles  Schmidt's 
Ute  glabra,  having  the  same  characteristic  silvery  sheen  on 
the  surface,  due  to  the  presence  of  the  huge,  longitudinally 
disposed  oxea.  The  Port  Jackson  specimens  are  shortly 
stipitate  and  one  of  them  consists  of  two  individuals  united 
below  for  about  half  their  length,  or  one  might  regard  it  as 
a  branched  individual. 

Localities. — Near  Port  Phillip  Heads  (Carter,  and  Station 
14,  coll.  J.  B.  Wilson)  ;  Watson's  Bay,  Port  Jackson  (coll. 
T.  Whiteiegge). 

24.    Ute  argentea,  Polejaeff. 

Ute  argentea,  Poldjaeff.  Report  on  the  Calcarea  of  the 
Challenger  Expedition,  p.  43. 

This  species  is  also  very  similar  to  Schmidt's  Ute 
fjlabra.  Although  the  skeleton  is,  as  PoJejaefF  points  out, 
"  inarticulate,"  there  are  no  subdermal  sagittal  triradiates 
with  inwardly  directed  basal  ray.  From  personal  examina- 
tion of  Polejaeff 's  type  I  believe  this  species  to  be  quite 
distinct  from  Ute  syconoides,  the  latter  differing,  amongst 
other  things,  in  its  much  longer  radial  chambers,  with  many- 
jointed  tubar  skeleton,  and  in  the  much  slenderer  and  less 
densely  packed  longitudinal  oxea. 

Locality.— Oft'  Twofold  Bay  (Polejaeff). 

25.    Ute  ."ipicidosa,  n.  sp. 

Sponge  colonial,  consisting  of  a  number  of  individuals 
(about  twent}^  in  the  specimen  under  examination)  united 
together  by  their  bases  so  as  to  form  a  spreading  colony. 
The  individuals  composing  the  colony  are  sessile,  ovoid, 
narrowing  above  to  the  small  terminal  osculum,  which  has 


S)jiiopsi's  of  the  Aadrailau  Oalotrea  Heterucwla.    93 

a  very  inconspicuous  fringe  ;  they  attain  a  height  of  about 
15  mm.  and  a  maximum  diameter  of  about  5  mm.  The 
texture  is  dense  and  firm,  and  the  sui-face  is  roughened 
by  the  projecting  ends  of  some  of  the  large  oxea. 

The  gastral  cavity  is  narrow  and  cylindrica],  occupying 
only  about  one-third  of  the  total  diameter  of  the  sjionge. 
The  flagelUited  chambers  are  long  and  naiTow  and  more  or 
less  radially  arranged  with  regard  to  the  central  gastral 
cavity  ;  they  do  not  extend  nearly  through  the  entire  thick- 
ness of  the  sponge  wall,  and  they  communicate  with  tlie 
gastral  cavity  through  long,  sometimes  branched  exhalant 
canals.  The  inhalant  canal  system  consists  of  scattered 
pores  on  the  dermal  surface  leading  into  elongated  canals 
which  lead  down  between  the  chambers,  but  the  typical 
syconoid  arrangement  of  the  canal  system  is  greatly 
obscured  by  the  strong  development  of  the  mesoderm 
and  the  dense,  irregular  skeleton.  There  is  a  ver}-  thick, 
dense  cortex  on  both  dermal  and  gastral  surfaces. 

The  skeleton  of  the  gastral  cortex  consists  of  a  densely 
felted  mass  of  irregularly  arranged  triradiates,  mostly  lying- 
parallel  to  the  gastral  surface.  These  spicules  are  sagittal, 
with  fairly  stout,  sti-aight,  conical  and  graduall}'  shar})- 
pointed  rays  ;  the  oral  rays  are  longer  than  the  basal  and 
the  oral  angle  wider  than  the  other  two  ;  oral  ra3'S  about 
()-lS  mm.  by  002  mm.,  basal  about  012  mm.  by  0010  mm. 
The  skeleton  of  the  chamber  layer  is  dense  and  irregular, 
but  shows  traces  of  the  articulate  tubar  arrangement  in  the 
usually  centrifugal  direction  of  the  basal  rays  of  the  tri- 
radiates. These  spicules  are  smaller  than  those  of  the 
gastral  cortex,  and  of  different  shape ;  there  is  not  much 
difference  in  the  length  of  the  rays,  though  the  basal  may 
be  slightly  longer  or  shorter  than  the  others  ;  all  the  rays 
are  fairly  stout,  conical  and  gradually  sharp-pointed  ;  the 
basal  is  straight  but  the  orals  are  more  or  less  curved 
towards  one  another;  dimensions  of  rays  about  012  by 
0-OlGmm. 

The  skeleton  of  the  dei-mal  cortex  consists  of  a  dense,, 
confused  mass  of  triradiates,  resembling  those  of  the 
chamber  layer  but  becoming  markedly  smaller  towards 
the  outside,  where  they  lie  parallel  to  the  surface ;  amongst 
which  are  found  oxea  of  two  kinds : — (1)  Very  large,  stout, 
fusiform,  slightly  curved  and  sharply  pointed  at  each  end  ; 
measuring  about  18  mm.  by  O'l  mm.,  and  arranged  parallel 
to  the  long  axis  of  the  sponge,  with  the  upper  end  often 


i)!         I'nn'ccdhnjH  of  I  la:    Hoijal   Sociel//  of   Victoria. 

slightly  projc'ctJnii-.  (2)  Sinn]],  lung  and  slender,  neaih' 
straight,  gradually  sharp-pointed  at  the  inner  end  and 
usually  more  or  less  hastate  or  lance-pointed  at  tlie  outei- ; 
measui'ing  about  ()-24  hy  0"008  mu).  These  spicules  occur 
in  the  outermost  portion  of  the  cortex,  and  their  outer  ends 
])roject  well  beyond  and  more  or  less  at  right  angles  to  the 
dermal  surface.  A  number  of  similar  but  longer  spicules 
insei'ted  around  tlie  inside  of  the  osculum  form  a  dense  but 
not  pi-ominent  oscular  fringe. 

/>of«?//v/.— Watson's  Bay,  Pt.  Jackson  (coll.  T.  Whitelegge). 

•20.    Ufe  spenceri,  n.  sp. 

Sponge  solitary,  sessile,  globular  or  sub-sphericai,  with 
correspondingly  dilated  gastral  cavity  and  nari'ow,  naked 
osculum.  The  texture  is  lirm  and  harsh  to  the  touch,  the 
dermal  surface  being  rather  uneven  and  slightly  i-oughened 
by  the  projecting  apices  of  some  of  the  large  oxea,  but  not 
hispid.  Diameter  of  entire  sponge  about  11  mm.;  thickness 
of  wall  about  2o  ujm.  Tlie  dermal  cortex  is  very  thick, 
occupying  more  than  one-third  of  the  entire  thickness  of  the 
wall. 

The  inhalant  pores,  scattered  over  the  surface  of  the 
sponge,  lead  into  wide,  ii-regulai-,  sub-dermal  cavities,  lying 
in  the  coitex,  from  which  narrow  inhalant  canals  lead  down 
between  the  radial  chambers.  The  radial  chambers  aie 
arranged  w^ith  considerable  I'egularity  parallel  to  one  anothei". 
They  are  long  and  narrow^  (about  1  •()  mm.  by  ()"14  mm.), 
and  at  their  distal  ends  they  branch  in  a  curiously  iri'egular 
manner,  the  branches  scmietimes  penetrating  for  some  little 
distance  into  the  dermal  cortex.  The  proximal  ends  of  the 
chambers  are  all  situate  about  at  the  same  level,  which  is 
some  little  distance  from  the  gastral  cavity  and  even  fron; 
the  gastral  cortex,  which  latter,  though  dense,  is  very  thin 
as  compared  with  the  dermal  cortex.  Hence  we  find  a 
number  of  rather  short,  cylindrical,  radially-arranged 
exhalant  canals,  which  look  like  continuations  of  the  i-adial 
chambers  without  the  collared  cells,  and  which  may  unite 
together  in  groups  before  opening  on  the  gastral  surface. 
The  I'adial  chambers  are  separated  from  the  exhalant  canals 
by  well-marked  diaphragms. 

The  .skeleton  is  very  dense  and  very  complicated  and 
consists  of  the  following  parts: — (1)  Quadriradiates  of  the 
gastral  cortex ;  sagittal,  with  straight,  conical,  gradually 
sharji-pointed  facial  ra3's  ;  the  oral  angle  is  wider  than  the 


Si/nopsls  of  the  Australian  Calcarea  Heterocosla.    95 

paired  angles  and  the  basal  ray  may  be  either  longer  or 
shorter  than  the  other  two,  which  measure,  say,  abont  0"09 
by  ()0()8  mm.  ;  tlie  apical  ray  is  well  developed,  conical, 
gradually  sharp-pointed,  slightly  curved,  and  nearly  as  long 
and  thick  as  the  oral  ray.s.  These  spicules  form  such  a 
dense  feltwork  that  it  is  difficult  to  make  out  the  details  of  in- 
dividual form  in  situ,  while  the  projecting  apical  rajs  thickly 
echinate  the  gastral  cavity.  {'2)  Quadritadiates  of  the  exha- 
lant  canals  ;  these  are  extremely  characteristic  and  peculiar 
spicules ;  the  basal  ray  is  reduced  to  a  mere  rounded 
tubercle,  while  the  oral  and  apical  rays  are  long,  straight 
and  very  slender,  and  finely  pointed ;  the  oral  rays  diverge 
at  an  angle  of  about  120"  and  the  apical  comes  off  between 
them  and  appears  to  lie  nearly  in  the  same  plane  ;  the 
oral  rays  measure  about  008  by  0  0027  mm.,  though 
occasionally  stoutei*,  and  the  apical  ray  is  about  one-third 
as  long ;  these  spicules  ai'e  found  around  the  exhalant  canals, 
with  the  apical  ray  projecting  into  the  cavity.  A  few  lai-ger 
and  stouter  quadni-adiates,  with  normal  basal  ray,  also 
occur  around  the  exhalant  canals.  (3)  Inner  sagittal  tri- 
radiates ;  under  this  term  we  may  pei'haps,  in  this  species, 
include  all  those  triradiates  which  lie  in  the  zone  between 
the  gastral  cortex  and  the  commencement  of  the  flagellated 
■cliambers,  although  they  lie  at  varying  depths  beneath  the 
gastral  cortex.  The  oral  rays  are  straight  or  nearly  so,  conical 
and  gradually  sharp-pointed,  about  0-09  by  0-0085  mm.;  the 
basal  ray  is  long,  straight,  conical  and  gradually  sharp- 
pointed,  measuring  about  0- 1 6  by  0-0085  mm. ;  the  oral  angle 
is  wider  than  the  paired  angles.  (4)  Tubar  triradiates  ; 
very  similar  to  the  foregoing  but  the  basal  r;iy  gradually 
diminishes  in  length  towards  the  dermal  cortex.  These 
spicules  form  an  articulate  tubar  skeleton  of  many  joints, 
which  is  continued,  as  already  indicated,  within  the  inner 
limits  of  the  chamber  layer  to  the  gastral  cortex,  while 
towards  the  outside  it  becomes  irregular  and  gradually 
passes  into  the  skeleton  of  tiie  dermal  cortex.  (5)  Triradiates 
of  the  dermal  cortex ;  slightly  sagittal  or  sub-regular, 
mostly  larger  and  stouter  than  the  tubar  triradiates,  with 
conical,  sharp-pointed  rays  measuring  about  O'l  6  by  0-02  mm., 
but  very  variable ;  towards  the  outside  they  lie  parallel  to 
the  dei-mal  surface,  but  otherwise  they  are  very  irregularly 
arranged.  (6)  Large  oxea  of  the  dermal  cortex  ;  fusiform, 
slightly  curved,  gradually  sharp-pointed  at  each  end  ; 
measuring  about  1-4   by  O'l   mm.,   but  sometimes  more  or 


90       Froceeclin<i6  of  the  Royal  Society  of  Victoria. 

less.  These  spicules  are  imbedded  in  large  numbers  in  the 
dermal  cortex  at  various  levels  ;  they  mostly  lie  more  or 
less  parallel  to  the  long  axis  of  the  sponge,  but  there  is  a 
good  deal  of  irregularity  in  their  arrangement  and  not 
infrequentl}"  one  end  of  the  spicule  projects  slightlj'-  beyond 
the  dermal  surface.  (7)  Minute  oxea  of  the  dermal  surface  ; 
short  and  slender,  usually  slightly  curved  ;  the  inner  end 
gradually  tapering  to  a  fine  point,  the  outer  end  thicker, 
more  or  less  hastate,  minutely  toothed  or  roughened.  These 
spicules  measure  only  about  004  by  0'003  mm.;  they  occur 
in  large  numbers  on  the  dermal  surface.  (S)  Minute  oxea 
of  tlie  gastral  surface  ;  similar  to  the  foregoing  but  not  so 
numerous. 

One  of  the  two  specimens  in  the  collection  was  attached 
to  a  crab's  back,  which  it  completely  covered  like  a  thick 
crust ;  it  resembled  a  specimen  cut  in  half  longitudinally, 
with  the  concave  gastral  surface  turned  towards  the  crab's 
back.  Hence,  as  the  gastral  cavity  was  no  longer  an 
enclosed  space,  there  was  no  oscuium  in  the  ordinary  sense 
of  the  word.  The  crab,  of  course,  occupied  the  gastral 
cavity,  and  the  exhalant  canals  of  the  flagellated  chambers 
must  have  discharged  their  contents  on  to  the  crab's  back. 
One  often  finds  sponges  growing  on  crab's  backs,  but  I 
never  before  saw  a  case  in  which  the  essential  torm  of  the 
sponge  was  so  strangely  modified  in  accordance  with  this 
habit.  Had  it  not  been  for  the  presence  of  the  other  and 
normal  specimen  in  the  collection  I  should  have  been  inclined 
to  regard  this  strange  modification  in  form  as  of  at  least 
specific  value.  The  species,  is,  however,  so  well  characterised 
by  spiculation,  &c.,  that  there  can  be  no  doubt  as  to  the 
identity  of  the  two  specimens. 

I  have  much  pleasure  in  dedicating  this  remarkable  species 
to  Professor  W.  Baldwin  Spencer. 

Locality. — Watson's  Bay,  Port  Jackson  (coll.T.  Whitelegge). 

27.   Synute  pulchella,  Dendy. 

Synute  'pulchella,  Dendy.  Proceedings  of  the  Royal 
Society  of  Victoria,  Vol.  IV  (New  Series),  p.  1. 

I  have  nothing  to  add  to  my  description  of  this  remark- 
able sponge  until  such  time  as  I  ma}^  be  able  to  publish 
illustrations  of  its  anatomy. 

Locality. — Near  Port  Phillip  Heads  (Dendy). 


Si/iiojJ'sis  of  f/tc   A  a,>ifn(lla)i  Cdlcarea  Heterocfelit.    97 

28.    Anainixilla  torresi,  PolejaefF. 

Aiiainixilla  torresi,  Polejaeff.  Report  on  the  Calcarea  of 
the  Challenoer  Expedition,  p.  50. 

I  liave  seen  no  specimen  of  this  s[)onge  except  a  portion 
of  the  type  from  the  British  Museum. 

Locality. — Torres  Straits  (Polejaeff). 

29.    Leucandra  australiensis,  Cartel-,  sp. 

Leuconia  Jistulosa,  var.  australiensis,  Carter.  Annals 
and  Magazine  of  Natural  History,  August  J  886,  p.  127. 

There  is  one  specimen  in  the  collection,  belonging  to  the 
National  Museum,  which  closely  resembles  in  external 
characters  and  spiculation  that  described  by  Mr.  Carter. 
The  sleiiclerness  of  the  radiate  spicules  gives  to  the  sponge 
a  soft  and  yielding  texture,  while  the  dermal  surface  is 
densely  hispid  from  the  long,  slender,  projecting  oxea.  At 
first  sight  the  specimen  looks  like  a  large  example  of 
Grantessa  hirsuta,  but  it  differs  markedly  in  the  arrange- 
ment of  the  canal  system  and  in  the  absence  of  the 
subderinal  sagittal  triradiates.  The  chambers  are  large  and 
irregularly  sac-shaped,  averaging  say  about  0'3  by  0"1  mm. 
(but  very  variable)  ;  not  arranged  radially  around  the 
central  gastral  cavity  of  the  sponge,  but  around  wide 
exhalant  canals  which  penetrate  the  thickness  of  the  wall  of 
the  sponge  and  are,  like  the  gastral  cavity  itself,  echinated 
by  the  apical  ravs  of  quadriradiate  spicules. 

Locality. — Near  Port  Phillip  Heads  (Carter,  and  Station  1 4, 
coll.  J.  B.  Wilson).- 

30.  Leucandra  alcicornis,  Gray,  sp. 

Aphroceyas  alcicornis,  Gray.  Proceedings  of  the  Zoolog- 
ical Society  of  London,  1858,  p.  114. 

Leucandra  alcicornis,  Hseckel.  Die  Kalkschwamme, 
Vol.  II,  p.  184. 

I  have  not  j^et  had  an  opportunity  of  examining  this 
widely  distributed  and  very  remarkable  species. 

Loccdity. — -Bass  Straits  (Hteckel).  Also  recorded  from 
various  localities  in  the  Pacific  and  Indian  Oceans  and  from 
the  Cape  (^vide  Hreckel). 

31.  Leucandra  catcqjhracta,  Hseckel. 

Leucandra  cataphracta,  Hseckel.  Die  Kalkschwamme, 
Vol.  II,  p.  203. 


i)8        Proceedings  of  ike   Roijal  Society  of  Victoria. 

Leucandra  catcqjhracta,  von  Lendenfeld.  Proceedings  ot" 
the  Liiiiiean  Society  of  New  South  Wales,  Vol.  IX,  p.  1129. 

I  am  indebted  to  Mr.  T.  Whitelegge  for  a  considerabh; 
number  of  tine  specimens  of  this  sponge  from  Watson's  Bay, 
Port  Jackson.  Neither  Hteckel  nor  von  Lendenfeld  have 
described  the  flagellated  chambers,  which  are  small,  appioxi- 
matel}^  spherical  and  scattered  abundantly  in  the  thick 
wall ;  measuring  about  0  09  mm.  in  diameter. 

Loc(dities. — Port  Jackson  (Heeckel,  von  Lendenfeld,  &c.)  ; 
Port  Denison  (von  Lendenfeld). 

32.    Leucandra  typlca,  Polejaeff,  sp. 

Leuconia  typica,  Polejaeff.  Report  on  the  Calcarea  of 
the  Challenger  Expedition,  p.  56. 

Leucandra  typica,  von  Lendenfeld.  Proceeding  of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1130. 

Locality. — Port  Jackson  (von  Lendenfeld.  Recorded  by 
Polejaeff  from  the  Bermuda  Islands). 

33.    Leucandra  "tneandrina,  von  Lendenfeld. 

Leucandra  meandrina,  von  Lendenfeld.  Proceedings  of 
the  Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1128. 

I  identify  with  this  species  a  somewhat  mas.sive  but  not 
large  specimen  collected  by  Mr.  Wilson,  which  seems  to 
agree  closely  with  a  fragment  of  the  type  from  the  British 
Museum,  but  the  species  is  by  no  means  an  easy  one  to 
characterise. 

The  chambers  are  approximately  spherical  and  about 
009  mm.  in  diameter. 

Localitieti.—  '^OYt  Jackson  (von  Lendenfeld)  ;  near  Port 
Phillip  Heads  (coll.  J.  B.  Wilson). 

34.  Leucandra  vaginata,  von  Lendenfeld. 

Leucandra  vaginata,  von  Lendenfeld.     Proceedings  of  the 
Linnean  Society'of  New  South  Wales,  Vol.  IX,  p.  1133. 
Locality. — Port  Jackson  (von  Lendenfeld). 

35.    Leucandra  conica,  von  Lendenfeld. 

Leucandra  conica,  von  Lendenfeld.     Proceedings  of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1 1 26. 
Locality. — Port  Jackson  (von  Lendenfeld). 


Si/no^jiii'^  of  fke  Aitstiullan  Galcarea  Heteroccela.    JJi) 

36.  Lcncandra  Jtisplda,  Carter,  sp. 
Leuconia   hlspida,    Carter.        Annals    and    Magazine    of 

Natural  History,  Auo-ust  1880,  p.  128. 

This  species  is  abundant  in  the  collectiou.  It  is  distin- 
guished by  its  elongated  cylindrical  form,  hispid  surface,  and 
the  long-ra37ed,  slender  triradiates  of  the  main  skeleton.  All 
the  specin'iens  are  solitary  and  sessile,  of  moderate  size  and 
with  a  well-developed  oscular  fringe.  The  large  oxea  of  the 
dermal  surface  are  long  and  comparatively  slender,  slightly 
curved.  The  flagellated  chambers  are  approximately  spherical 
and  average  about  0'09  mm.  in  diameter. 

Mr.  Carter  appears  to  have  had  oid}-  a  single  specimen 
(collected  by  Mr.  Wilson),  which  was  exceptionally  short  and 
"  conoglobular  ; "  I  judge  from  his  description  and  manu- 
script illustrations  of  the  spiculation  that  it  is  speciticalh' 
identical  with  the  specimens  described  above. 

Localities. — Near  Port  Phillip  Heads  (Carter,  and  Stations 
0,  10,  14,  coll.  -J.  B.  Wilson)  ;  Port  Jackson  (coll.  Professor 
Spencer). 

37.  Leucandra  echiwda,  Carter,  sp. 
Leuconia    echiimta,    Carter.      Annals   and    Magazine   of 

Natural  History,  August  188G,  p.  129. 

This  species  is  abundant  in  the  collection.  The  sponge 
usually  has  the  form  of  a  rather  small,  ovoid,  sessile,  thick- 
walled  individual,  with  terminal  fringed  osculum  and  coarsely 
echinated  dermal  surface.  The  species  exhibits  a  good  deal 
of  variation  in  spiculation,  especially  in  the  size  of  the 
irregularly  arranged  triradiates  of  the  main  .skeleton,  which 
are  often  very  much  larger  than  those  of  the  deimal  cortex. 
The  other  forms  of  spicule  present  are  gastral  quadriradiates, 
large  dermal  oxea  (echinating  the  sui-face),  and  long,  slender, 
hair-like  oxea  of  the  oscular  fringe.  The  flagellated  chambers 
are  approximately  spherical  and  densel}"  scattered  through- 
out the  thickness  of  tlie  wall ;  they  measure  about  009  mm 
in  diameter. 

Locality. — Near  Port  Phillip  Heads  (Carter,  and  Stations 
1,9,  10  and  outside  the  Heads,  coll.  J.  B.  Wilson);  Watsons 
Ba^^  Port  Jackson  (coll.  T.  Whitelegge). 

38.    Leucandra.  niultifida,  Carter,  sp. 
Leuconia   iiiidtijida,  Carter.     Annals   and    Magazine    of 
Natural  History,  August  1880,  p.  1-il. 

Localitij. — Near  Port  Phillip  Heads  (Carter). 

H  2 


100      Froceediri<js  of  the  Royal  Soeietij  of  Vicforia. 

3!).    Leucaiuira  lohata,  Carter,  sp. 

Leuconia  lohata,  Carter.  Annals  and  Magazine  of  Natural 
History,  August  188{i,  p.  14o. 

Loealiti/.—^eaY  Port  Phillii)  Heads  (Carter). 

40.     Laacajuhxi  compacta,  Carter,  sp. 

Leuconia  compacta,  Carter.  Annals  and  Magazine  of 
Natural  History,  August  1886,  p.  144. 

Locallfi/.—Neav  Port  Phillip  Heads  (Carter). 

41.    Leucandra  ■phUlipensls  n.  sp. 

The  single  specimen  in  the  collection  is  a  solita)-y,  sessile, 
irregularly  sac-shaped  sponge,  with  a  constricted  terniinal 
osculum  provided  with  a  feebly  developed  oscular  fringe. 
The  outer  surface  of  the  sponge  is  slightly  hispid  and  the 
wall  of  the  sponge  is  ratlier  thin,  enclosing  a  wide  gastral 
cavity.  The  height  of  the  spoiige  is  about  40  mm.,  the 
greatest  width  20  mm.,  and  the  thickness  of  the  wall  nearly 
;]  mm.  There  is  a  very  thin  dermal  and  gastral  cortex. 
'J'he  canal-sj'stem  is  very  typical  ;  thickly  scattered  groups 
of  dermal  pores  lead  into  wide,  more  or  less  lacunar  inhalant 
canals,  which  penetrate  dee]>  into  the  substance  of  the  wall. 
The  exhalant  canals  are  also  wide  and  deep  and  unite  to- 
gether in  groups  before  opening  into  the  gastral  cavity. 
Between  these  wide  inhalant  and  exhalant  canals  the 
tiagellated  chambers  are  thickly  scattered ;  these  are 
generally  more  or  less  ovoid  in  shape  but  onlyaboi.it  0-14 
mm.  in  longer  diameter. 

The  skeleton  is  rather  weak  owing  to  the  prevailing 
slenderness  of  the  spicule-rays,  the  spiculation  being  as 
follows: — (1)  Gastral  quadriradiates ;  usually  more  or  less 
sagittal ;  with  very  long,  slender,  straight  or  neai-ly  straight, 
sharp-pointed  facial  rays  ;  the  oral  angle  wider  than  the 
paired  angles  and  the  oral  rays  somewhat  longer  than  the 
basal;  oral  rays  about  04  by  O'Ol  mm.;  basal  ray  about  0'3 
by  O'Ol  mm.  The  apical  ray  is  straight  or  slightly  cui'ved, 
conical  and  finely  pointed,  measuring  about  01  (i  by  O'Ol  nun. 
These  spicules  are  very  abundant  and  form  a  thin  gastral 
cortex,  the  apical  rays  projecting  into  the  gastral  cavity  in 
large  numbers.  The  walls  of  the  larger-  exhalant  canals  are 
aJso  provided  with  very  similar  spicules.  Near  the  osculum 
tlie  gastral  spicules  become  much  more  strongly  sagittal  and 


Si/riopsis  of  the  Australian  Calcarea  Heterocada.   ]()] 

the  a[)ical  ray  is  often  absent.  (2)  Triradiates  of  tlie  main 
skeleton  ;  varying  from  nearly  regular  to  slight!}'  sagittal  ; 
with  ver}-  long,  slender,  straight  or  slightly  curved  rays, 
sharply  pointed  and  measuring  about  Oo'i  by  0()](j  nma. 
These  spicules  ai-e  veiy  irregularly  arranged  but  many  of 
them  have  one  ray  pointing  centrifugal!}^  towards  the  dermal 
surface.  In  many  of  them  a  small  apical  ray  is  developed. 
(3)  Triradiates  of  the  dermal  surface  ;  similar  to  the  fore- 
going but  decidedly  smaller  ;  arranged  parallel  to  the  surface 
to  form  a  thin  dermal  cortex,  (i)  .Large  dermal  oxea  ; 
rather  slender,  fusiform,  symmetrical,  very  slightly  curved, 
gradually  sharp-pointed  at  each  end;  measuring  about  1 '4 
by  0  03  mm.;  occasionally  however  they  are  much  largei- 
and  they  may  then  have  a  hastately  pointed  inner  end. 
These  spicules  are  scattered  singly  and  irregularly  at  right 
angles  to  the  dermal  surface,  with  the  outer  end  projecting 
for  a  short  distance.  (5)  Long,  fine,  hair-like  oxea';  these 
are  arranged  in  loose,  iri-egular,  scattered  bundles  between 
the  large  oxea  and  they  also  form  the  feebly  developed 
oscular  fringe. 

Locality.— ^e-Av  Port  Phillip  Heads  (coll.  J.  B.  Wilson). 

42.    Leucandra  gladiator,  n.  sp. 

The  single  specimen  in  the  collection  forms  an  extremely 
irregular,  contorted  crust,  with  a  number  of  deeply  convex 
surfaces,  bordered  by  prominent  margins,  as  if  it  had  grown 
over  some  irregularly  cylindrical  body.  A  few  small  oscula 
are  irregularly  scattered  over  the  convex  upper  surface.  The 
surface  is  slightly  hispid,  the  hispid  character  becoming 
mucli  more  strongly  developed  at  the  margins  of  the  crust. 
The  .specimen  has  been  broken,  but  it  must  have  Ijeen  about 
50  nan.  in  greatest  diameter.  The  growth  has  been 
extremely  irregular,  and  it  has  enclosed  various  foreign 
objects.  The  texture  is  coarse  and  fragile.  The  dei'mal 
cortex  is  strong,  but  not  very  thick. 

The  canal-system  is  difficult  to  work  out  in  detail,  owing 
to  the  strong  development  of  the  skeleton,  wdiich  renders 
section-cutting  very  difficult.  There  is  no  large,  ct.;ntral, 
gastral  cavity,  but  a  number  of  tolerably  wide  exhalant 
canals  converge  towards  each  osculum.  The  flagellated 
chambers  are  irregularly  scattered,  approximately  sjiherical, 
and  about  0  09  mm.  in  diameter. 

The  skeleton  is  composed  of  the  following  spicules  : — (!) 
Gastral  quadriradiates;  minute,  cruciform  or  dagger-sha[>ed ; 


102      ProceediiKjs  of  the  Royal  Societi/  of   Victoria. 

till'  ;n)ii'al  luy  long,  slender,  .straight  and  graduall}'  sliarp- 
])ointed,  nearly  in  a  line  with  the  basal  ray  ;  the  facial  rays 
short,  stout,  conical  and  sharp-pointed,  the  basal  rather 
longer  than  the  other  two  and  often  slightly  crooked,  the 
orals  being  straight;  basal  vav  about  003  by  0007  mm.  ; 
orals  002'by  0007  mm.  ;  apical  008  (or  le.ss)  by  0000  mm. 
These  spicules  are  found  in  the  walls  of  the  larger  exhalant 
canals,  but  they  are  not  very  abundant.  (2)  Enoi-mous 
sub-regular  or  ii-i-egular  triradiates,  with  conical,  gradually 
shar})-pointed  rays  which  measure,  when  fully  developed, 
about  ]  -8  by  0-16  nun.  These  spicules  form  the  bulk  of  the 
skeleton  and  are  irregularly  and  abundantly  scattered 
throughout  the  thickness  of  tlie  sponge  ;  they  vary  con- 
siderably in  size.  (3)  Small,  straigiit  oxea,  of  hair-like 
fineness  and  up  to  about  O'l  mm.  in  length  ;  scattered 
through  the  interior  of  the  sponge,  either  separately  or  in 
dense  sheaves  (trichodragmata).  (4)  Triradiates  of  the 
dermal  cortex  ;  strongly  sagittal,  with  long,  nearly  straight, 
very  widely  extended,  gradually  sharp-pointed  oral  rays, 
and  much  shorter,  straight,  gradually  sharp-pointed  basal 
ray  ;  these  spicules  form  a  dense  feltwork,  the\^  are  quite 
irregularly  arranged,  excejjt  that  they  all  lie  parallel  to  the 
dermal  surface.  They  vary  greatly  in  size,  the  oi-al  rays, 
which  are  extended  almost  in  a  line,  measuring  up  to  about 
O-tio  by  0-()3G  mm.,  with  the  basal  about  0-3  by  0036  mm. 
(5)  Dermal  oxea  ;  straight,  .slender,  gradually  sharp-pointed 
at  each  end.  In  most  parts  of  the  surface  these  spicules 
are  comparatively  few  in  number,  projecting  at  right  angles 
from  the  dermal  cortex  and  measuring  only  about  O'-t-  by 
O'Ol  mm.  They  viivy  greatly  in  size,  hovvevei",  and  around 
the  margins  of  the  sponge  they  become  very  greatly  elong- 
ated, forming  a  thick,  den.se  fringe. 

This  veiy  remarkable  species  is  obviously  very  closely 
i-elated  to  the  European  Leucandra  nivea,  as  described  by 
Hfeckel  in  "  Die  Kalkschwamme  "  ;  in  both  we  find  colossal 
triradiates,  smaller  dermal  triradiates,  dagger-shaped  quadri- 
radiates  and  trichodragmata  (which  ai-e  extremely  rare  in 
calcareous  .sponges),  and  in  botli  we  meet  with  the  character- 
istic encrusting  habit.  There  are,  however,  certain  marked 
differences  in  spiculation,  as  in  the  shape  of  the  dermal 
triradiates  and  of  the  dagger-shaped  quadradiates,  and 
especially  in  the  presence  in  L.  gladiator  of  the  projecting 
dei'mal  oxea,  which  seem  to  be  entirely  wanting  in 
L.  inrea. 


Si/nopsis  of  tJie  Avf;fr(dia)i   Calcarea  Hetevocaela.  103 

It  is  important  to  notice  that  Mr.  Carter's  Leuconia 
iiivea  var.  austra.Heiislfi  appears  to  be  totally  distinct  both 
from  the  true  Li'ucavdra  nivea  and  from  L.  gladiator.  It 
is  carious  that  Mr.  Cai'ter  should  have  chosen  tliis  name  for 
onej^of  Ml'.  Wilson's  sponges  and  that  later  on  Mr.  Wilson 
should  have  obtained  from  the  same  locality  another  s})ecies 
which  really  is  ver}"  closely  related  to  the  remarkable 
Leucandm  nivea. 

Localitfj.—  OntH'xde  Port  Philli})  Heads  (coll.  J.  B.  Wilson). 

43.    Leucandra  carteri,  n.  sp. 

Leucaltis  Jioridana,  var.  australiensis,  Carter.  Annals 
and  Magazine  of  Natural  History,  August  1886.  p.  l-io. 

This  species  appears,  from  Mr.  Carter's  description,  to  be  a 
large,  massive  Leucandra,  resembling  L.  microrapkis  in 
form  but  distinguished  by  the  presence  of  minute  oxea  on 
the  surface.  As  the  name  australiensis  is  already  occupied 
in  the  genus  I  pi'opose  to  call  the  species  Leucandra  carteri. 

Locality.  —  Near  Port  Phillip  Heads  (Carter). 

44*.    Leucandra  schuhei,  Polejaetf,  sp. 

Eilhardia  scJudzei,  Polejaetf.  Report  on  the  Calcarea  of 
the  Challenger  Expedition,  p.  70. 

Localities. — Otf  Pt.  Jackson  and  Twofold  Bay  (Polejaeff). 

4-5.    Leucandra  loricata,   Polejaetf,  sp. 

Leuconia  loricata,  Polejaeff.  Report  on  the  Calcarea  of 
the  Challenger  Expedition,  p.  03. 

Leiicoriis  loricata,  von  Lendenfeld.  Proceedings  of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1123. 

Locality. — Off  P(n-b  Jackson  (Polejaetf). 

46.    Leucandra  palvinav,  H^eckel,  sp. 

I^eucortis  irulviuar,  Hieckel.  Die  Kalkschwamme,  Vol.  II, 
p.  162. 

This  species  ranges,  according  to  Hasckel,  from  the  Red 
Sea  to  the  west  coast  of  Australia.  I  have  not  yet  met 
with  it. 

Locality.— West  coast  of  Australia  (Hfeckel.  Also  recorded 
from  the  Red  Sea  and  various  paits  of  the  Indian  Ocean). 


104     Proceedings  of  the  Royal  Society  of  Victoria. 

47.    Leucandra  Jtelena,   von  Lendeiifeld,  sp. 

Leucaltis   helena,  von    Lendenfeld.     Proceedings   of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1119. 
Locality. — Port  Jackson  (von  Lendenfeld). 

48.    Leucandra  puiuila,  Bowerbank,  sp. 

Leacoma  pumila,  Bowerbank.  Monograph  of  British 
Sponges,  Vol.  2,  p.  41. 

Leucaltis  'pumila,  Hpeckel.  Die  Kalkschvv'amme,  Vol.  '2, 
p.  148. 

Locality. — Bass  Straits  (Ha?ckel.  Also  recorded  from 
various  localities  in  the  Atlantic  Ocean,  vide  Bowerbank  and 
Haickel). 

49.  Leucandra  hatkybia,  Hai-cke],  sp. 

Leucaltis  batJtybia,  Hpsckel.  Die  Kalkschwamnie,  Vol.  2, 
p.  loG. 

Leucaltis  bcdkybia  var.  australiensis,  Ridley.  Zool.  Coll. 
H.M.S.  '^Aleit,"  British  Museum,  p.  482. 

Leucaltis  bathybia,  von  Lendenfeld.  Pi'oceedings  of  the 
Linnean  Societ}^  of  New  South  Wales,  V^ol.  IX,  p.  i]21. 

Loccdity. — Port  Jackson  (Ridley.  Recorded  by  Hseckel 
from  the  Red  Sea). 

50.  Leucandra  pandora,  Hpeckel,  sj). 

Leucetta  pandora,  Hseckel.  Die  Kalksehwarnme,  Vol.  2, 
p.  127. 

Localities. — Bass  Straits  and  Gulf  of  St.  Vincent  (Hseckel). 

51.   Leucandra  microraphis,  Hjieckel,  sp. 

Leucetta  microraphis,  Hseckel.  Die  Kalkschwamme, 
Vol.  2,  p.  119  {= Leucetta  primigenia  var.  microraplt.is). 

Leucetta  microrapJiis,  von  Lendenfeld,  Proceedings  of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1J17. 

Leuconia  dura,  Polejaeff.  Report  on  the  Calcarea  of  the 
Challenger  Expedition,  p.  65. 

I  identify  with  this  species  a  number  of  large  specimens 
of  very  irregular  external  foi-m.  They  are  sometimes  com- 
pressed, sometimes  massive  and  sometimes  sac-shaped,  with 
thick   walls ;    usually  Mnth    wide    naked    oscula   and    large 


Synopi:ils  of  the  Au'^irallan  Calcarea  Heterocwla.   105 

exhalant  canals.  The  texture  is  very  coaise,  bard  and  dense; 
the  surface  is  irregular,  and  often  characteristically  ridged  ; 
frequently  the  huge  triradiate  spicules  can  be  seen  with  the 
naked  eye  on  the  dermal  suiface.  Some  specimens  have  a 
few  quadriradiate  spicules,  while  in  others  I  cannot  find  any. 

Some  of  the  s])ecimens  measure  four  or  five  inches  in  their 
longest  diameter,  and  one  was  remarkable  from  having  been 
of  a  green  colour  in  life,  probably  due  to  symbiotic  algre. 

The  flagellated  chambers  are  approximately  spherical  ; 
thickly  scattered  through  the  sponge,  and  about  ()'12  mm. 
in  diameter.  In  some  specimens  the  mesoderm  around  the 
chambers  is  very  strongly  developed,  giving  to  the  sponge  a 
very  dense  texture.  The  inhalant  pores  are  scattered  thickly 
over  the  dermal  surface,  at  any  rate  in  parts. 

The  skeleton  is  dense  and  very  irregular,  consisting  of 
scattered  triradiates  of  two  very  different  sizes,  i-ather  small 
and  enormously  large,  the  former  being  most  abundant. 

I  ccuisidor  Pol^jaefi's  Leuconia  dura  to  be  identical  with 
this  species,  because  I  do  not  think  the  presence  of  sagittal 
spicules  in  the  neighbourhood  of  the  osculum  is  a  specific 
character,  as  it  is  of  such  extremely  general  occurrence. 

Localities. — Near  Port  Phillip  Heads  (Stations  1  and  .9 
and  outside  the  Heads,  coll.  J.  B.  Wilson)  ;  Torres  Straits 
(Ridley,  Pol^jaefi)  ;  Port  Jackson  (von  Lendenfeld). 

52.    Leucandra  Jimtkeliaiia,  Polejaefi",  sp. 

Leucetta  hceckeliana,  Polejaeff.  Rejiort  on  the  Calcarea 
of  the  Challenger  Expedition,  p.  69. 

Voamaeria  hceckeliana,  von  Lendenfeld.  Proceedings  of 
the  Linnean  Society  of  New  South  Wales,  Vol.  IX,  p  ]  1 14. 

Loc(ditj/.—Oii'  Port  Jackson  (Polejaefi). 

53.    Lelapia  aastralis,  Carter. 

(?)  Lelapia  austral  is,  Gray.  Proceedings  of  the  Zoological 
Society  of  London,  18G7,  p.  557. 

Lelapia  austrcdis,  Carter.  Annals  and  Magazine  of 
Natural  History,  August  1 88G,  pp.  1 38  and  1 48. 

This  sponge  appears  tt)  be  of  exce|)tional  interest,  and  I 
greatly  regret  that  I  have  never  had  an  opportunity  of 
examining  it. 

Localitij.—]<leAv  Port  Philli))  Heads  (Carter). 


lOG      Proceeding.'^  of  flie  Royal  Society  of  Victoria. 

54".    Gvantesfia^  sacca,  von  Lendenfekl. 

Gninies.'ia  aucca,  von  Lendenfekl.  Proceedings  of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1098. 

Hypoyi-antia  aacca.  Carter.  Annals  and  Magazine  of 
Natural  History,  July  188(),  p.  42. 

This  very  beautiful  sponge  is  well  represented  in  the 
collection.  All  the  specimens  which  I  have  seen,  six  in 
number,  are  more  or  less  compressed,  in  the  case  of  large 
specimens  veiy^  much  so.  The  finest  specimen  is  GO  mm.  in 
height  by  50  mm.  in  greatest  breadth.  Von  Lendenfeld 
repi-esents  the  radial  chambers  as  being  perfectly  straight 
and  unbranched,  whereas,  in  the  Victorian  specimens,  they 
l)rancli  ie))eatedly,  the  branches  running  parallel  with  one 
another  to  the  dermal  surface.  This  may  possibly  constitute 
a  specific  difference,  but  I  am  more  inclined  to  think  that 
the  tigui-e  leferred  to  is  incorrect. 

Localities. — Port  Jackson  (von  Lendenfeld)  ;  near  Port 
Pliillip  Heads  (Carter,  and  outside  the  Heads,  coll.  J.  B. 
Wilson). 

55.    tirantes.sa  hlrsuta,  Carter,  sp. 

Hyp()</faiiti'i  hlrsuta,  Cartel-.  Ann-ils  and  Magazine  of 
Natural  History,  July  18S(i,  p.  41. 

In  an;itomical  chaiacters  this  species  closely  i-esembles 
G.  sacca,  but  differs  markedly  in  external  appearance  and  in 
the  less  regularly  arranged  tubar  skeleton,  which,  though 
composed  of  slender  spicules,  is  very  dense  and  confused. 
The  suiMlermal  sagittal  triradiates,  with  inwardly  directed 
basal  r;  y,  are  not  mentioned  by  Mr.  Carter;  they  are  clearly 
present  in  my  specimens  thougli  less  obvious  than  in  G. 
sacca  on  account  of  tlie  somewliat  confused  character  of  the 
tubar  skeleton.  The  oxea  of  the  dermal  tufts  are  straight, 
or  only  very  slightl}^  curved.  The  manuscript  illustration 
which  Mr.  Carter  has  kindly  sent  me  shows  only  a  very 
slight  curvature,  though  he  describes  them  as  "  curved." 
The  species  is  abundant  in  the  collection. ' 

Localities. — Near  Port  Phillip  Heads  (Carter;  and  Station 
1  and  outside  the  Heads,  coll.  J.  B.  Wilson)  ;  King  Island 
(coll.  Prof  Sjiencer)  ;  Holiart,  Tasmania  (coll.  A.  Dendy). 

5().    Gratitessa  klsplda,  n.  sp. 

Small,  cylindrical  or  slightly  compressed,  solitary  persons, 
with   moiv  or  less  distinct  fringt-  around  the  terminal  oscu- 


Sijnojms  of  the  Austr-dian  Calcarea  Heterocoelc.   107 

lum  and  strongly  hispid  surface.  The  largest  specimen  .in 
the  collection  is  about  40  mm.  high  by  4  mm.  in  greatest 
diameter,  the  wall  of  the  sponge  being  only  about  O'oG  mm. 
thick.  The  canal  system  closely  resembles  that  of  G.  sacca, 
but  the  chambers  are  shorter  and  less  branched.  Tlie  skele- 
ton is  arranged  as  in  G.  sacca,  but  the  dermal  tufts  of  oxea 
are  less  definite  and  less  regularly  aiTanged,  and  the  tubar 
skeleton  is  composed  of  much  fewer  joints.  The  spiculation 
is  as  follows:— (I)  Gastral  quadriradiates  ;  very  rare,  with 
short  apical  ray.  (2)  Gastral  triradiates  ;  strongly  sagittnl, 
with  very  long  and  slender  rays ;  gradually  sharp-pointed  •, 
basal  ray  straight,  about  0-3  by  O'OOSS  mm. ;  oral  rays 
straight  or  slightly  crooked,  often  unequal  in  length,  about 
0-2  by  0008  mm.  "  (3)  Subgastral  sagittal  triradiates  ;_  oral 
rays  widelv  extended,  slightly  recurved,  sharply-pointed, 
about  0-12'by  0-0082  nun.  ;  basal  ray  long,  straight,  sharp- 
pointed,  varying  in  length  up  to  about  0-3  mm.,  and  about  as 
thick  as  the  orals,  extended  in  a  centrifugal  direction  through 
the  chamber  layer.  (4)  Tubar  triradiates,  resembling  the 
foregoing,  with  similar  very  long  basal  rays.  (5)  Subdermal 
sagittal  triradiates  ;  with  veiy  widely  extended  oral  rays 
lying  in  the  dermal  cortex,  and  long  straight  basal  ray 
extending  inwards  through  the  chamber  layer  ;  oral  rays  up 
to  about  0-2  by  O'Ol  mm.  ;  basal  ray  up  to  about  0-32  by 
001  mm.;  all  sharply-pointed.  (6)  Dermal  triradiates; 
sagittal,  resembling  the  foregoing,  but  often  very  irregular 
and  with  shorter  basal  ray.  "(7)  Dermal  oxea;  long,  straight 
or  slightly  curved,  spindle-shaped,  gradually  sharp-pointed 
at  each  end,  length  variable,  up  to  about  07  by  0-UlG  mm.  in 
the  largest  specimen,  but  much  longer  and  slenderer  in  one  of 
the  smaller  ones.  The  spicules  of  the  oscular  fringe  do  not 
<lifFer  markedly  from  these. 

Locality.— i^ em-  Port  Phillip  Heads  (Station  9,  coll.  J.  B. 
Wilson). 

57.    Gmnte.ssa  j^oculum,   Polejaeff,  sp. 

AmijhorisciDi  pocidum,  Polejaeff.  Report  on  the  Calcarea 
of  the  Challenger  Expedition,  p.  46. 

Hetero^na  patulosculifera,  Carter.  Annals  and  Magazine 
of  Natural  History,  July  188G,  p.  49. 

A.  careful  examination  of  portions  of  Carter's  and 
Polejaeff 's  type  specimens  from  the  British  Museum  has 
convinced   me   that  the   two  are  specifically  identical,  and 


108     Proceedings  of  the   Royal  Sockty  of  Victoria. 

I  therefore  revert  to  the  earlier  specific  name.  I  have  also 
two  other  specimens  collected  by  Mr.  Wilson,  one  of  which 
exhibits  very  beautifully  the  "  agglomerated "  character 
mentioned  by  Carter,  while  the  otlier  is  only  a  fragment. 
The  quadriradiates  mentioned  by  Carter  are  scarce  and 
inconspicuous,  and  I  have  not  noticed  them  in  the  other 
specimens. 

Localities. — Off  Port  Jackson  (Polejaeff )  ;  near  Port 
Phillip  Heads  (^Carter  and  coll.  J.  B.  Wilson). 

58.    Grantessa  eriiiaceus,  Carter,  sp. 

Leuconia  erinaceus,  Cartel'.  Annals  and  Magazine  of 
Natural  History,  August  1886,  p.  180. 

This  species  is  readily  recognised  by  its  external  appear- 
ance and  the  peculiar  arrangement  of  the  dermal  oxea.  The 
flagellated  chambers  are  elongated  and  radial,  but  very 
irregular  and  branching,  and  they  communicate  with  the 
gastral  cavity  by  unusually  long  exhalant  canals,  which 
unite  together  in  groups.  The  tubar  skeleton  is  very 
iriegular,  but  still  presents  clear  traces  of  the  ty])ical 
"  articulate  "  arrangement.  Subgastral  sagittal  triradiates 
are  present,  and  the  subdermal  sagittal  triradiates,  with 
inwardly  directed  basal  rays,  are  very  conspicuous.  There 
is  a  dense  dermal  cortex  of  much  smaller  triradiates,  and  a 
less  well-developed  gastral  cortex.  Endogastric  septa,  with- 
out spicules,  are  pi-esent  in  both  my  specimens,  and,  as  Mr. 
Carter  also  mentions  them  in  his,  they  would  seem  to  be 
characteristic  of  the  species. 

Locality. — Near  Port  Phillip  Heads  (Carter,  and  Station  7, 
coll.  J.  B.  Wilson). 

59.    Grantessa  intusariicidata.  Carter,  sp. 

Hy pograntia  intusarticulata,  Carter.  Annals  and  Maga- 
zine of  iS^atural  History,  July  188G,  )).  45. 

Hypograntia  medioarticidata,  Carter,  loc.  cit.  p.  46. 

I  have  eleven  specimens  which  I  believe  to  be  all  referable 
to  this  species,  and  I  am  strongly  of  opinion  that  Mr.  Carter's 
Hypograntia  mecUoarticukUa  is  specifically  identical  with 
his  intusarticulata.  The  minute  details  of  spiculation  vary 
considerable  in  different  specimens,  the  most  charactei'istie 
features  being  the  dermal  ci'ust  of  minute  oxea  or  "  mortar 
spicules,"    and     the    subdermal    sagittal    triradiates.        The 


Si/nopfii-'s  of  tit e  AuMvuVian  Calcarea  Heterocuda.   109 

radial  chambers  are  laucli  Viraiiclied,  which  I  believe  is 
what  Mr.  Carter  means  when  he  says  that  they  intercom- 
municate by  lai-ge  holes.  The  branches  run  parallel  with 
one  another  to  the  dermal  cortex  ;  the  exhalant  canals  are 
short. 

Localities. — Near  Port  Fliillip  Heads  CL'ai'ter,  and  Stations 
3,  5,  cS,  9  an.l  outside  the  Heads,  coll.  J.  B.  Wilson)  ;  Wat- 
son's Ba3%  Port  Jackson  (coll.  T.  Whitelegge). 

60.    GntntessK  (?)  polupeyhtoraia,  Carter,  sp. 

Heteropia  polype listoinUt,  Carter.  Annals  and  Magazine 
of  Natural  History,  July  1 886,  p.  4-7. 

Locality.— ^en.Y  Port  Phillip  Heads  (Carter). 

61.    Grantessa  (?)  compressa,  Cartel',  sp. 

Heteropia  compressa,  Carter.  Annals  and  Magazine  of 
Natural  History,  July  1886,  p.  ol. 

Locality. — Near  Port  Phillip  Heads  (Cartel-). 

62.    Grcmtessa  (?)  plwrloscidifeva,  Carter,  sp. 

Heteropm  pliirloscullfera.  Carter.  Annals  and  Magazine 
of  Natural  History,  July  1886,  p.  o2. 

Locality. — Near  Port  Phillip  Heads  (Carter). 

63.  Giuntessa  (?)  ei-ecta.,  Carter,  sp. 

Heteropia  erecta,  Carter.  Annals  and  Magazine  of  Natural 
History,  July  1886,  p.  53. 

Locality. — Near  Port  Phillip  Heads  (Carter). 

64.  Grantessa  (?)  splssa,  Carter,  sp. 

Heteropia  splssa,  Carter.  Annals  and  Magazine  of 
Natural  History,  July  1886,  p.  54. 

Locality. —  Near  Port  Phillip  Heads  (Carter.) 
The  last  five  species  are  described  by  Mr.  Carter  apparently 
from  single  specimens,  all  collected  by  Mr.  Wilson.  It  appears 
to  me  very  doubtful  whether  they  are  all  specifically  distinct, 
and  also  whether  some  of  them  at  any  rate  are  not  mere 
varieties  of  G.  inoculum  or  Vosmaeropsls  macera,  which 
they  resemble  in  spiculation.  Unfortunately,  I  have  not 
seen  any  of  the  types. 


110      I'rocccOi  iKjs  of  the   lh>ij(i.l  Socicfij  of   Vlcfori'i. 

Go.    Vo.smacfoihsi.s  laoct'rd,  Carter,  sp. 

Hetei'opla  macerd,  Carter.  Antials  ami  Magazine  of 
Natural  History,  July  J 880,  p.  oO. 

This  species  is  well  represented  in  the  collection  I  have 
been  able  to  convince  myself  of  the  correctness  oi  the  identi- 
fication by  a  niinnte  comparison  of  a  piece  of  Mr.  Carter's 
type  fi'oni  the  Britisii  Museum.  It  is  remarkable  for  its 
densely  agglomerated  or  colonial  habit.  Specimens  may 
attain  a  large  size,  consisting  of  veiy  numerous  individuals 
almost  completely  fused  together,  usually  in  linear  series, 
which  are  inter-connected  by  cioss-bars.  The  oscula  are 
i-aised  on  conical  prominences,  and  each  indicates  a  separate 
gastral  cavity.  The  canal  system  is  remarkable.  The 
chambers  are  thimble-shaped  and  mostly  widely  separated 
from  the  gastral  cavity,  with  which  they  conuiiunicate  by 
a  strongly  developed  system  of  exhalant  canals,  each  being- 
separated  from  its  exhalant  canal  by  a  well  developed 
diaphragm.  Those  chambers  which  lie  next  to  the  dermal 
suriace  still  exhibit  a  radial  arrangement  with  regard  to 
the  long  axis  ot  the  individual.  Both  subdermal  and 
subgastral  sagittal  trii'adiatcs  arc  strongly  developed. 

Locality. — Near  Port  Phillip  Heads  (Cartel',  and  coll. 
J.  B.  Wilson). 

06.     Vosmaeropslti  (Jepreaaa,  n.   sp. 

Specimen  flattened,  cushion-shaped,  with  flat  under  and 
convex  up])er  surface.  About  12  mm.  in  horizontal  diameter, 
and  only  4  mm.  thick  in  the  middle.  Maigin  rounded, 
i-OLighly  cii'cnlar  in  outline.  There  is  no  wide  gastral  cavit}', 
but  several  large,  branching  exhalant  canals  converge  to  a 
single  small  osculum  situate  near  the  middle  of  tlie  uppei' 
surface.     Sinface  smooth  ;  no  distinct  oscular  fringe. 

The  inhalant  canal-sy.stem  is  quite  irregulai-,  commencing 
in  wide  lacunar  spaces  situated  beneath  the  thin,  pore-bear- 
ing dermal  cortex.  The  flagellated  chambers  are  irregularly 
but  thickl3' scattered  throughout  the  thickness  of  the  sponge, 
with  no  trace  of  radial  airangement  around  a  central  gastral 
cavity.  They  are  irregular-ly  sac-shaped  or  thimble-shaped, 
measuring  about  0'2  by  009  mm. 

The  bulk  of  the  skeleton  is  made  up  of  fairly  large,  sub- 
regular  or  slightly  sagittal  triradiates,  scattered  witliout 
definite  ordei-  throughout  the  thickness  of  the  sponge,  but 
many    with    one  .slightly   longer   ray   jiointing   towards   the 


SynojJsi.i  of  the  Australian  Calcarea.  Heterocada.   Ill 

dermal  surface.  Beneath  the  dermal  surface,  but  apparent 
only  on  the  upper  surface  of  the  sponoe,  is  a  distinct  lawyer 
of  subdermal  sagittal  triradiates  with  inwardly-directed 
basal  ray.  The  dermal  skeleton  is  made  up  principally  oi" 
subi-eguiar  triradiates  of  various  sizes,  placed  horizontally, 
but  with  no  definite  ariangement ;  amongst  these  very 
minute,  slender  oxea  are  scattered,  rare  on  the  upper  surface 
of  the  sponge  but  abundant  on  the  lower ;  around  the 
osculuin  these  oxea  are  numerous  and  a  few  are  much  lai-ger 
than  the  rest.  Around  the  main  exhalant  canals  there  is  a 
layer  of  small  sagittal  triradiates.  The  forms  and  dimen- 
sions of  the  different  spicules  are  as  follows: — (1)  Triradiates 
of  the  exhalant  canals  ;  sagittal,  rays  conical,  fairly  sharply- 
pointed  ;  basal  straight,  orals  usually  slightly  incurved  or 
recurved  ;  basal  commonl}^  somewhat  shorter  than  orals, 
which  measui-e  about  ()i6  b}'  0012  mm.  Just  below  the 
osculum  I  have  seen  short  apical  rays  in  a  few  of  the  sagittal 
radiates.  (2)  Triradiates  of  the  main  skeleton  ;  subregular 
or  slightly  sagittal ;  rays  usually  straight,  conical,  gradually 
sharp-pointed,  rathei-  slender,  up  to  about  0"3G  by  0-024  mm. 
(3)  Subdermal  sagittal  triradiates  ;  similar  to  the  foregoing 
but  a  good  deal  smaller,  and  with  the  basal  ray  much  longei- 
than  the  others.  (4)  Dermal  triiadiates  ;  subregular,  with 
long,  conical,  gradually  sharp-pointed  rays  varying  greatly  in 
size, up  to  about  0-54-  by  0-045  mm.  (5)  Oxea;  mostly  very- 
minute  and  slende]-,  sharjdy-pointed  at  each  end,  with  one 
end  rather  thicker  than  the  other  ;  straight ;  often  slightly^ 
rougliened ;  usually  short,  but  varying  greatl}^  in  length  ; 
around  the  osculum  a  few  much  stouter  ones  occur,  Init  still 
very  small. 

Localhy. — Near  Port  Phillip  Heads  (Sorrento  Reef,  coll. 
J.  B.  Wilson). 

67.    Vosmaeropsis  ivilsoni,  n.  sp. 

Sponge  colonial,  consisting  of  short,  thick,  sub-cylindrical 
or  truncatedly  conical  individuals  united  together  by  their 
bases  into  smaller  or  larger  agglomerations,  which  may 
attain  a  diameter  of  nearly  five  inches.  Each  fully  grown 
individual  has  a  circular. osculum  at  its  summit,  which  may 
or  may  not  have  an  oscular  fringe,  adjacent  individuals  of 
the  .same  colony  sometimes  differing  in  this  respect.  The 
osculum  is  often  provided  with  a  very  distinct,  membranous 
diaphragm,  situated  a  short  distance  within  its  margin.     Tlie 


112      FroccediiHj.s  of  the  Royal,  Society  of  Victoria. 

iiidixiduals  vaiy  in  size,  and,  owing  to  their  peculiar  colonial 
and  bi-ancliing  habit,  it  is  difficult  to  give  exact  measure- 
ments, but  we  may  put  down  the  average  adult  size  as 
about  20  mm.  long  and  5  mm.  in  diameter.  A  large  colony 
contains  dozens  of  such  individuals  united  together  in  a 
complicated  and  irregular  manner.  The  outer  surface  is 
smooth,  except  for  a  slight  unevenness  due  to  the  presence 
of  large  triradiates,  visible  to  the  naked  eye.  The  colour  of 
spirit  specimens  varies  from  pure  white  to  pale  brown,  but 
one  specimen  which  I  observed  as  it  came  out  of  the  dredge 
was  then  of  a  violet  purple  colour. 

The  gastral  cavity  is  wide  and  cylindrical  and  the  wall  is 
about  2'5  ram.  thick.  There  is  a  dense,  thick  coi'tex  on  both 
gastral  and  dermal  surfaces. 

The  irdialant  pores  are  thickly  scattered  over  the  surface 
of  the  sponge  ;  each  leads  separately  into  a  .shoi't,  narrow, 
cylind]-ical  canal,  situate  in  the  outer  portion  of  the  dense 
dermal  cortex ;  these  canals  soon  unite  to  form  lai'ger,  but 
still  very  well-detined,  cylindrical  canals,  which  anastomose 
with  one  another  by  cross-branches  and  finally  lead  down  to 
the  chamber  layer  between  the  dermal  and  gastral  cortex, 
where  the  canal  system  becomes  more  or  less  lacunar.  The 
flagellated  chambers  are  thickly  scattered  through  the  meso- 
derm of  the  chamber  layer  ;  they  vary  much  in  shape  and 
size,  from  approximately  spherical  and  only  about  0"072  mm. 
in  diameter  to  elongatedly  sac-shaped  and  as  much  as  O'oT 
by  013  mm.*  The  exhalant  canals  unite  together  into 
tolerably  large  trunks,  which  penetrate  the  gastral  cortex 
and  open  into  the  gastral  cavity. 

The  skeleton  is  divisible  into  four  portions,  that  of  the 
gasti'al  cortex,  that  of  the  chamber  layer,  that  of  the  dermal 
cortex  and  that  of  the  osculum.  The  gastral  cortex  is  about 
0'3  mm.  thick  and  its  skeleton  consists  entirely  of  a 
dense  feltwork  of  medium-sized  triradiate  spicules,  arranged 
irregularly  but  parallel  to  the  gastral  surface.  These 
spicules  are  sagittal,  the  oral  angle  being  rather  wider  than 
the  paired  angles  and  the  oral  rays  rather  longer  than  the 
basal;  oral  rays  straight  or  very  slightly  curved  towards  one 
another,  conical  and  gradually  sharp-pointed,  measuring 
about  03  by  0'024  mm.  ;  basal  ray  straight,  conical, 
graduall}^  sharp-pointed,  a  little  shorter  than  the  orals.     The 

*  These  measurements  were  taken  from  different  specimens,  but  it  would  be 
difficult  to  make  a  mistake  as  to  the  species  in  this  particular  case,  and  we 
also  find  considerable  variation  in  the  chambers  even  in  the  same  section. 


Sijnop.si'^  of  the  A  ((■^fralian  Cahiarea  Hefevocud't.  113 

skeleton  of  tlie  chamber  l;t3^er  is  nuule  up  of  lai-ii'e  Mib- 
dermal  and  subgastral  .sagittal  tri radiates,  whose  basal  rays 
penetrate  the  chamber  layer  in  opposite  directions.  These 
spicnles  vary  greatly  in  size,  the  basal  rays  often  extend 
completely  through  the  chamber  layer  aiicl  are  very  thick; 
the  oral  rays  are  shorter,  more  or  less  curved  and  widely 
extended.  Frequently  many  of  those  which  have  centrifugal 
basal  rays  are  not  strictly  subg.istral  but  ha\-e  the  oral  i-ays 
situate  at  various  levels  in  the  chamber  la^^er.  The  i-ays 
are  conical  and  gradually  sharp-pointed.  The  dermal  cortex 
is  about  0-i  mm.  thick  and  its  skeleton  is  rande  up  almost 
entirely  of  triradiate  s{)icules  ot  various  shapes  and  sizes. 
Towards  the  inside  we  find  large,  I'eguiar  or  subregular 
triradiates,  arranged  parallel  to  the  dermal  surface,  with 
conical,  gradually  sharp-pointed  rays  which  measure  up  to 
about  10  by  0' 17  muj.;  man\^,  however,  being  much  smaller. 
On  the  outside  is  a  much  thinner  layer  of  veiy  different, 
small  triradiates.  These  spicules  are  irregular  in  sha])e  and 
irregularly  arranged  ;  their  rays  are  conical  and  gr;iduall\' 
sharp-pointed,  but  crooked  ;  one  of  them  commonly  projects 
inwai'ds  at  right  angles  to  the  <lermal  surface  ;  the  rays 
measure  about  ()-083  bv  000'i2  mm.  We  also  find  in  tlit' 
outermost  part  of  the  dermal  cortex  a  few  very  minute, 
straight,  slender  oxea,  whose  exact  size  and  shape  are  ver\^ 
difficult  to  determine.  The  osculai-  skeleton  consists  of  a 
fringe  (not  always  visible  to  the  naked  eye  but  sometimes 
strongly  developed)  of  very  long  and  veiy  slender  oxea. 

I  i)ave  much  pleasure  in  dedicating  this  very  remarkable 
and  abundant  species  to  Mr.  J.  Biucebridge  Wilson,  who  lias 
collected  all  the  specimens  at  present  known. 

Locality.— OntHide  Port  Phillip  Heads  (coll.  J.  B.  Wilson). 

08.    IJ('ieroj)egin(i   nodus  <jordii,  Polejaeff. 

Heteroperpmi  /^(^tl((NW/u/'(7//,  Polejaefi!  Re[)ort  on  the  Cal- 
carea  of  the  Challenger  Expedition,  p.  4-."3. 

The  only  specimen  which  I  have  seen  of  this  species, 
unless  indeed  H.  latitubulatd,  be  considered  specifically 
identical,  is  a  portion  of  Pol^jaeff's  type  specimen  in  the 
British  Museum.  I  have  nothing  to  add  to  Polejaeff 's 
excellent  description. 

Locality. — Torres  Straits  (Polejaeff.  Polejaefi"  also  records 
the  species  from  the  Bermudas). 

I 


114      Proceediiui-s  of  the   Roi/(U  Society  of   Victoria. 

(59.    Hetcropeijma  lutitulnildta,  Cartel',  sj). 

Clafhrina  latitubidatd  (|)i-ovi,sional,  iiicertie  sedis),  Carter. 
Annals  and  Magazine  of  Natural  History,  June  188G,  p.  515. 

After  describing  the  external  form  and  spicnlation  of  this 
lemarkalile  sponge,  Mr.  Carter  remarks  that  in  general  form 
it  is  very  nuich  like  Vo]eyAeii''i<.  IIefero2'egnta  itodtts  gordii, 
l)ut  totally  different  amitomieally.  I  have  fortunately  been 
able,  owing  to  the  kindness  of  the  authorities  of  the  British 
Museum,  to  make  a  minute  anatomical  examination  both  of 
Polejaeff's  type  of  Hcte ropegina  'nodii^i  gordii,  and  also  of 
Carter's  type  of  Cldthriiui  liitituhvlata,  and  I  have  also 
received  the  latter  species  direct  from  Mr.  Wilson.  I  find 
that  in  external  forn),  canal-system,  and  also  in  the  ari-ange- 
ment  of  the  skeleton,  the  two  species  are  identical,  agreeing 
with  the  admii-able  figures  given  by  Polejaeff'in  his  Challenger 
Rc[>oit.  The  only  difference  which  I  have  been  able  to 
detect  concerns  the  size  and  shape  of  the  minute  quadriradiate 
spicules  of  the  chamber  layer,  which  are  even  further  reduced 
in  //.  liditnhuhda  than  thev  are  in  li.  nodus  gordii. 

Locality.— l^em-  Port  Phillip  Heads  (Carter,  and  Station  1, 
coll.  J.  B.  Wilson). 

70.    Amphorisciis  cyathiftcus,  HaH:kel. 

Arui>horiscus  cyatJd-scas,  Hseckel.  Prodromus  cines 
Systems  der  Kalkschwamme.  Jenaische  Zeitschrift,  Vol.  5, 
part  2,  ]\  238. 

Sycilla  cyatJrii<cus,  Hai^ekel.  JJie  Kalkschwamme,  Vol.  2, 
p.  250. 

Locality.  —  South  Australia  (Haickel). 

71.    Aiiiphorimns  cylindrus,  Hseckel,  sp. 

Sycilla  cylindrus,  Hajckel.  Die  Kalkschwamme,  Vol.  2, 
p.  254. 

Amphoriscus  cylindriL^;  von  Lendenfeld.  Proc.  Linn. 
Soc.  N.S.W.,  Vol  iX,  p.  1103. 

Locality. — Port  Jackson  (von  Lendenfeld.  Recorded  by 
Hoeckel  from  the  Adriatic). 

72.   Leucilla  uter,  Polejaefi". 

Leucilla  uter,  Polejaefi".  Report  on  the  Calcarea.  of  the 
Challenger  Expedition,  p.  53. 

Folejna  uter,  von  Lendenfeld.  Proceedings  of  the 
Linnean  Societ}'  of  New  South  Wales,  Vol.  IX,  p.  1115. 


Si/nopsis  of  the  ALudfallan  Calcarea  HeterocmUi.  115 

Localities. — Torres  Strjiits  (von  Lendenfeld.  Recorded  by 
Polejaeff  f roui  the  Phillipine  Islnuds  and  the  Bermudas). 

73.   Lev,cill(i  imperfecta,  Polejaeft',  sp. 

Leueetta  irtvpevfecta,  Polejaeft'.  Report  on  tlie  Calcarea  of 
the  Challenger  Expedition,  p.  67. 

Vosriiaeria  imperfecta,  von  Lendenfeld.  Proceedings  of 
the  Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  11 13. 

Localitii.—O^^  Port  Jackson  (Polejaeft). 

7-t.   Leucilla  audraliensis,  Carter,  s[). 

Leuconia  johnstonii,  var  aiistraliensis,  Carter.  Annals 
and  Magazine  of  Natural  History,  August  1886,  p.  133. 

This  beautiful  little  species  nearly  always  presents  itself 
under  the  form  of  a  small,  ov(Md,  sessile,  solitary  person,  with 
single,  circular,  naked,  terminal  osculum.  The  sponge-wall 
is  comparatively  thick,  and  the  dermal  surface  smootii  and 
iiard  owing  to  the  large  quadriradiates.  One  very  large 
specimen  in  the  collection,  hoAvever,  is  conical  in  shape,  and 
has  a  very  irregular  surface,  but  this  is  very  exceptional. 
The  species  is  sometimes  social,  and  rarely  consists  of  two 
or  more  individuals  united  together,  or  of  a  single  branched 
individual  ;  but  the  small  egg-like  form  is  highly  character- 
istic.    There  are  numerous  specimens  in  the  collection. 

The  ftagellated  chambers,  thickly  scattered  through  the 
thickness  of  the  wall,  are  usually  approximately  spherical, 
and  about  O'l  mm.  in  diameter  ;  immediately  l^eneath  the 
dermal  cortex,  however,  they  are  commonly  rather  lai'ger 
and  more  or  less  sac-shaped. 

Locality. — Near  Port  Phillip  Heads  (Carter,  and  Stations 
1,  5,  6,  9,  coll.  J.  B.  Wilson,  and  oft"  Geelong,  coll.  H. 
Grayson). 

75.    Leucilla  prolifera.  Carter,  sp. 

Telchouella  prolifera,  Carter.  Annals  and  Magazine  of 
Natural  History,  July  1878,  p.  85,  and  August  1886,  p.  146. 

This  beautiful  sjjecies  is  represented  in  the  collection  by  a 
number  of  hue  examples,  one  of  which  I  have  already  ftgured 
in  n)y  paper  "  On  the  Anatomy  of  Gixtntia  labyriiifhica, 
Carter,  and  the  so-called  Family  Teichonidse "  {Quarterly 
Journal  of  Microscopical  Science,  Vol.  32,  N.S.)  The 
ftagellated  chambers  are  approximately  spherical  and  about 
0-09  mm.  in  diameter,  thickly  scattered  through  the  sub- 
stance of  the  sponge.  With  the  exception  of  the  small 
quadrii-adiates  in  the  walls  of  the  oscular  tubes,  and  the  very 

I  2 


116      Proceedinfjs  of  ihe  Rojjdl  /S'oc/VV//  of  Vlctorld,. 

large  quadriradiat.es  of  tlie  dennal  surface,  the  skeleton  is 
quite  irregularly  an-anged,  consisting  of  scattered  triradiate 
spicules.  On  account  of  the  large  subdermal  quadriradiates, 
thougli  the  inwai'dly-directed  apical  ray  is  but  short,  i 
propose  to  include  the  species  in  the  genus  Leucilla. 

LocaUtie.s. — Near  Port  Phillip  Heads  (Cai-ter,  and  outside 
the  Heads,  coll.  J.  B.  Wilson)  ;  Freemantle,  W.A.  (Carter). 

70.   Leucilla  saccharata,  Hseckel,  sp. 

Leucandi\t  saccharata,  Hseckel.  Die  Kalkschwiimme, 
Vol.  2,  p.  22S. 

Leucandra  saccharata,  von  Lendenfeld.  Proceedings  of 
the  Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1137. 
'■  This  remarkable  species  exhibits  a  singular  irregularity  in 
external  form,  varying  from  com])ressed,  irregularly-folded 
plates  to  elongated  C3dindrical  tubes,  and  often  attaining  a 
large  size.  It  is  common  in  Port  Jackson,  whence  1  have 
received  specimens  from  Professor  Spencer,  but  I  have  only 
seen  a  single  specimen  from  Port  Phillip,  collected  by  Mr. 
Wilson. 

The  flagellated  chani])ers  are  a])pr(jximately  spherical, 
scattered  irregularly,  about  OOD  mm.  in  diameter.* 

Localities. — Bass  Straits  (Hseckel) ;  Port  Jackson  (von 
Lendenfeld,  and  coll.  Prof.  Spencer)  ;  Port  Denison  (von 
Lendenfeld)  ;   Port  Phillip  (Station  14,  coll.  J.  B.  Wilson). 

77.    Leucilla  rillom,  von  Lendenfeld,  sp. 

Leucandra  villosa,  von  Lendenfeld.  Proceedings  of  the 
Linnean  Society  of  New  South  Wales,  Vol.  IX,  p.  1131. 

The  only  s})ecimen  of  this  sponge  which  I  have  seen  is  a 
piece  of  the  type  from  the  British  Museum,  in  which  sub- 
dermal  quadriradiates,  with  long,  inwardly-directed  apical 
ray,  are  abundant.  I  therefore  include  the  species  in  the 
genus  Leucilla. 

Ijocality. — Port  Jackson  (von  Lendenfeld). 

78.    Parcdeucilla  cucuriiis,  Hreckel,  sp. 

Leucandra  cucuinis,  Hasckel.  Die  Kalkschwiimme,  Vol. 
2,  p.  205. 

Localities. — Bass  Straits  and  Gulf  of  St.  Vincent  (Haeckel. 
Also  recorded  by  Htekel  from  Ceylon). 

•  Von  Lendenfeld  gives  tbe  diameter  as  0-04  mm.,  but  this  is  probably  an 
error,  for  ho  also  says  that  in  Leucandra  ti/pica  tbe  chambers  "are  smaller 
than  in  any  other  ease,  their  diameter  rarely  exceeding  0-04  mm."  (loc.  cit., 
p.  1130. 


Art.  VII. —  On  Two  Nein  Tertiary  Stylasterids.. 


(With  Plate  XIII.) 

By    T.    S.    Hall,    M.A. 

[Bead  October  IS,  1892.] 


No  members  of  the  family  Stylasteridm  have,  I  beheve, 
been  recorded  as  fossil.s  in  Austi-alia,  then-  small  size  having 
caused  them  to  be  overlooked  by  collectors.  The  specimens 
I  have  found,  were  obtained  by  washing  the  clays  which 
are  so  characteristic  of  our  Eocene  deposits. 

The  airangement  of  the  pores  in  the  cyclosystems  of  both 
species  seems  to  warrant  the  formation  of  new  genera  for 
their  reception  ;  at  any  rate,  they  will  not  fit  into  any  of 
the  genera  defined  by  Mosele3^ 


Genus,  Deontopora  (gen.  nov.) 

Dactylopores  arranged  in  an  arc  of  about  three-quarters 
of  a  circle  round  a  gastropore  at  the  centre,  and  absent  on 
the  inner  or  attached  edge  of  the  c\'closystem.  There  are 
no  styles  visible  on  a  superficial  examination  in  the  dactylo- 
pores, and  the  presence  of  matrix  in  the  gastropoi-es  pre- 
vented the  seai'ch  for  them  there  without  mutilation  of 
what  is  at  present,  the  oidy  specimen  I  have. 


D.  mooraboolensis  (sp.  nov.) 

The  coenosteum  is  branched,  but  its  general  form  is  as 
yet  unknown.  The  portion  found  is  about  1  cm.  long  and 
2  mm.  in  diameter.  The  surface  is  composed  of  dense  cal- 
careous tissue,  and,  as  in  Astylas  suhviridis  (Moseley),* 
is    marked    by    conspicuous    longitudinal    rounded    ridges, 

*  "  On  the  Structure  of  the  Stylasteriche."     Phil'.  Trans.,  187S,  p.  4.57. 


118      Proceedings  of  the  Royal  Society  of   Victoria. 

sepaj'sited  by  grooves.  The  ridges  are  pai-allel  and  can  be 
traced  separately  for  some  distance.  In  sonje  places,  brandl- 
ing and  anastonu/sing  of  the  ridges  takes  place.  Over  the 
.surfaces  of  the  ampulla^  the  ridges  are  contorted  very  much, 
and  tlieir  individuality  is  preserved  for  a  very  shoit  distance. 
In  the  groo\es  between  the  ridges,  under  a  strong  light, 
narrow  slit-like  pits  can  be  seen  with  a  hand  lens.  Pourtales, 
as  quoted  by  Moseley,*  mentions  small  linear  pores  arranged 
in  rows  and  scattered  over  the  whole  coenenchyma  in 
Pliobothrus  symmetricus  (Pourtales).  These  pores,  Moselej' 
states,  are  occupied  by  canals  of  the  coenosarcal  meshwork 
in  the  recent  condition.  The  pits  in  Deontopora  are 
probably  the  mouths  of  pores  with  the  same  function,  as 
the  mici-oscope  shows  they  have  considerable  depth.  In 
two  places,  where  smaller  branches  had  been  broken  off,  the 
ccenosteum  had  a  cellular  appearance,  but  this  was  not 
visible  on  the  ends  of  the  specimen. 

The  cyclosystems  are  arranged  alternately  on  opposite 
sides  of  the  branch,  and  are  a  little  over  1  nnn.  in  diameter. 
The  axis  of  the  system  and  that  of  the  bi'anch,  form  an  angle 
of  about  45°.  The  inner  wall  of  the  gastropore  is  confluent 
with  the  side  of  the  branch,  and  the  cord-like  ridges 
mentioned  above,  run  right  down  into  the  mouth  of  the 
pore.  The  dactylopores  are  placed  on  an  elevated  flattened, 
horse-shoe-sha[)ed  ridge  overlooking  the  gastropore.  Each 
dactylopore  is  situated  in  a  broad,  deep  gi-oove,  at  about 
one-third  of  its  length  from  the  outer  end  of  the  groove. 
This  groove  cuts  the  outer  wa\\  of  the  cyclosystem,  wliile 
its  inner  end  runs  out  on  to  the  level  floor  pierced  by  the 
gastropore.  As  in  Astylus  isiibvirldis,-\-  the  upper  edge  of 
the  system  slightly  overhangs  the  outer  wall,  and  shows 
marginal  indentations  corresponding  with  the  centres  of  the 
outer  ends  of  the  pseudosepta,  or  walls  between  neighVjouring 
dactylopores.  The  broken  edges  of  the  pseudosepta  are 
cellular  in  appearance.  The  number  of  dactylopores  in  each 
S3^stem  is  variable,  twelve  or  thirteen  usually  occurring. 
In  one  place,  where  a  small  branch  is  given  off",  a  gastropore 
is  in  the  axil,  and  four  dactylopores  are  placed  on  eacli  of 
the  opposite  sides,  none  being  found  on  the  sides  of  either 
the  main  or  the  secondary  branch.  In  another  case,  the 
cyclos3^stem  is  at  one  side  of  the  base  of  a  small  branch, 
slightly  below  the  level  of  the  axil,  so  that  the  position  of  a 


III,  p.  440.  +  Moseley,  op.  cit.,  p.  457. 


Oil  Two  Neiv  Tertlari/  Sfijlasterids,  111) 

branch,  in  reference  to  a  cyclosystem,  is  not  quite  constant. 
In  the  latter  case,  six  dactylopores  are  present.  In  botli 
tliese  cases  the  dactylopores  are  rather  indistuict,  and  the 
systems  are  possibly  in  process  of  obliteration,  such  oblitera- 
tion being  recorded  by  Moseley,  as  occurring  in  oldei"  portions 
of  colonies  oi'  £!rrina* 

The  gastropore  is  approximately  circular  in  outline,  slightly 
funnel-shaped  above,  and  cylindrical  below.  The  presence 
of  the  matrix  prevented  a  closer  examination  of  its  deeper 
parts. 

The  ampulla;  form  comparatively  large  hemispherical  pro- 
jections above  the  surface  of  the  coenosteum.  Their  position 
is  not  constant,  though  they  lie  near  the  base  of  a  cyclo- 
system. Some  systems  have  no  ampulla  near  them,  while 
one  has  two,  placed  one  on  each  side  of  the  base,  tho\igh  one 
seems  the  usual  number.  In  some  cases,  a  small  aperture 
at  the  base  leads  through  the  outer  wall  of  the  ampulla, 
which,  as  a  fractured  portion  shows,  is  thin.  In  one  instance, 
this  aperture  is  surrounded  by  a  slightly  projecting  neck, 
with  a  thick,  definite  wall.  The  only  external  openings 
leading  into  ami)ulliTe  mentioned  by  Moseley, -f*  occur  in  male 
colonies  of  Sporadopora,  where  the}^  are  small  and  olit-like, 
and  are  placed  at  the  bottom  of  irregular  depressions,  which 
are  seen  with  difficulty. 

In  Cryptokella  the  female  colony  has  but  one  ampulla 
associated  with  each  cyclosystem,  but  in  what  are,  perhaps, 
male  colonies,  several  ampullre  occur  with  each.^  In  the 
female  colonies  of  most  of  the  stylasteridtie  desctibed  by 
Moseley,  there  is  only  one  planula  developed  in  each  ampulla, 
and  tiie  only  means  of  escape  seems  to  be  by  absorption  and 
breaking  down  of  the  outer  wall  of  the  ampulla.§ 

In  Cryptokella,  however,  there  are  several  gonophores 
present  in  each  gonangium  in  all  stages  of  deve]opment.|| 
In  the  latter  case,  a  definite  constant  means  of  escape  for  the 
planulse  would  seem  to  be  necessary,  though  I  can  find  no 
mention  of  this.  The  large  size  of  the  ampullfe  in  the 
present  example  <d1  Deontopora,  would  be  a  strong  argument 
in  favour  of  the  colony  being  a  female  one,  and  the  ])resence 
of  such  a  definite  opening  into  the  ampullae  would,  I  think, 

*  Op.  cit.,  p.  444.  t  Op.  cit.,  p.  431. 

:  Op.  cil.,  p.  46H.  .  §  Id.,  p.  441. 

il  Op.  cil.,  p.  477. 


120      Proceed uif/s  of  tli.e  Royal  Society  of   Victoria. 

.show  tliut  it  would  be  requii-ed  frequently.  Possibly  then, 
vvc  have  in  this  instance  a  near  ap]»roach  to  the  conditions 
found  in  the  gonangia  of  Cryptohe/ld. 

When  other  specimens  of  this  stylasterid  are  available  for 
examination,  the  exact  position  of  the  genus  in  the  group 
will,  probably,  be  determinable.  The  suppression  of  some 
of  the  dactylozoids  of  a  c3'closystem  on  one  side  of  a 
gastrozoid,  occurs  at  times  in  Stylaster,  and  is  constant  in 
GryptoJielia.  In  the  latter  case,  however,  the  suppression 
is  accompanied  by  the  production  of  a  calcareous  lamina 
overhanging  and  pi'otecting  the  cyclosystem  ;  but  it  must 
be  noted  that  the  abortion  occurs  on  opposite  faces  of  the 
system  in  Cryjifohelia  and  Beontopora.  In  the  formei-,  it 
occurs  on  the  side  of  the  system  towards  the  proximal  end 
of  a  branch,  while  in  the  latter,  it  is  towards  the  distal  end. 

In  Astylus,  which  is  ])robably  the  nearest  ally  of  Crypto- 
hclia,  the  hoinologue  of  the  external  lid  of  the  latter  is  a  small 
tongue-like  projection  placed  somewhat  deeply  in  the  gas- 
tropore,  and  dividing  it  into  an  inner  and  an  outer  chamber.* 
Sections  would,  of  course,  require  to  be  rubbed  down  to 
settle  wlietiiei-  or  not  such  a  process  is  found  in  Deonfopora, 
or  whctlie)-,  on  the  other  hand,  a  style  is  developed,  in  the 
former  c;ise  its  nearest  ally  would  be  Asfylas,  and  in  the 
latter  Stylaster.  If  neither  stjde  nor  process  occur,  its 
affinities  would  be  with  (Jonopora .-^  The  external  characters 
point,  I  think,  to  an  alliance  with  Astylus  and  GryptoheUo. 

Locality. — Grey  clays,  Orphanage  Reserv^e,  Fyansford, 
Geelong.     Only  one  specimen  found. 

Genus  Lsptobotlu-as  (gen.  nov.) 

The  poi'es  are  grouped  in  cyclosystems  ;  dactylopores  not 
in  radial  grooves. 

The  absence  of  grooves  containing  the  dactylo[)ores  is  a 
feature  not  ap})arently  occurring  in  any  genei'a  in  which 
cyclosj'stems  are  found,  though  it  occurs  in  Sporadopor a  and 
Disticliopora. 


L.  spenceri  (s[).  nov.) 

The    specimen   tigiu-ed  consists  of  a  portion  of  a  branfli 
which  is  circular  in  section.     Length  b  mm.,  diameter  1  mm. 


Moseley,  op.  cit.,  p.  458.  t  Id-,  V-  503. 


On   Two  iS'eiv  Teiihu'H  Shjh(derid.s.  121 

The  regularity  of  its  forui  is  disturbed  by  cyclosy.steins, 
which  are  scattered  irregularly  over  its  surface.  The  surface 
is  marked  by  minute  pores,  which  are  slit-like,  oval,  or 
circular.  They  are  larger  and  more  distinct  than  the 
corresponding  pores  of  Deontojiora,  and  are  irregularly 
scattered. 

The  cyclosystems  appeal"  as  cylindrical  elevations  at  right 
angles  to  the  axis  of  the  branch,  and  irregular  in  position. 
The  gastropores  have  well  defined  walls  of  similar  texture  to 
the  surface  of  the  branch.  They  are  cylindrical  in  shape, 
and  maintain  the  same  diameter  right  to  the  top  of  the 
cyclosystem,  not  as  in  most  other  genera,  opening  into  a 
basin-shaped  depression.  The  dactjdopores  are  about  eight 
or  nine  in  number,  and  open  directly  on  the  surface  ot  the 
ring  which  forms  the  boundar}'  of  the  gastropore,  not  V)eing 
placed  in  radial  grooves.  The  apertures  are  about  midway 
between  the  inner  and  the  outer  walls.  No  styles  were  seen 
in  either  kind  of  pore. 

The  ampullae  are  not  noticeable  externally,  but  a  large 
distinct  pore,  with  a  slightly  expanded  external  opening,  is 
present  at  a  small  distance  below  the  cj'closystem  in  nearly 
every  instance.  In  Cryptohelia*  the  ampullae  are  always 
developed  in  connecticjn  with  the  cj-elosy-stems,  and  the 
invariable  presence  of  a  pore  in  this  position  in  the  present 
specimen,  renders  it  probable  tliat  it  leads  into  an  ampulla. 
One  or  two  pores  which,  though  somewhat  smaller,  have  a 
similar  appearance,  are  placed  without  relation  to  any 
cyclosystem.  Named  as  a  compliment  to  Professor  W. 
Baldwin  Spencer. 

Localities — A  well-sinking  in  the  Eocene  beds  at  Belmont, 
near  Geelong,  and  at  Schnapper  Point. 

My  thanks  are  due  to  Protessor  Spencer  for  suggestions, 
and  for  the  loan  of  works,  without  which  this  paper  could 
not  have  been  prepared. 

*  Moseley,  ojj.  cit.,  p.  477. 


122      Proeeedinc/.s  o/  tJte  Royal  Socletij  of  Victoria. 

DESCRIPTION    OF    PLATE. 

Fig.  1. — Deoiitopora  mooraboolensis,  enlarged. 

Fig.  2, — Two  ainpulLie  seen  somewhat  from  below. 

((. — Broken  end  of  a  small  bx-anch  showing- 
cellular  appearance,  due  to  the  cut 
ends  of  the  coenosai'cal  tubes. 

b.  h. — Ampul l?e,    the   one   on    the   left   being 

broken. 

c.  —  Pore  of  ampulla,   with    detinite    ring-like 

wall. 

d.  d. — Slit-like  pores  in  gi-ooves  on  surface  of 

coenosteum. 
e. — Pore  of  ampulla,  its  wall  being  broken. 

The  pai't  shown  in  this  Figui'e  is  seen  from  above  in  the 
lowej-  left-hand  corner  of  Fig.  1. 

Fig.  3. —  Diagram  of  cyclosystem  of  Deonto'pora. 

Op. — Gastropore. 

Dp. — Dactylopore  in  groove. 

Ps.  /-**%— Pseudose})ta. 
Fig.  4. — Leptobotkrus  spenceri,  enlarged. 
Fig.  o. — Diagram  of  cyclosystem  of  the  same. 

(t/). — Gastropore. 

Dp. — Dact3dopore. 


Fie,  1. 


Proc.  R-5.  Victoria  Plale 


s 


f 


f 


'  i^ 


^  rV'U\'i::;;;'iy 


Piy   4. 


Fig. 3. 


f      I 


RWenddUth.  C  Troedel iC? print. 


Art.     VIII. — Three,     rare   Species   of    E<j(js   liliherto   only 
described  from  the  Oviduct  of  the  Bird. 

By  A.  J.  Campbell,  F.L.S. 

(Coininuiiicated  by  Professor  W.  Baldwin  Spencer,  M.A.) 

[Read  September  8,  1892.] 

1.    Serici^^us  melinus,  Latham  (Regent-bird). 

Localiijj. — Somewhat  restricted,  being  ehietly  confined  to 
the  sub-tropical  coastal  scrubs  of  the  Northern  portion  of 
New  South  Wales  and  Southern  Queensland  ;  but  its 
extreme  limit  appears  to  be  Port  Jackson  in  the  South, 
where  the'  bird  has  been  occasionally  observed,  and  the 
Fitzroy  River  in  the  North.  I  have  recorded  having 
received  a  skin  of  a  young  male  from  Duaringa,  near  that 
Rivei-. 

Observations. — The  Regent-bii-d,  especially  the  adult  male 
with  glorious  black  and  yellow  plumage,  Gould  has  well 
said  is  one  of  the  finest  of  Australian  Fauna.  Last 
November  I  undertook  an  excursion  to  the  Richmond  River 
District,  New  South  Wales,  with  a  view  of  obtaining, 
amongst  other  items,  the  eggs  of  the  Regent-l)ird.  I  found 
the  luxuriant  scrubs  abounding  with  Regent-birds,  in  fact, 
they  were  as  plentiful  there  as  the  Wattle-birds  about  the 
Banksia  groves  of  our  Southern  coast.  We  experienced  no 
difficulty  in  yjrocuring  our  few  specimen  skins.  All  that 
was  necessary  was  to  select  a  balmy  day  and  recline  under 
a  Canthium  tree,  wliere  the  birds  (males  in  various  stages  of 
plumage  and  females)  came  to  regale  themselves  on  the 
bunches  of  hard  yellow  berries.  Nevertheless,  although 
well  aided  with  a  hardy  companion,  I  prosecuted  a 
vigorous  and  toilsome  search  through  dense  labyiinths 
of  hot  scrub  and  thorny  brakes  of  prodigal  growth, 
whei'e  the  thick  foliage  of  the  trees  caused  a  perpetual 
twilight  underneath,  but  returned  without  the  eggs.  It 
was  an  experience  akin  to  seeking  for  the  proverbial 
needle  in  a  haystack.     From  evidence  gained  by  dissection 


124      Proceedings  of  ihe  Royal  Society  of  Victoria. 

and  otherwise,  it  appears  that  November  was  too  early  for 
the  ninjority  of  the  birds.  However,  just  prior  to  leaving 
(19th  ^'ovember)  we  detected  a  female  carrying  a  stick,  and 
after  much  laborious  work  we  succeeded  in  tracing  her 
through  an  entanglement  of  wild  ras[)beri-ies  and  stinging 
trees,  and  were  satisfied  that  she  was  building  in  a  certain 
bushy  Buoyong  {Tarrietia)  tree,  after  seeing  her  return 
sevei-al  times,  each  time  with  a  twig  in  her  bill.  Marking 
the  tree,  we  {pointed  it  out  to  two  young  farmers,  directing 
them  to  send  the  eggs  after  us.  Some  weeks  afterwards,  I 
received  a  doleful  letter  stating  they  were  unable  to  climb 
the  tiee.  However,  the  next  month  another  farmer,  whose 
scrub  paddock  I  had  scoured,  following  up  my  instructions, 
fnind  therein  a  Regent's  nest  containing  a  pair  of  fresh  eggs, 
which  I  now  have  pleasure  in  describing. 

The  Eyg.s. —  {a)  A  beautiful,  well-shaped  s[)ecimen,  with  a 
fine  texture  of  shell  of  a  light  yellowish-stone  colour,  with  a 
fvint  green isli  tinge,  and  marked  with  blotches  and  spots  of 
sienna,  but  principally  with  hair-like  markings  of  the  same 
colour  in  I'anciful  shapes  and  figures,  as  if  a  person  had 
painted  them  on  with  a  fine  brusli.  Intermingled  are  a  few 
gre3dsh  streaks,  dull,  as  if  under  the  shell's  surface.  All  the 
markings  are  fairly  distributed,  but  are  more  abundant 
around  the  upper  quarter  of  the  egg.  The  dimensions  are 
1  cm.  long  hy  i\  breadth  of  2'8  cm.,  somewhat  large  compared 
with  the  size  of  the  jjarent.  The  character  of  the  markings 
resemble  nnich  the  egg  of  its  close  ally,  the  Spotted  Bower- 
bird  (Ohlamydodera  nuicidata),  which  1  ibund  near 
Wentworth,  River  Darling,  October  1887,  with  the  difference 
that  the  ground  colour  of  the  Regent  is  more  yellowish  and 
not  of  tiie  greenish  shade  of  the  Bower-bird,  (fj)  Similar  to 
the  other  s[)ecimen,  but  markings  less  pi'onounced  and  finer 
in  charactei',  with  a  g'eater  proportion  of  the  dull  greyish 
hair-like  streaks,  also  a  little  smaller;  length  .S-9o  cm.  by 
a  breadth  of  2  75  cm. 

The  jS'ed. — It  was  discovered  during  the  last  week  in 
December,  was  placed  about  15  feet  from  the  ground,  and 
was  observed  by  the  bird  sitting  thereon.  The  structure 
was  of  such  a  loose  nature — merely  a  few  twigs  forming  a 
flat  shelf  about  five  inches  across — that  it  fell  to  pieces  on 
removal  from  the  tree.  It  was  accounted  remarkable  how 
the  eggs  could  retain  their  position  in  it.  The  description 
of  the  nest  verifies  the  statement  found  in  Gould,  that  "  it  is 
rudely    constructed    of    sticks  ;    no    other    material    being 


Three  Rare  Species  of  Efifj^.  125 

einploj'ed,  not  even  a  lew  i-oots  as  a  lining,"  Ijut  is  at 
variance  with  Mr.  North's  statement,  which  precedes  his 
description  of  the  egg  taken  from  the  oviduct  of  a  bird  by 
Mr,  C'ockerell,  the  collector,  the  only  other  egg  at  present 
known. 

Incidentally,,  the  sti'eaky  markings  of  the  eggs  o})en  np  a 
speculation  in  reference  to  coloration.  A  clever  paper  read 
before  this  Society  some  time  ago,  suggested  the  bowers  or 
pla}^  houses  of  the  birds  as  attributive  to  the  cause.  Perliaps 
Mr.  Lucas  had  in  his  mind  Jacob  and  the  flocks  that  con- 
ceived before  the  rods  and  bi'ought  fortli  cattle, 
"  ringstreaked,  speckled,  and  spotted."  But  all  bower- 
building  birds  do  not  la}^  streaky-coloured  eggs,  to  wit,  the 
Satin  Bower-bird  {Ptilonorlcy ncltits) .  I  succeeded  in  taking 
a  good  |)hotograph  of  the  Regent-bird's  bower. 

2.    ScYTHROPS    N()V.E-HoLLA.NDLE,    Latham    (Channel-bill) 

Locality. — This  bird  is  a  wanderer  over  the  whole  of 
Australia,  but  has  not  yet  been  received  from  the  South 
West  ])ortion,  and  sometimes  reaches  Tasmania.  Is  recorded 
from  New  Guinea. 

Observations. — Th.  Channel-bill  is  manifestly  interesting, 
because  it  is  the  lai'gest  of  Australian  Cuckoos.  It  is  some- 
times known  inland  as  the  "flood"  bird,  ai'riving  with  such 
occurrences.  Gould  described  an  egg  from  the  oviduct.  Mr. 
North  described  a  similar  immature  egg  from  a  bird  shot  on 
the  Macleay  River  during  the  first  week  in  November  1884. 
An  egg  collected  for  me  (taken  from  a  crow's  nest,  if  I 
recollect  rightly)  at  Coopei-'s  Creek,  was  unfortunately 
broken  in  transit. 

The  Egg. — A  mature  example,  in  tiie  possession  of  Mr. 
D.  Le  Souef,  Zoological  Gardens,  Melbourne,  may  be 
described  as  light  Vjufl'or  pinkish-brown  in  colour,  mediumly 
spotted  with  pinkish-red  and  chestnut,  with  a  number  of 
light  purplish  markings  under  the  shell's  surface.  In  shape 
and  in  general  coloration,  it  is  not  unlike  a  Strepera's  (Crow- 
shrike)  small  egg.  Texture  of  shell  a  little  coarse  ;  surface 
almost  lustreless  ;  length  -t-2  cm.  by  a  breadth  of  2'84  cm. 
The  egg  was  taken  in  October  1880,  near  Inglewood, 
Queensland,  where  the  Channel-bills  were  fairly  numei'ous, 
by  Mr.  Herman  Lau,  an  observing  naturalist,  and,  as 
remarkable  as  it  may  appear,  from  the  nest  of  the  Sparrow- 
hawk  {Accipiter),  together  with  an  egg  of  the  bird  of  prey. 


12(5      Proceeding:^  of  the  Roynl  Socieftj  of  Victoria. 

On  anotliei-  occasioi),  Mi-.  Lau  took  a  pair  of  Channel-bill's 
eggs,  together  with  a  ])air  of  the  common  magpie's 
Gi/mno)'hlng  tlhiceu),  all  fresh  from  the  nest  of  the  latter; 
wliile  the  previous  season  he  took  a  pair  of  young  Channel- 
bills  from  a  Strepera's  nest,  and  forwarded  them  to  the 
Queensland  Museum.  It  would  be  interesting  indeed  to 
learn  if  the  same  Channel-bill  deposit  two  eggs  in  the  foster 
bird's  nest,  or  were  they  laid  b}'  separate  birds. 

3.    LoPHOLAiM[Ts   Antarticus,    Shaw    (Topknot  Pigeon). 

Locality. — The  coastal  scrub  generally  from  Cape  York  to 
Gippsland  Lakes.  Occasional  stragglers  reach  Tasmania. 
This  handsome  bird  is  persistently  omitted  from  the 
Victorian  list,  notwithstanding  it  has  been  recorded  from 
Eastern  Gippsland,  and  I  possess  a  note  of  a  flock  having 
appeared  at  Tyrell  Creek,  near  Charlton,  ISovember  1889. 

Observations. — T  enjoyed  am])le  opportunity  of  ol)serving 
these  pigeons  at  home  in  the  Richmond  River  scrub  last 
November,  but  was  much  too  early  for  their  breeding  season. 
It  was  delightful  as  the  rising  sun  was  gilding  the  tops  of 
the  taller  trees  of  the  scrub  to  steal  along  the  leafy  avenues 
to  some  favoured  native  Tamarind  tree  {Diploglottis),  there 
to  watch  the  Topknot  Pigeons,  in  company  with  the 
gorgeously  dressed  Magnificent  and  Swainson's  Fruit 
Pigeons,  ravishing  the  agreeable  acrid  bunches  of  fruit. 
The  flight  of  the  Topknot  Pigeon  is  rapid  and  powerful.  At 
times  they  eongiegate  in  large  numbers,  hence  they  are  some- 
times called  "flock  "  pigeons  by  the  dwellers  of  the  scrub. 
The  egg  I  am  jiermitted  to  describe  is  in  the  collection  of 
Mr.  Le  Souef  It  was  taken  from  the  nest  at  the  end  of 
January  1887,  by  Mr.  Herman  Lau,  Vandilla,  Queensland. 
Incubation  was  about  a  week  old,  therefore  it  is  probable 
that  this  species  lays  one  egg  only. 

Tlce  Egg. — Is  dull  white,  somewhat  granulated.  In  shape, 
inclined  to  oval,  with  peculiarly  pointed  extremities, 
especially  the  smaller  end  which  nips  ofl'  suddeidy.  Length 
4-5  cm.  by  a  breadth  of  S'OO  cm. 

The  Nest. — The  bird  was  shot,  not  knowing  it  flew  from 
the  nest,  which  was  immediateh-  discovered  about  forty 
feet  from  the  ground  on  a  thick  branch  of  a  Eucalypt,  near 
the  outskirts  of  the  Bunya  Mountain  scrub.  Like  those  of  the 
majoritj'  of  pigeons,  the  nest  was  of  the  usual  scanty  nature 
of  coarse  sticks,  a  few  finer  inside. 


Art.  IX. — Notet^  on  the  Mode  of  Reprodactlov  of  Geone- 
■mertt's  austraHenNis. 

By  Arthur  Dendy,  D.Sc,  Univevsit}'  of  Melbourne. 

[Read  October  13,  1892. j 

In  the  paper  on  Geonemertes  austral  lens  Is*  which  I  had 
the  honour  of  reading  before  this  Society  hist  year,  I  shewed, 
by  anatomical  examination,  that  in  this  w^orni  the  sexes  ai'e 
not  united  in  the  same  individual,  but  that  distinct  males 
and  females  exist.  The  males,  however,  appeared  to  be 
much  less  common  than  the  females  and  the  single  one  which 
I  obtained  was  considerably  below  the  average  size,  though, 
from  the  insufficiency  of  the  data,  it  was  impossible  to  found 
any  generalization  upon  this  fact.  At  the  time  when  I  wi'ote 
I  had  made  no  observations  either  as  to  the  mode  of  copula- 
tion of  male  and  female  or  as  to  the  manner  in  wliich  the 
eggs  are  deposited.  Relying  upon  m}^  anatomical  investiga- 
tions, however,  I  ventured  to  indulge  in  certain  speculations 
on  these  points,  which  ai'e  contained  in  the  following 
paragraph  : — 

"  The  ova,  as  already  stated,  grow  to  a  very  large  size, 
Ineasuring  up  to  about  O'G  mm.  in  diameter.  It  seems  to  me 
ahiiost  impossible  that  they  should  be  discharged  through 
the  narrow,  preformed  genital  ducts.  I  believe  that  they 
escape  by  rupture  of  the  body  wall  and  that  the  ducts  merely 
serve  to  convey  spermatozoa  to  them.  That  these  ducts  do  so 
convey  the  spermatozoa  I  conclude  from  the  fact  that  I  have 
found  spermatozoa  in  them.  Probably  the  process  of 
fertilization  is  effected  by  the  male  crawling  over  the  female 
and  passing  out  the  sperm  as  he  crawls."-f- 

Since  this  was  printed  my  friends  Messrs.  C.  C.  Brittlebank 
and  H.  Giles  have  collected  specimens  of  Geonemertes 
australiensis  and  also  made  some  extremely  interesting 
observations  upon  the  method  of  copulation  and  egg-laying. 
I  have  to  thank  these  gentlemen,  not  only  for  an  account  of 
their  observations,  but  also  for  sending  me  the  specimens 

*  "  On  an  Australian  Land  Nemertine  {Geononcrtes  aii^traUensix,  n.  sp.)" 
Proc.  Koyal  Soc.  Victoria,  Vol.  IV,  N.S.,  p.  85. 
+  Loc.  cit.,  p.  115. 


12(S      Proceed  ill  (js  of  liie  Roijol  SocieJi/  oj    Victoria. 

n|»()ii  vvliicli  tliese  o1).servati()iis  wei'c  based,  so  that,  in  the 
case  of  the  egg-laying  liabite,  I  was  able  to  continue  the 
observations  on  1113^  own  account.  It  will  be  seen  in  the 
sequel  that  my  suggestion  as  to  the  manner  in  which  the 
eggs  are  discharged  from  the  body  was  incorrect,  while,  on  the 
(jtljcr  hand,  m_y  views  as  to  the  method  of  copulation  receive 
support.  The  new  oltservations  reieried  to  above  are  as 
follows: — 

On  the  22nd  of  May  last  Mr.  Brittlebank  found,  near 
Myrniong,  two  specimens  of  Geovemertes,  apparently  male 
and  female  in  copulation.  The  supposed  male  was  very 
much  smaller  than  the  female,  and  was  riding  on  the  back  of 
the  latter.  The  female  was  about  three-quarters  of  an  inchand 
and  the  male  only  about  one-quartei'  of  an  inch  in  length  when 
ci'awling.  Mr.  Brittlebank  observed  the  specimens  for  an 
hour  and  then  posted  them  to  me,  but  unfoitunately  tliey 
were  lost  in  the  post,  so  that  I  M^as  unable  to  determine  the 
sexes  by  microscopical  examination.  The  notes  and  sketches 
made  by  this  careful  observer,  however,  point  strongl}'  to 
the  conclusion  that  the  specimens  were  really  male  and 
female.  Again,  on  August  5,  Mr.  Brittlebank  wrote  to  me 
that  he  had  found  anothei'  })air  coujiled  and  he  adds  "  the 
male  onl}^  crawled  over  the  dorsal  surface  of  the  female." 
Unfortunately  tliese  specimens  also  are  not  forthcoming  for 
microscopical  examination,  but  Mr.  Brittlebank  informs  me 
that  he  watched  them  for  a  long  time. 

The  above  evidence,  though  not  absolutely  conclusive, 
points  stT'ongly  to  the  conclusion  thah  mj^  suggestion  as  to 
the  manner  in  which  the  eggs  are  fertilized  is  correct.  "We 
have  next  to  deal  with  the  manner  in  wliich  the  eggs  are 
deposi  ted . 

On  July  4th  Mr.  H.  Giles,  of  Oreekside,  Nar-Nar-Goon, 
found  a  very  fine  specimen  of  Geoiieoiiertes,  which  he  kept, 
intending  to  send  it  to  me  alive.  He  forgot,  it,  however,  for 
some  days,  and  meantime,  on  July  7th,  it  de])Osited  a  mass 
of  eggs,  and  on  the  J  3th  it  was  found  coiled  around  a  second 
mass  of  eggs.  On  the  loth  July  I  i-eceived  from  Mr.  Giles 
the  parent  worm  and  the  two  masses  of  eggs  which  it  had 
laid,  the  woi-m  apparently  in  good  health,  and  without  any 
signs  of  I'upture  of  the  body  wall,  and  still  containing  a 
number  of  eggs  visible  through  the  integument.  I  kept  this 
specimen  under  observation  foi-  a  long  time,  and  on  August  1  st 
found  it  lying  by  the  side  of  yet  a  third  mass  of  eggs  which 
it  had  evidently  just  deposited  underneath  some  moss  in  the 


Mode  of  Reprodncthnt  of  Geonemerte.^  australiensls.   120 

vivarium.  Tlie  parent  animal  survived,  apparently  in 
perfect  iiealtli  and  condition,  until  September  lOtli,  when  I 
killed  and  preserved  it  lor  future  reference. 

It  will  be  obsei-ved  that  all  these  three  lots  of  eggs  (whifh 
I  shall  describe  presently)  were  laid  by  an  animal  in 
ca|itivity,  and  if  this  wei'c  all  the  evidence  forthcoming  some 
critic  might  perhaps  suggest  that  the  laj'ing  of  the  eggs  was 
due  to  the  abnoi-mal  conditions  ot  life,  as  has  been  suggested 
in  the  case  of  Feripatus.  Fortunately,  howevei-,  about  the 
same  time  two  other  observers,  Mr.  Hennel  and  Mr.  Fiddian, 
found  similar  masses  of  eggs  in  a  state  of  nature,  which  they 
kindly  brought  to  me,  and  which  subsequently  proved  to  be 
undoubtedly  eggs  of  Geonemertes.  Mr.  Hennel  obtained 
his  specimen  on  July  18th,  in  the  damp  bark  of  a  gum  tree 
on  the  Dandenong  Creek,  and  Mr.  Fiddian's  specimen  was 
found  beneath  a  stone,  at  Creswick,  at  the  end  of  July. 

The  newly  deposited  eggs  of  Geoncnierfe.s  austraUensls 
are  opaque  spherical  bodies  about  O'O  mm.  in  diameter  and 
of  a  white  or  nearly  white  colour.  Some  thirty  of  these  eggs 
are  enclosed  together  in  a  sausage-shaped  mass  of  colourless 
transparent  jelly,  about  half  an  inch  in  length,  the 
individual  eggs  being  scattered  through  the  jelly.  The 
surface  of  the  gelatinous  matrix  is  smooth,  and  the  jelly 
appears  to  be  common  to  all  the  eggs,  instead  of  forming  a 
special  envelope  around  each,  as  in  the  case  of  frog-S}uiwn. 
One  such  mass  of  eggs  is  deposited  at  a  time,  and,  as  is 
evident  from  the  observations  recorded  above,  at  least  three 
can  be  deposited  in  succession  by  the  same  animal,  at 
intervals  of  several  days,  the  animal  itself  remaining  perfectly 
uninjured.  Hence  it  appears  almost  certain,  although  the 
actual  deposition  of  the  eggs  has  not  been  (observed,  that 
they  leave  the  body  separately,  each  by  the  narrow  duct 
which  leads  from  the  sac  or  capsule  containing  it  to  tlie 
exterior.  This  duct,  then,  appears  to  serve  both  for  the 
admission  of  the  speraiatozoa  and  for  the  extrusion  of  the 
fertilized  eggs.  The  source  (jf  the  gelatinous  material  in 
which  the  eggs  are  deposited,  and  also  the  manner  in  which 
the  whole  mass  is  moulded  into  shape,  have  yet  to  be 
discovered.  Probal^ly  the  animal  discharges  the  eggs  and 
pours  out  the  jelly  as  a  secretion  from  the  surface  of  the  body 
simultaneously.  If  this  were  done  while  the  animal  was 
slowly  crawling  along  the  result  would  certainly  be  one  of 
the  curious  egg-masses  described  above.  We  may  con) pare 
this  hypothotical  process  with  the  formation   of  the  slimy 

K 


130     Proccedinys  of  ike  Royal  Society  of  Victoria. 

track  wliicli  viiider  ordinary  circumstances  the  animal  leaves 
behind  it  wlien  it  crawls,  only  in  the  latter  case  the  secretion 
of  slime,  and  consequently  the  slimy  track,  are  continuous. 
1  do  not  mean  to  suggest  by  this  comparison  that  the 
gelatinous  matrix  is  identical  witli  the  ordinary  slime,  for 
that  I  think  highly  impi-obable. 

With  regai'd  to  the  development  of  the  eggs  my  observa- 
tions have  been  attended  with  very  little  success.  The 
o))acity  of  the  embryos,  due  to  the  pi'esence  of  a  large 
quantity  of  food  yolk,  renders  investigation  of  them  in  the 
living  state  extremely  difficult,  and  the  cutting  of  sections, 
which  I  also  attempted,  has  not  so  far  yielded  satisfactory 
results  either.  As  might  have  been  expected,  I  have  not 
foimd  any  ti'ace  of  the  Pilidium  stage  so  characteristic  of 
some  marine  nemertines.  80  far  as  I  can  judge  at  present 
the  development  appears  to  be  direct. 

On  August  Sfith  I  examined  some  of  tlie  embiyos  from 
the  mass  of  eggs  which  was  found  freshl)'  deposited  in  the 
vivarium  on  August  1st.  It  was  easy  to  distinguish,  two 
stages  of  development.  In  the  first  the  embrj-os  were 
spherical  and  eacli  enclosed  in  a  very  delicate  transparent 
membrane.  Each  was  about  0'6  mm.  in  diameter,  opaque  and 
solid-looking,  and  clothed  with  short  cilia.  They  revolved 
slowly  inside  their  delicate  envelopes  and  sometimes  slightly 
changed  their  shape.  In  tlie  second  stage  the  embryos  had 
emerged  from  their  delicate  envelopes  and  under  the 
microscope  they  slowly  crawled  about,  constantly  changing 
their  shape  in  an  amoeboid  fashion,  elongating  as  they  crawled. 
They  were  clothed  with  short  cilia  and  were  still  perfectlj' 
opaque.     No  eye  spots  were  yet  visible. 

The  next  stage  observed  was  in  the  mass  of  eggs  collected 
by  Mr.  Fiddian  at  Creswick,  some  of  which  I  examined  on 
August  26th,  about  a  month  after  they  had  been  found. 
When  removed  from  the  soft,  investing  jelly  these  embryos 
elongated  themselves  greatly  and  crawled  about  pretty 
freely,  much  after  the  fashion  of  the  adult  worm.  A  single 
pair  of  eye  s|)ots  was  visible  at  the  anterior  end.  Micro- 
scopic examination,  by  means  of  sections  and  otherwise, 
showed  that  the  proboscis,  alimentary  canal  and  nervous 
system  were  all  well  developed,  even  the  characteristic 
stylets  of  the  proboBcis  being  present.  Hence,  although  these 
animals  had  not  yet  left  the  investing  jelly,  the  development 
was  nearly  complete.  The  alimentary  canal  still  contained, 
however,  a  ver}^  large  quantity  of  unabsorbed  yolk-granules. 


Art.  X.—The  Bluff  at  Bariuon  Heads. 

(Witli  Plate  XIV.) 

By  G.  S.    Griffiths,  F.G.S. 

[Read  November  6,  1891.] 


This  bold  headland,  at  the  uioutli  of  the  Riv^er  Barwon, 
presents  some  features  of  geological  interest.  A  crag  of 
grey  sandstone,  it  owes  its  preservation  to  the  circumstance 
that  its  seaward  extremity  stands  upon  a  basement  of  hard 
lava,  which  rises  just  above  the  level  of  high-water.  The 
result  of  such  an  arrangement  of  the  rocks  is  illnstrated  bj- 
the  profiles  of  two  of  the  cliffs,  one  of  which  consists  wholly 
of  calcareous  sandstone,  the  lava  foundation  being  wanting. 

Where  the  base  is  of  lava,  as  it  is  in  the  cliff  at  B  in 
Section  A  B,  the  profile  has  an  inclination  of  about  45°,  and 
can  easily  be  scaled.  This  shows  that  the  rate  of  recession 
of  the  face  of  the  cliff  is  much  faster  than  that  of  the  foot. 
Now,  the  foot  of  such  cliffs  is  cut  back  b}^  both  the  sea  and 
the  weather,  while  the  face  is  cut  back  by  the  weather  alone. 
As  the  waves  and  weather  together  work  into  the  cliff  much 
more  quickly  than  the  unaided  atmospheric  agencies  can,  we 
seek  for  some  special  condition  in  the  cliff  itself,  to  explain 
the  slanting  profile,  and  we  find  it  in  the  toughness  of  the 
lava  base,  which  here  retards  the  encroachments  of  the 
waves. 

If  we  now  turn  to  the  cliffs  near  D,  we  see  that  one  is 
vertical,  and  another,  which  I  have  not  drawn,  is  deeply 
undercut  at  the  sea  level,  so  that  it  continually  falls  in  great 
slabs,  which  encumber  the  beach.  Here  the  entire  face  of 
the  cliff  is  of  homogeneous  material,  and  the  greater  wasting 
jiower  of  the  sea  over  the  atmosphere  shows  itself  in  the 
profile,  which  is  vertical  where  it  does  not  overhang  its  foot. 
The  aerial  destruction  is  not  less  here,  but  the  sea  scour  is 
much  greatei'.  Hence  the  difference  between  the  profiles  of 
the  two  cliffs. 

K  2 


1*32      Proceedings  of  the  Jio;/al  Society  of  Victoria. 

The  next  circumsiraiice  illusti-ated  by  the  local  features 
is  the  effect  of  a  lava  flow  upon  tlie  distribution  of  shallow 
v/ater  deposits. 

Tlie  tongue  of  rock  projected  in  a  molten  state  across  a 
submarine  plain  of  shifting  sands,  forms  a  permanent  ridge 
against  which  the  swift  currents  at  once  heap  up  bai-s  of 
sand.  When  these  become  very  thick,  so  that  the  lower 
portions  are  not  disturbed  for  long  periods,  the  base  of  the 
ma&s  may  become  cemented  into  hard  rock  by  the  percolation 
of  lime  in  solution,  or  from  the  moment  sand  is  heaped  over 
the  vincooled  lava,  the  gases  and  acidulated  waters  may  slake 
the  mass  into  compact  strata.  This  has  occurred  here,  and 
thus  a  spit  seems  to  have  been  formed,  over  which  is  spread 
a  bed  of  clay  which  may  be  volcanic  ash  decomposed  in  -situ. 
or  an  ordinary  littoral  deposit.  Upon  the  top  of  this  clay 
bed  is  a  very  horizontal  soil  bed  ;  just  such  a  sandy  loam  as  / 
is  now  seen  to  be  capping  the  clitfs,  very  fine,  and  darkened 
with  abundant  carbonaceous  matter.  The  next  stage  is  that 
this  land  surface — which  may  have  been  no  more  than  the 
muddy  fore-shore  of  the  Barwou,  or  Lake  Connewarre — gets 
covered  with  sand,  which  is  false  bedded,  and  as  far  as  I  can 
see,  unfossiliferous.  Whether  then  this  is  a  sedimentary  (ji- 
an  eblian  deposit  it  is  hard  to  say,  as  false  bedding  occurs  in 
rocks  originating  in  either  way.  There  aie,  however,  thin 
beds  of  watei'-worn  conglomerates  intercalated  between  these 
false  bedded  sandstones,  which  lead  me  to  believe  that  the 
coast  was  sinking  and  that  the  sands  were  spread  over  this 
sf»it  by  the  sea  currents.  The  old  land  surface  humus, 
although  it  has  been  compressed  by  the  overlying  sandstone, 
IS  still  about  two  feet  thick,  and  its  upjier  margin  is  very 
sharply  marked  off  from  the  dejx^sit  above  it.  This  latter 
rises  as  a  cliff  face  to  a  height  ot  from  seventy  to  ninety  feet. 
]t  is  divided  into  at  least  three  greater  divisions,  and  these 
again  are  resolvable  into  lesser  beds,  all  current  bedded. 
There  are  differences  to  be  observed  which  distinguish  the 
l;u-ger  masses  from  each  other.  The  middle  bed  at  one  part 
of  the  cliff  especially,  contains  so  much  lime  that  every 
piojection  of  the  rock  wall  carries  its  group  of  stalactites. 
At  a  considerable  height  up  the  cliff  face  there  is  a  bed  of 
conglomerate,  or  breccia,  marked  E  in  the  sections.  The 
stones  are  small  sized,  some  are  basaltic  pebbles  water-worn, 
the  rest  are  of  sandstone,  some  loUed  and  some  not,  many 
having  a  black  burnt  look.  The  whole  mass  is  very  strongly 
cemented  together    by  carborate  of  lime.     It  is  worthy  of 


Proc.   R  S    Victoria.  Plate  XIY. 


The  Bluff  at  Barwon  Heads.  133 

notice,  that  while  the  old  land  .surface  at  the  base  is  quite 
level,  the  conglomerate  bed  rises  from  A  to  B  in  that  section. 
Other  similar-  beds  lie  above  this  one,  none  lying  horizontal^. 

The  lava  flow  is  not  seen  to  the  west  of  the  Barwon  Head- 
land (B),  but  some  beds  of  clay,  of  volcanic  origin  probably, 
though  destitute  of  basalt  boulders,  are  to  be  seen  at  two 
levels. 

On  the  beach  at  t\  which  is  about  three  hundred  yards 
distant  west  of  the  last  clitf,  a  bed  of  rough  conglomerate 
is  being  broken  up  by  the  waves.  This  is  beneath  the 
horizon  of  the  lava  flow  at  B,  but  it  nevertheless  contains 
plenty  of  basalt  boulders.  Amongst  other  rocks  there  are 
quartz  pebbles,  and  broken  shells  are  plentiful.  The  stratum 
is  about  three  feet  thick,  ami  it  lies  between  beds  of  sand- 
stone. Where  the  scour  of  tlie  .sea  has  worn  down  the 
conglomerate  to  a  smooth  flat  floor,  its  varied  materials  set 
in  a  hard  matrix  give  to  it  the  appearance  of  a  gigantic 
brawn. 

The  Barwon  Head  beds  must  be  either  late  Tertiary  or 
Pleistocene.  When  they  were  forming,  Bass's  Strait  extended 
northwards  of  the  present  coastline.  Subsequently  the  area 
rose  again,  and  when  the  present  clifl"  tops  were  probably 
.seventy  to  ninety  feet  lowei'  than  they  are  now,  one  extension 
of  Lake  Connewarre  southward  covered  them,  and  the  silt 
from  its  floods  spread  their  soil  over  the  barren  sand  rock. 
More  tilting  up  of  the  strata  drove  back  the  lake  waters  to 
the  north  ;  or  it  may  be  that  the  Barwon  cut  through  the 
sandstone  ridge  that  lay  between  the  lake  and  the  sea,  and 
thus  let  out  and  lowered  the  waters  of  Connewarre,  until 
they  approximated  their  surface  levels  to  that  of  the  ocean 
into  which  thev  were  di.scharo;ed. 


Art.  XI.  -On  the  Conductivlf^/  of  o   Solution  of  Copper 
Sulphate. 

(With  Plates  XV  and  XVI.) 

By  W.  HuEY  Steele,  M.A. 

[Read  August  11,  1892.] 

The  following  observations  were  made  with  the  intention 
of  examining,  under  various  conditions,  the  conductivity  of 
a  salt  solution,  which  is  of  some  impo)'tance  at  present, 
owing  to  the  attention  being  paid  to  solutions  now  by 
Ostwald,  van't  Hoft,  and  others.  I  chose  copper  sulphate 
(CuSO^+S  HjO)  to  work  with,  as  that  was  the  most  con- 
venient. It  is  plentiful  and  easily  purified,  and  cojjper  is  a 
convenient  metal  to  use  for  making  the  electrodes. 

All  the  methods  of  measuring  electrolyte  resistance  by  the 
ordinary  Wheatstone  bridge  and  galvanometer  are  more  or 
less  unsatisfactory,  the  only  satisfactory  method  being  that 
suggested  by  Kohlrausch,  namely^  of  using  rapidly  alternat- 
ing currents  and  a  telephone,  instead  of  steady  currents  and 
a  galvanometer.  The  alternate  current  may  be  produced  by 
a  small  dynamo,  but  much  more  conveniently  by  an 
induction  coil  maintained  by  a  few  cells.  A  small  coil 
is  preferable  to  a  large  one,  as  the  statical  charge  on  the 
electrodes,  especially  it  they  be  small,  is  liable  to  introduce 
a  sei'ious  error,  besides  which  is  the  annoyance  of  receiving 
shocks  on  touching  exposed  ]mrts  of  the  circuit,  if  one 
works  with  such  high  E.M.F.'s  as  ai'e  produced  in  a  large 
coil.  The  coil  I  used,  when  maintained  by  fou^-  freshly 
charged  Grove  cells,  gave  a  spark  of  rather  more  than  a 
centimetre,  but  I  generally  used  a  much  weaker  primaiy 
current.  A  slide  wire  bridge  is  generally  recommended,  but  1 
found  a  resistance  box  more  sensitive  and  more  convenient. 
The  greatest  sensitiveness  I  ever  obtained  was  about  1  in 
1500,  that  being  with  a  resistance  of  1500  ohms.  The  dis- 
tribution of  resistances  which  is  most  advantageous  in  the 
arms  of  the  ordinary  Wheatstone  bridge  is  by  no  means  the 
best    in    Kohlrausch's   ari-aiiuement.     In    the    former,    it    is 


Conduct  I  vliy  of  a  Solution  of  Copper  Sidphate.    135 

necessary  to  ari-ange  the  aims  so  that  when  the  resistances 
are  balanced,  the  niaxiimnn  current  shall  pass  through  the 
galvanometer,  and  generally  the  variable  aim  can  be  so 
arranged  that  there  is  no  perceptible  deflection  of  the  needle. 
But  in  Kohlrausch's  method,  one  cannot  get  complete  silence 
in  the  telephones,  and  a  variation  of  say  1  per  cent,  is  moie 
noticeable  in  a  feeble  sound  than  in  a  loud  one,  and  so 
(unless  the  currents  be  very  weak)  the  arms  have  to  be 
arranged  to  send  the  minimum  current  through  the 
telephones.  This  method  also  differs  from  the  ordinary  in 
its  inability  to  measure  with  accuicicy  low  resistances,  less 
than  10  ohms,  neither  can  it  measure  very  high  resistances 
more  than  50,000  ohms,  although,  with  the  box  I  had,  I  conld 
otherwise  have  measured  1,000,000  ohms.  Where  one  tries 
to  measure  these  high  or  low  resistances,  it  is  found  that 
when  the  resistances  are  approximately  balanced,  it  takes  a 
considerable  alteration,  say  5  per  cent,  in  the  variable  arm,  to 
produce  any  perceptible  change  in  the  sound  in  the  telephone, 
and  when  the  change  is  produced,  it  is  not  so  much  a  change 
in  intensity  as  in  quality — it  almost  seems  like  an  alteration 
in  pitch,  though  that  could  not  be.  Besides  overcoming  the 
ditHculty  introduced  by  polarisation,  there  is  an  enormous 
advantage  in  Kohlrausch's  method  in  the  way  of  rapidity. 
Making  an  observation  is  the  matter  of  seconds,  instead  of 
minutes. 

The  cell  I  used  to  examine  the  effect  of  change  of  tempera- 
ture on  conductivity  was  a  glass  tube  (see  Fig.  1),  about 
20  cm.  in  length,  and  1  cm.  in  diameter,  slightly  bent.  The 
ends  of  tliis  fitted  into  twi^  flat  copper  cups,  with  holes  in  the 
sides,  fitted  with  slightly  conical  necks.  These  cups  were 
about  7  cm.  x  5  cm.  x  2  cm.  The  space  between  the 
glass  and  the  copper  necks  was  tightly  packed  with  loose 
hemp,  and  formed  a  perfectly  water-tight  joint.  Wires 
soldered  to  the  cups  gave  a  means  of  connection,  the  cups, 
or  rather  their  iuterioi-  being  the  electrodes,  the  surfaces 
exposed  to  the  solution  being  about  80  square  cm.  The 
cups  were  closed  at  the  tops  b}^  blocks  of  indiarubber  cut  to 
fit.  I  had  some  diflticulty,  however,  in  making  tliese  quite 
water-tight,  and  tried  several  methods  of  stopping  up  the 
cracks.  It  was -easy  enough  to  stop  them  at  ordinary 
temperatures,  but  the  difficulty  was  to  find  some  cement 
that  did  not  soften  at  100°  C.  Sealing-wax  and  putty  were 
among  the  things  I  tried,  but  neither  remained  water-tight 
at  100°  C.     A  solution  of  indiarubber  in  naptha  was  finally 


18o      Proceedinys  of  the  Royal  Society  of   Victoria.    . 

tried,  and  with  ccmplete  success,  and  I  found  the  wliole  cell 
now  water-tight  even  under  considerable  hydrostatic 
pressure.  Glass  tabes  were  passed  through  the  indiarubber 
blocks,  and  Liebig  condensers  were  attached  to  these  by 
pieces  of  indiarubber  tubing.  The  inner  tubes  of  the 
(U)ndensers  were  closed  at  the  top  b}^  corks.  The  condensers 
were  held  vertically  by  clamps,  and  the  cell  was  thus 
sus])ended.  It  was  inunersed  in  oil  to  a  depth  of  about 
()  cm.  (dotted  line  in  figure).  I  had  to  keep  it  hung,  as  my 
bath  was  copper.  I  tested  the  insulation  of  the  oil,  and 
could  get  no  current  through  a  very  slight  thickness  of  it. 
When  heating"  a  solution,  air  bubbles  began  to  form  aJfc 
about  75°  C.  The  bending  of  the  tube  was  to  allow"  the 
escape  of  these  when  they  became  large  enough,  as  well  as 
to  allow  the  steam  to  escape  more  readily  when  the  temi>era- 
ture  rose  to  boiling-point.  The  conden.sers  were,  of  course, 
intended  to  keep  the  solution  at  a  constant  strength. 
Observations  of  the  resistance  above  70°  were  made  after  the 
solution  had  been  well  boiled,  so  that  there  were  no  air 
bubbles  to  increase  the  resistance  of  the  system. 

To  observe  the  temperature,  I  took  a  glass  tube  of  the 
same  section  and  thickness  as  that  of  the  cell,  and  corking 
one  end,  I  partially  filled  it  with  the  same  solution  as  that 
with  which  I  was  working,  and  putting  the  thermometer 
into  this,  I  put  the  tube  in  a  slanting  ].)Osition  in  the  bath. 
Under  tliese  conditions,  I  considered  tliat  tlie  temperature  of 
the  solution  in  the  second  tube  ought  not  to  differ  much 
from  the  temperature  of  the  solution  in  the  cell.  For  extra 
security,  however,  I  always  kept  the  temperature  within  a 
degree  or  two  for  several  minutes,  and  within  i  degree  for 
about  half  a  minute  before  taking  a  reading  of  the  resistance. 
The  salt  I  used  was  ordinary  commercial  copper  sulphate 
which  I  purified  by  making  strong  super-saturated  solutions 
in  distilled  water,  and  taking  the  crystals  which  were 
deposited  before  the  .solution  became  cold.  I  obtained  the 
strength  of  each  solution  by  weighing  the  amounts  of  salt  and 
water  in  it,  and  checked  the  results  by  taking  the  density 
with  hydrostatic  balance,  using  a  glass  sinker,  and  then 
comparing  these  values  with  a  series  previously  obtained  and 
l>lMtted. 

I  made  a  very  great  number  of  obsei'vations  altogether, 
but  finally  have  drawn  my  conclusions  from  eight  sets, 
which  were  the  last  made,  and  on  which  I  spent  more  time 
and  pains  than  on  the  others.     In  the  results  which  follow. 


ConductiviUi  of  a  Solution  of  Copper  Sidpli.ate.     137 


T  is  the  temperature  centi<^ia(]e,  R  the  observed  resistance  of 
the  system  in  legal  ohms,  aiul  /.•  is  the  conductivit}',  i.e.,  the 
rrciproeal  of  the  specific  resistance.  Taking  «  as  the  specific 
resistance  r  =  ^^.  where  r  is  the  mean  radius,  and  I  is  the 
length  increased  by  "8  /■  at  each  end,  ^72  i«  a  constant  for 
tile  instrument  determined  by  measurement  once  for  all. 
Thus  k  = 


logl 


loo-  R  =  l-2:)4() 


j,  and  log 

log  R^  so  tliat  the  calculation  of  /•  from  the  observed  resist- 
ances is  veiy  simple.  The  following  tables  show  all  the 
observations  used  from  whicli  I  calculated  my  results  : — 


5-98 

R 


1155 

•0155i  ; 

887 

2025  i 

im 

2335  [ 

C7.9 

2615  ! 

629 

2855  j 

305   i 

589 

5a7 

3343  i 

537-5 

334   1 

5^0 

332   i 

0-18 

328   [ 

556 

323   1 

581 

309 

3-34  % 

2-51  % 

1^258  ; 

; 

T 

R 

1800 

-00997 

T 

li 

k 

7' 

R 

k 

18-9 

18-3 

2290 

-00784 

15-S 

4030 

•00445 

31 

1459 

•012.30 

:U) 

1837 

•00977 

35^2 

2800 

641 

15 

1225 

1465 

40 

1598 

•01123 

52 

2350 

764 

(iO 

1066 

1684 

50 

1410 

•01246 

65 

21^20 

847 

70 

999 

1796 

60 

1335 

1344 

80 

1990 

902 

80 

962 
928 

1866 
1934 

70 
81) 

1266 
1218 

1118 
1474 

100 

80 

1957 

917 

99-7 

929 

1932 

97 

1174 

1529 

90 

1915 

937 

98-6 

930 

1930 

94 

1175 

1528 

94 

1925 

932 

95 

932 

1926 

90 

11S2   ; 

1519 

97 

1915 

937 

92 

933 

1924 

85 

1192 

1506 

100 

1901 

944 

90 

935 

1920 

80 

1212  i 

1481 

97 

1903 

943 

87 

940 

1909 

1 

94 

1901 

944 

85 

943 

1903 

80 

957 

1875 

1 

138     Pfoceediiif/s  of  ike  RoyaL  Society  of  Victoria. 


•597  % 

■262  % 

T 

Ji 

Ic 

T 

A^ 

k 

DO 

3270 

•00549 

99 

6270 

•00286 

'.17 

3250 

552 

96^5 

6260 

287 

96 

3235 

555 

94 

6260 

287 

9;5 

3230 

556 

91 

6J80 

286 

89 

3230 

556 

86-5 

6330 

283 

87 

3235 

555 

80-5 

6430 

279 

84 

3250 

552 

70-3 

6740 

266 

82 

3260 

550 

IS  i 

13850 

1296 

80 

3280 

547 

30-5 

10870 

1651 

70 

3400 

528 

45 

8900 

2017 

16-3 

6650 

270 

60  1 

7550 

2377 

30-2 

5060 

355 

70  , 

7000 

2564 

46 

4130 

435 

80 

6550 

2740 

6'J 

3650 

492 

70  , 

6900 

2601 

70 

3380 

531 

I  had  now  to  find,  first,  the  law  of  variation  of  conduc- 
tivitj'-  with  temperature,  and  second,  its  variation  with 
streno-th  of  solution.  In  working  out  the  former,  I  took 
20°  C.  as  my  standard,  and  in  what  follows,  t  is  the  excess 
of  temperature  over  20°.  I  found  that  each  set  of  observa- 
tions was  given  within  the  limits  of  errors  of  observation 
by  the  foiinula  k^  =  h^^  (I  +  at  —  /3  t'^)  l\  and  l'^^  being 
the  conductivities  at  20°  +  f  C.  and  20°  C.  respectively.  Tn 
determine  a  and  /3  as  accurately  as  possible,  I  worked  it  out 
in  each  case  by  the  "  method  of  least  squares,"  working 
from  the  conductivities  at  20,  30,  40,  50,  00,  70,  80,  9('), 
and  100°  ('.  found  by  interpolation  from  the  results  given 
above. 

The  values  I  found  are  shown  in  the  foUowinu'  table  : — 


n 

h. 

a 

P 

(Solution 

Conceutration). 

25-7  % 

•0458 

•0254 

•000100 

9-24 

•0224 

•0237 

140 

5-98 

•0165 

243 

144 

3-34 

•0102 

211 

125 

2-51 

•00808 

221 

136 

1-26 

•00482 

231 

138 

•597 

•00293 

215 

115 

•262 

•00135 

220 

69 

Coiidactlult)/  of  a  Solution  of  Copper  Sulphate.     189 

From  this  table  it  will  be  seen  that  a.  and  /3  are  fairly 
constant  for  all  solutions,  though  perhaps  a  increases  slightly 
with  the  concentration.  The  errors  in ^ aie  too  great  and 
too  irregular  to  indicate  any  law  of  variation.  Assuming 
then  that  a  and  /?  are  constant,  we  find  the  mean  values  are, 
a  =  022!)  ;  (i  =  000121.  In  a  the  probable  error  of  the 
result  is  -OOOo^,  or  a  little  less  than  2|  per  cent,  of  the  whole. 
Although  the  values  of  a  and  /S  thus  found  give  the  conduc- 
tivity with  ftiir  accuracy,  yet  they  fail  in  one  particular.  It 
will  be  seen  on  examining  the  results  in  the  case  of  the  last 
two  solutions,  that  there  is  a  temperature  of  maximum  con- 
ductivity somewhere  between  90  and  100°  C.  In  previous 
experiments,  however,  I  got  maxima  between  90  and  100°, 
with  solutions  of  3  and  0  per  cent.,  it  being  very  marked  in 
the  latter  case.  It  is  possible  that  there  may  be  a  maximum 
in  every  case,  but  generally  above  100°  C,  and  that  its 
position  may  vary  considerably  with  veiy  small  impurities 
in  the  solution,  though  I  do  not  know  what  impurity  I  could 
have  introduced  in  the  one  case  and  not  in  the  other,  as  in 
each  case  I  used  water  distilled  in  the  same  way,  and  salt 
from  the  same  vessel. 

I  should  remark  that,  in  calculating  a  and  (3  in  the  case  of 
the  solutions  that  have  a  maximum  under  100°,  I  only  used 
the  results  between  20°  and  80°. 

It  now  remained  to  determine  the  law  connecting  con- 
ductivity and  concentration  {k  and  n).  After  trying  various 
formuhe  and  plotting  sevei-al  functions  of  k  and  n,  I  at  last 
suspected  that  k  varied  as  some  powei-  of  n,  and  on  taking 
logai-ithms  and  plotting  them,  I  found  the  resulting  curve 
very  nearly  a  straight  line,  the  deviations  from  it  being  sucli 
as  might  arise  from  errors  of  observation.  Putting  tc  ^  a  n'', 
we  have  log  A;  =  log  a  +  h  log  n.  This  is  a  very  simple 
form  to  work  out  by  "least  squares,"  and  I  found  the 
constants  were  a  =  -OOiOS,  b  =  -700,  the  avei-age  error 
being  34  per  cent.  The  general  expression  for  the  conduc- 
tivity thus  becomes  k  =  -00403  X  ii^'^  (1  +  -0229  t  - 
•000121  t^).  The  curves  I,  II,  and  111  show  the  relations 
between  the  conductivity  and  temperature  for  three  different 
solutions,  and  may  be  taken  as  typical.  The  curves  them- 
selves are  plotted  from  the  mean  values  of  the  temperatuie 
coefficients,  and  the  crosses  show  the  actual  observations. 
As  I  remarked  previously,  the  coefficients  are  probably  some 
function  of  the  concentration,  but  my  results  are  not  accurate 
enough  to  determine  it.      Curve  IV  shows  the  logarithms  of 


140     Proceedings  of  the  Royal  Society  of  Victoria. 


the  diffl^rent  values  of  tlie  conductivity  and  concentration  ; 
as  before,  the  curve  showing  the  mean  calcuhited  vaUies,  and 
the  crosses  the  observed  vakies. 

The  following  table  gives  the  conductivities  for  several 
concentrations  and  temperatures,  and  may  be  useful  for 
reference  : — 


CONCENTEATION. 

Tejip. 

•1% 

•5  % 

1% 

5% 

10% 

20% 

30% 

20 

•000690 

■00237  ! 

•0040;! 

•013W 

•0235 

•0398 

•0543 

30 

•000840 

403  i 

490 

168 

286 

484 

661 

40 

•000975 

335 

569  ' 

195 

332 

562 

767 

50 

•001090 

374  1 

6H5 

•218 

371 

628 

857 

60 

•00119 

4(.8  ! 

693 

238 

405 

685 

936 

70 

•00127 

436 

741 

254 

433 

733 

1000 

80 

•001.^3 

459  ! 

780 

267 

455 

771 

1050 

90 

1 

472 

800 

1090 

100 

1 

483 

819 

1120 

DESCRIPTION    OF    PLATES    XV  AND   XVI. 


cups ;  c  c,  indiarubber 
e  e,  Liebig  condensers ; 


Fig.   ]. — a,  glass  tube;  b  6,  copper 
blocks;  r?,  h«^iiip  Packing 
/,  level  of  solution  in  cell ;  (/,  level  of  bath   in 
which  cell  is  immersed. 

Fig.  2. — Curves  I,  II,  III,  showing  agreement  between 
mean  value  of  temperature  coefficients,  and 
values  in  typical  cases.  Abscissae  represent  tem- 
perature centigrade  ;    ordinates,  conductivity. 

Fig.  3. — Curve  IV,  showing  that  the  connection  between  the 
logarithms  of  tlie  concentration  and  conductivity 
is  linear,  and  consequently,  that  the  conductivity 
varies  as  a  power  of  the  concentration.  Abscissae, 
logarithms  of  conductivity;  ordinates,  logarithms 
of  concentration. 


Proc-  R  S     Victoria   Plate  XV. 


Frg.l 


Froc   R  S.  Victoria    Plate  XV/. 


^^-^ 

--r- 

.03 

^^ 

- 

^^ 

Ji^ 

^^ 

' 

__E— — r 

2 

0                         3 

0                        4 

C                      5 

0                      6 

0                         7 

0                       so                      9 

0                     !00 

Fig. 2. 


1 1 1 I I ^ 


Fig.  3. 


Art.  XU.—The  Lichens  of  Victoria.     Part  I. 
By  Rev.  F.  R.  M.  Wilson. 

[Eead  November  10,  1892.] 

INTRODUCTION. 

1.— Thk   Structure  ov  Lichens. 

Lichens  are  cellular  plants,  and  consist  generally  of 
thallus,  apothecla,  and  spermagones. 

1.  The  thallus  is  usuall}^  composed  of  layers  (a)  cortical, 
(6)  gonidial,  (c)  medullar^/,  and  (d)  hypothaUine. 

(a)  The  cortical  layer,  which  occurs  on  the  upper  surikce 
of  most,  and  also  on  the  under  surface  of  mau}^  lichens, 
varies  in  composition,  but  is  generally  formed  of  closely 
compacted  cellules.  It  varies  also  in  colour,  in  thickness, 
in  degree  of  tenacit}^  and  in  smoothness  of  surface.  The 
inferior  cortex  of  many  lichens  is  furnished  with  vhizincB,  or 
root-like  filaments,  by  which  the  plants  are  attached  to  their 
substratum. 

(6)  The  gonidial  system,  which  generally  occurs  immedi- 
ately under  the  cortex,  is  specially  characteristic  of  this  class 
cf  plants.  When  the  gonidial  cells  are  completely  filled 
with  bluish  or  olive-green  matter,  they  are  called  granula 
gonlma;  but  when  the  yellowish-green  contents  are 
suiTounded  by  a  hj-aline  .space  within  the  cell,  they  are 
called  gonidia.  Both  kinds  are  usually  sphteroidal.  In 
some  genera  the  gonidia  are  flat  and  irregular  in  outline, 
chroolepoid.  Some  lichens  are  chrysogonimic,  with  golden 
yellow  gonidia. 

(c)  Tiie  medulla,  which  is  found  beneath  the  gonidial 
sj'stem  of  many  lichens,  consists  of  colourless,  tubular,  and 
articulate  filaments,  more  or  less  closely  compacted  or 
interlaced. 

(d)  The  hypothallus,  which  is  spread  under  the  thallus  of 
some  lichens,  is  usually  (jf  a  dark  colour,  and  is  formed  of 
the  filamentous  growth  arising  from  the  germinated  spores, 
on  which  the  other  parts  of  the  thallus  are  deposited. 


142     Froceedinrjs  of  the  Boy  a  I  Sociefy  of  Victoria. 

2.  The  apotliecium,  or  reproductive  oi-gan,  coDsists  of 
(1)  an  exci]}ulum ,  either  pale  or  dark,  on  whicli  lies  (2)  the 
hyjjothechim,  also  either  dark  or  coloiu-less.  From  the 
surface  of  the  latter  rises  (3)  the  thechnrt  or  Itymenium, 
which  contains  tlteccv,  generally  sniroiinded  by  'jniva/physes 
or  club-shaped  filaments,  all  which  are  usually'  glued 
togethei-  by  the  (/elathut  liymenia.  The  surface  jormed  by 
the  conglutinated  apices  of  the  paraphyses  is  termed  the 
epitlieciwni  or  disk.  The  hypothecium  of  nucleated  apothecia 
is  styled  a  i^evithecmnt. 

The  various  forms  of  apothecia  are  («)  leco/norhie,  i.e., 
orbicular  and  bordei-ed  by  a  tli;  Uine  margin  ;  (h)  lecideine, 
i.e.,  orbicular  with  no  thalline  margin,  but  often  bordered 
l)y  the  edge  of  the  excipulum  or  hypothecium,  which  is 
called  a  proper  margin,  i.e.,  a  margin  propei-  to  the  apothe- 
cium  ;  (c)  graphicline,  i.e.,  like  writing,  iiTegular  in  form, 
l)ut  typically  naiTow  and  horizontally  lengthened  ;  and  {d) 
pyo'etwdiiw,  i.e.,  globular  and  nucleated. 

The  theca?  contain  spores,  usually  8,  but  sometimes  1,  or 
2,  or  4,  or  (3,  or  sometimes  innumerable.  The  spores  vary 
much  in  size  and  form  and  colour  and  contents.  Home  are 
divided  into  two  or  man}^  cells,  and  some  are  simple.  The 
outer  and  inner  walls  are  called  the  e^yispjore  and  evdospore. 

8.  The  spermagones  are  small  thalline  tubercles,  containing 
a  colourless  receptacle,  within  which  there  arise  minute 
filaments,  eithei-  simple,  when  they  are  termed  sferigmata, 
or  articulate,  when  they  are  termed  arthrosteri(/viata. 
These  filaments  carry  upon  their  apices  very  minute  bodies, 
called  spermatio,  which  are  of  various  shapes,  but  generally 
(ylind)-ical,  and  which  are  readily  separable  and  pressed 
foi'th  in  great  numbeis  through  a  pore  in  the  a[)ex  of  the 
spermagone.  The  spermatia  are  supposed  to  fertilise  the 
apothecia,  but  their  function  is  not  known. 

There  are  also  other  bodies  occasionally  found  on  the 
thallus.  Pycvidei<  are  small  organs  of  a  dark  colour, 
(■oniaining  filaments  called  hasidia,  which  bear,  singly  at 
their  summits,  minute,  generally  oblong,  bodies,  called 
dylospores.  These  are  by  many  authors  supposed  to  be  a 
sort  of  secondaiy  fructification,  and  by  others  affirmed  to  be 
minute  fungi.  Cyp)heJla;  are  minute  excavations  in  the 
under  surface  of  certain  lichens.  Their  function  is  unknown. 
When  the}'  are  ap])arently  filled  with  white  or  yellow 
powder,  they  are  called  ph<e;vdoryp)hella'.  CepliaJodia  are 
tubercles  of  various  shapes,  which  are  found  on  many  species. 


TJw.  Lichens  of  Victoria.  143 

and  contain  cells  or  granula  gonima.  Their  functions  are 
unknown.  Soredia  are  powdery  protrusions  of  gonidia  and 
I >f  portions  of  the  interior  through  the  surface  of  the  thallus. 
In  some  species,  the  npothecia  are  often  conv^erted  into 
soredia  and  are  sterile.  This  formerly  constituted  a  genus 
Vrriolaria,  from  which  this  formation  is  styled  vaviolarioid. 
The  surface  of  the  thallus  is  often  roughened  by  minute 
tlialline  excrescences,  which  formerly  gave  rise  to  a  genus 
Isidium,  by  which  name  this  formation  is  still  distinguished. 
The  obsolete  genus  Lepraria  was  formed  of  lichens  whose 
thallus  is  s'terile  and  pulverulent.  The  obsolete  genus 
Spiloma  has  been  foinid  to  consist  of  certain  small  fungi 
parasitic  on  lichens.  Various  other  foreign  growths  are 
occasionally  detected  on  tneir  thallus  and  apothecia.  Some- 
tijnes  minute  alga?,  or  fungi,  or  portions  of  mosses  will  come 
into  view  under  the  microscope;  but  their  structure  is 
exidentl}'  very  different  from  that  of  the  lichen  with  which 
they  are  found. 

2. — The  Usks  of  Lichens. 

Their  chief  design  in  nature  seems  to  be  to-  form  a 
vegetable  soil  for  the  growth  of  higher  plants.  It  is 
lemarkable  that  no  ])oisonous  principle  has  ever  been 
found  in  any  species  of  lichen  ;  yet,  with  the  exception 
oi  a  few  kinds,  they  are  seldom  eaten  by  animals.  Snails 
devour  them,  and  there  are  microlepidopterous  larvae  which 
feed  upon  them.  On  a  coi-al  island  in  the  Indian  Ocean, 
I  found  lichens  with  the  traces  of  the  claws  of  crabs,  which 
had  evidently  sought  nourishment  from  them,  especially 
from  their  apothecia.  'J^he  omnivorous  Acarus  destructor 
seems  to  infest  lichens,  both  in  the  field  and  in  the  herbarium. 
.Some  species  afford  nourishment  to  the  higher  animals. 
What  is  called  the  reindeer  moss  is  a  lichen,  and  is  well- 
known  as  the  chief  food  of  the  reindeer  in  Lapland.  There 
are  also  species  in  other  lands,  which  are  useful  as  fodder  for 
domestic  animals.  Even  human  beings  occasionally  use 
some  kinds  for  food,  others  are  employed  for  medicine,  and 
others  again  have  proved  valuable  for  the  dyeing  of  cloth. 


3. — The  Geographical  Distribution  of  Lichens. 

The  growth  of  lichens  in  Victoria  depends  chiefly  on  the 
moisture  of   the   various   districts   of  the  colony.     On  the 


1+4      Proceed iv(/s  of  the   Royal  Society  of  Victoria. 

plains,  and  even  on  the  hills,  north  of  the  Dividing  Range, 
they  are  much  fewer  than  on  the  southern  slopes  of  tlie 
Range,  and  on  the  hills  and  in  the  forests  towards  the  coast. 
Suh-A]i)ine  species  are  found  on  the  lofty  mountains  of 
the  Victorian  Alps,  and  sub-tro])ical  species  in  east  Gip])s- 
land.  This  latter  district,  indeed,  from  its  position  near  the 
warm  currents  of  the  Pacific  Ocean,  and  sheltered  from  the 
.Southern  Ocean  by  Tasmania,  is  physically,  rather  a  part  of 
New  South  Wales,  thari  a  province  of  Victoria. 

The  annual  firing  of  the  forests  has  destroyed,  and  is 
destroying,  many  spots  which  used  to  be  good  collecting 
grounds.  As  lichens  live  only  by  the  influence  of  air 
and  moisture,  their  growtli  is  intermittent  ;  and  man 3^  of 
them  increase  verj^'  slowly,  pi'obably  continuing  in  life  foi' 
centuries,  and  their  reproductive  organs  are  supposed  to 
remain  fertile  for  ages.  The  destruction  of  such  plants  is  a 
loss  which  cannot  be  readily  made  good.  Many  of  them 
are  of  more  rapid  growth,  and  some  of  them  are  ainiual. 

In  favoui-able  localities  they  are  found  on  the  earth,  on 
stones,  on  rocks,  on  the  bark  and  leaves  of  trees,  and  on 
other  plants,  even  on  other  lichens,  on  dead  wood,  on 
decayed  moss,  on  fallen  leaves,  &c.,  on  dry  bones,  on  leather, 
on  glass.  Some  prefer  one  substratum  and  some  another, 
and  some  grow  indifferently  on  an}'.  Some  saxicole  lichens 
gi'ow  only  on  calcareous  rocks,  some  on  siliceous  rocks,  and 
some  on  any.  Certain  kinds  love  the  mountain,  and  some 
grow  oidy  on  Alpine  or  sub  Alpine  heights  ;  others  love  the 
plain.  Some  grow  only  in  wet  places,  others  in  the  dens«^ 
sunless  shade,  either  on  trees  or  in  caves,  or  under  ovei- 
hanging  rocks.  Some  like  the  neighbourhood  of  the  sea. 
othei'S  of  rivers  or  lakes.  Some  live  in  the  water,  either 
constantly  or  occasionally  submerged,  in  the  channels  of 
streams,  or  on  the  seashore  ;  and  some  flom'ish  on  the  slate 
roofs  of  houses. 

4. — The  Collecting  <jf  Lichens. 

The  collecting  of  lichens  is  best  done  in  tine,  but  not  too 
dry,  weather.  Those  which  grow  closely  attached  to  the 
bark  of  plants,  may  be  secured  by  cutting  off  the  bark  with 
a  strong  sharp  knife.  If  a  piece  of  wood  is  taken  along 
with  th.e  bark,  so  much  the  better,  as  it  v/ill  prevent  the 
bark  curling  up  when  dried.  The  tough  timl)er  of  our 
fences   and   decorticated    hardwood    trees,    on    which    some 


Tlie  Lichens  of  Victoria.  145 

grow,  will  need  the  use  of  cold  chisel  and  hammer.  Rocks, 
especially  granitic  and  basaltic,  require  a  light  well-tempered 
steel  chisel,  by  which  tolerably  thin  pieces  can  be  detached 
from  almost  any  rock  by  the  exercise  of  a  little  skill. 

All  that  is  necessary  to  preserve  the  specimens  is,  to  fold 
them  up  at  once  in  soil  ])aper  (newspaper  will  do),  to  prevent 
them  rubbing  against  one  another  in  the  bag  in  which  they 
are  carried  home.  Those  which  grow  on  earth  require  more 
careful  management.  They  need  to  be  collected  with  a 
sufficient  piece  of  earth,  and  tenderly  wrap})ed  up.  When 
brought  home,  the  earth  needs  to  be  pared  off  under  the 
specimen  to  a  level  surface,  and  then  solidified  by  the 
application  of  a  solution  of  isinglass  in  spirits  of  wine.  The 
solution,  when  liquefied  in  a  bottle  under  a  heat  of  25° 
to  80^  C,  or  77°  to  86°  Fahr.,  is  dropped  with  a  camel  hair 
pencil  on  to  tlie  earth  till  saturation,  taking  care  not  to  let 
it  touch  the  thallus,  which  it  would  discolour.  It  should  he. 
applied  underneath.  When,  after  a  day  or  so,  the  earth, 
thus  saturated,  has  become  dry  on  the  surface  by  exposure 
to  the  air,  the  specimen  should  be  placed  for  a  few  days 
under  sufficient  pressure  to  keep  it  in  shape  ;  it  will  thus 
harden  into  a  form  suitable  for  glueing  on  to  jniper,  as 
described  below. 


5. — The  Herjurium. 

The  mounting  and  arrangement  of  lichens  will  be  most  con- 
veniently carried  out  by  glueing  each  specimen  (with  Russian 
glue)  on  to  the  centre  of  a  piece  of  writing-paper,  with  a  space 
below  to  record  the  name  of  the  ])lant,  the  substratum  on 
which  it  was  found,  the  place  and  date  of  finding,  and  the 
name  of  the  collector,  and  with  a  space  above  to  record  notes 
of  examination.  These  pieces  of  writing-paper  may  be  then 
pinned  at  each  end  on  to  quarto  single  sheets  of  white 
cartridge  paper  with  "  lill  "  |)ins,  six  specimens  of  the  same 
species  to  tlie  sheet,  if  small,  or  two  if  larger.  They  can 
thus  be  easily  detached  for  special  examination.  The  largest 
specimens  may  be  glued  on  to  the  cartridge  paper  itself 
These  sheets  of  specimens  should  be  enclosed  in  a  quarto 
cover  of  ca.rtridge  paper,  one  species,  or  even  one  variety  to 
each  cover,  and  the  covers,  put  loose,  with  the  open  side 
inwards,  into  a  quarto  book  covei  of  pasteboard  (three  ply), 
joined  together  by  a  strip  of  strong  white  binder's  cloth,  of 
such  width  that  each  cover,  when  filled,  is  two  inches  thick. 

L 


146     Proceed'nigH  of  the  Royal  Society  of  Victoria. 

For  convenience  of  moving  them  to  fumigate,  &c.,  the  book* 
should  be  arranged  in  open  boxes  in  an  upright  row.  The 
most  convenient  size  of  box  is  that  of  J.D.K.Z.  Geneva  cases. 
The  boxes  being  arranged  like  shelves,  the  names  of  the 
family,  series,  genus  and  sub-genus  are  written  in  large 
characters  on  the  backs  of  the  bo!ok  covers,  and  thus  at  a 
glance  down  the  herbarium,  the  needed  book  can  be  readily 
seen  and  easily  taken  out.  The  names  of  the  specie.'i 
enclosed  in  the  books  should  be  written  on  the  left  hand 
lower  corner  of  each  doubled  cover,  and  thus  any  specimen 
can  be  conveniently  found  and  re]')laced  without  delay. 

To  preserve  tlie  specimens  from  the  ravages  of  insects, 
they  need  to  be  occasionally  exposed  to  the  fumes  of  bi- 
sidphide  of  carbon  in  a  covered  water-tight  case.  The 
quantity  of  fluid  required  depends  on  the  completeness  with 
which  the  case  is  tilled  by  the  boxes.  A  few  ounces  in  a 
small  c\\\)  will  serve  for  a  case  measuring  inside  3  ft.  x  2  ft. 
X  1  ft.  8  in.,  which  will  contain  three  boxes. 


6. — The   Examination    of   Lichens. 

The  examination  of  lichens  for  ordinary  purposes  is  most 
simply  and  expeditiously  carried  out  b}^  detaching  a  small 
portion  of  thallus  or  apothecium,  or  a  spermagone,  and 
]>utting  it  with  a  drop  or  two  of  water  on  a  glass  slide  for  a 
short  while  to  soak,  then  bruizing  it  down  gently  with  a 
pen-knife,  till  it  is  apparently  dissolved.  A  dry  cover  is 
applied,  and  gently  pressed  down  with  a  dry  knife.  The 
slide  is  then  put  under  a  micro.scope  having  a  good  |  inch 
object  glass,  and  an  eye  piece  magnifying  from  250  to  300 
diameters.  When  more  careful  examination  is  needed  for 
drawings  of  structure,  a  fine  section  will  need  to  be  made 
of  the  moistened  apothecium,  &c.,  M'ith  a  section  cutter,  or 
with  a  sharp  surgeon's  knife,  under  a  watchmaker's  lens. 
It  will  require  great  nicety  to  make  a  good  section,  neither 
too  thick  and  opaque,  nor  too  thin  and  depri\'ed  of  large 
spores.  Drawings  and  measurements  may  be  made  with  a 
camera  lucida  and  a  micrometer.  A  home  made  camera 
lucida  can  be  easily  constructed  by  cementing  a  half  of  a 
glass  cover  on  to  the  end  of  a  thin  plate  of  brass,  having  at 
the  other  end  an  aperture  to  correspond  with  that  of  the 
eye  piece,  and  bent  in  the  middle  at  an  angle  of  45 
degrees. 


The  Lichens  of  Victoria.  147 

The  chemicnl  le-agents  used  in  examining  specimens  are 
tlecrieil  by  some  liclienologists  as  being  unreliable.  Tliey 
are,  however,  v'aluable  assistants  in  determining  species, 
although  they  may  not  be  absolutely  conclusiv^e  taken  b}'- 
themselves.  The  usual  formula  by  which  the  solution  of 
iodine  (signified  by  the  lettei- 1),  is  prepared  is — iodine,  1  gr., 
iod.  potass.,  3  grs.,  distilled  water,  |  oz.  For  all  practical 
purposes,  however,  a  strong  enough  solution  is  made  by 
j)utting  a  few  grains  of  i(xline  into  a  small  phial  of  water 
and  allowing  it  to  stand  a  day  or  so.  The  solution  needs  to 
be  kept  in  a  glass-stoppered  bottle  of  dark  colour,  or  covered 
with  tin  foil  to  exclude  light.  Hypochlorite  of  lime  (signi- 
fied by  the  letter  C),  is  prei-»ared  by  putting  a  small  j»ortion 
of  chloride  of  lime  into  a  phial  of  water,  and  shaking  it. 
When  the  fluid  clarifies,  it  is  read}^  for  use.  Hydrate 
of  potash  (signified  by  the  letter  K),  is  composed  of  equal 
weights  of  water  and  caustic  potash.  It  may  be  well 
to  intorm  the  beginner  that  when  the  water  is  added 
to  the  caustic  potash,  a  good  deal  of  heat  is  evolved.  It  is 
well,  therefore,  to  previously  warm  the  bottle,  lest  the 
sudden  heat  should  break  it.  The  supply  of  caustic  potash 
needs  to  be  kept  from  the  air  by  beeswax  round  the  stopper 
of  the  bottle  in  which  it  is  preserved.  The  hydrate  should 
also  be  kept  in  a  stoppei-ed  ]ihial,  and  must  be  used  care- 
fully, as  it  corrodes  clothing,  »Scc.  These  re-agents  may  be 
applied,  a  drop  at  a  time,  by  means  of  thin  rods  of  glass ; 
keeping  each  rod  for  its  own  solution,  and  wiping  them  dry 
on  an  old  rag  after  using  them. 

The  application  of  C  and  K  is  either  to  the  surface  of  the 
plant  or  to  the  medulla.  The  younger  part  of  the  thallus  is 
the  best  for  examination.  In  cold  weather,  a  little  heat 
needs  to  be  applied  to  hasten  the  action.  This  may  be  done 
either  by  placing  the  ])hial  with  the  solution  in  a  cup  of 
warm  water,  or  by  putting  the  ])art  under  examination  close 
to  the  mouth  and  breathing  heavily  and  repeatedly  on  it 
after  touching  it  with  the  solution.  First  apply  C  to  a 
portion  of  the  thallus,  and  note  the  result.  Then  to  another 
portion  apply  K,  and,  after  watching  the  effect  a  short 
while,  add  C  and  note  the  results.  To  examine  the  action 
on  the  medulla,  scrape  off'  a  portion  of  the  cortex  from 
another  part  of  the  thallus,  and  apply  K  and  C  in  the 
same  manner.  The  more  freshly  made  the  solutions  are, 
and  the  more  carefully  they  are  kept  from  the  air,  the  more 
reliable  are  the  results. 

L  2 


]4<S     Proceedings  of  the  Roijal  Socieft/  of  Victoria. 

7. — The   Classification  of   Lichens. 

The  dassificatiou  of  lichens  adopted  in  the  following 
])ages,  is  that  of  Njdander,  as  the  most  natural,  being  based 
upon  the  consideration  of  all  tlie  parts  and  organs  of  the 
plants,  and  exhibiting  their  place  in  reference  to  the  neigh- 
i)ouring  classes  of  Algas  on  the  one  side,  and  Fungi  on  the 
other. 

8. — The   History  of   Victorian   Lichenology 

Begins  with  this  century.  The  first  lichens  collected  in 
Victoria  ai-e  recorded  in  an  a})pendix  to  Flinders'  Voyage 
to  Terra  Austraiis,  published  in  1814.  The  collection  was 
made  in  various  parts  of  Australia  and  Tasmania  by 
Mr.  Robert  Brown,  who  accom|)anied  Captain  Flinders  in 
his  investigation  of  the  coasts  of  New  Holland  in  1802. 
Brown's  specimens  were  afterwards  re-examined  by  Rev.  J. 
M.  Crombie,  and  the  result  recorded  in  the  Jouinal  Lin. 
Soc,  1880. 

In  1848  and  1849,  Dr.  Ferdinand  Mueller,  now  Baron 
von  Mueller,  collected  a  number  of  lichens  in  Victoria,  and 
sent  them  to  Dr.  Hampe,  who  determined  the  species.  The 
list  appeared  in  the  Report  of  the  Government  Botanist  to 
the  Victorian  Council,  1854.  A  second  parcel  of  specimens 
collected  in  Gippsland  and  the  Australian  Alps,  was  sent  to 
Dr.  Hampe,  and  enumerated  by  him  in  Schlechtendals 
Linnpea,  1856.  This  list  was  transcribed  into  the  Govern- 
ment Botanist's  Report  to  the  Victorian  Legislative 
Assembly,  1858.  These  namings  by  Hampe  need  revisal, 
in  view  of  the  more  minute  examination  of  later  lichen- 
ology. 

A  few  lichens  collected  by  a  visitor  from  Glasgow,  Mr. 
Hugh  Paton,  were  named  by  Dr.  Stirton,  and  published  l)y 
him  in  the  Proceedings  of  the  Royal  Society  of  Victoria, 
September  1880.  They  are  five  in  number,  and  all  new  to 
science. 

Collections  have  been  made  bv  Messrs.  R.  Wilhehni, 
D.  Sullivan,  C.  Walther,  MerralC  C.  French,  and  Mrs. 
McCann,  and  forwarded  by  Baion  von  Mueller  to  Europe. 
The  earlier  collections  were  sent  to  Dr.  Krempelhuber,  of 
Blankenberg,  on  the  Hartz  Mountains,  by  whom  their 
names  and  the  descri])tions  of  new  species  weie  ]iublished 
in  Ben  Verhandl.  des  Kois.  Keen.  Zool.  Bat.  OeseL  in  Wien., 


The  Lichens  of  Victoria.  149 

1880.  A  list  of  the  names  was  printed  in  the  Supplement 
to  the  eleventh  volume  of  the  "  Fragmenta  Phyt.  Austral." 
Authentic  named  sj)ecimens  of  most  of  these  are  pre.served 
in  the  Melbourne  Botanic  Museum.  These  determinations 
of  Krempelhuber  have  been  revised  by  Professor  Jean 
Mueller,  of  Geneva,  in  the  Ratisboii  Flora  or  Bot.  Zeit. 
1887.  The  later  collections  received  by  Baron  von  Mueller 
were  sent  to  Professor  Mueller,  by  whom  their  names  and 
the  descriptions  of  new  species  are  recorded  in  the  Ratisbon 
Flora  from  time  to  time.  Authentic  named  specimens  of 
most  of  them  are  preserved  in  the  Melbourne  Botanic 
Museum,  and  a  list  of  those  named  from  1881  to  1887 
is  given  by  Baron  von  Mueller  in  the  Victorian  Naturalist, 
October  1887. 

Collections  made  by  Miss  F.  M.  Campbell  (now  Mrs. 
Martin),  by  Mr.  F.  Reader,  and  by  Rev.  F.  R.  M.  Wilson 
were  sent  for  determination  to  Dr.  C.  Knight,  of  New 
Zealand.  Subsequently,  some  have  been  named  and  des- 
ciibed  by  Rev.  F.  R.  M.  Wilson,  and  lists  of  them  have 
appeared  from  time  to  time  in  the  Victorian  Naturalist, 
October  1887,  June  1888,  August  and  September  1889,  and 
April  1890  ;  and  latterly  many,  especially  of  the  crustaceous 
kinds,  have  been  submitted  to  Professor  Mueller,  whose 
determinations  have  not  yet  been  published. 

In  1891,  a  paper  entitled  "Lichens  Collected  in  the 
Colony  of  Victoria,  by  Rev.  F.  R.  M.  Wilson,"  was  published 
})y  the  Linntean  Society  of  London.  Many  of  the  names 
and  descriptions  there  given  are  reproduced  in  the  present 
paper,  but  some  are  altered.  The  alterations  of  names  are 
noted  in  each  case. 

Those  localities  to  which  no  name  is  appended  haA'e  been 
ascertained  by  the  author. 


CLASS   LICHENES.     Micheli. 

Thullus  containing  gonidia  or  granula  gonima  variously 
disposed,  and  very  often  also  crystals  of  oxalate  of  lime. 
Fructification  consisting  of  spores  in  theca3;  gelatina  hymenia 
in  most  species  becoming  blue,  in  others  reddish,  and  seldom 
unaffected  by  the  application  of  an  aqueous  solution  of 
iodine.  S])ermagonia  in  minute  thalline  tubercles  distinct 
from  the  apothecia. 


!•")()      Proceedings  of  the  Royal  Society  q/   Victoria. 
FAMILY   I.-COLLEMACEI. 

Thallus  usually  dark  in  colour,  black,  brown  or  olive, 
sometimes  ashy  or  bluish,  various  in  form,  gelatinous  in 
substance,  enclosing  granula  gonima,  which  are  variously 
arranged,  moniliform  or  enclosed  in  sacs  or  dispei'sed. 
Apothecia  usually  rufescent  or  pale,  seldom  black,  genei-ally 
lecanorine  or  biatorine,  rarely  endocarpoid. 

Tribe  I. — Lichinei. 

Thallus  blackish  or  brown,  small,  filiform,  caispitoso-fruti- 
culose  or  depresso-radiate.     Saxicole. 

Genus  1. — Ephebe,  Fr.  Born. 

Tiui-llus  fruticulose,  filiform,  branched  and  entangled  ; 
granula  gonima  large,  arranged  chiefly  under  the  cellulose 
coi-tex  sub-transversely,  two  or  four  or  more  together. 
Apotliecia  endocarpoid  in  thickened  poi'tions  of  the  thallus. 
Spermatia  cylindrical. 

1.    E.  i^uhesceiifi,   Fr. 

Thallus  blackish  brown,  small  (about  3  millimetres  higli, 
■1  mm.  thick),  much  branched,  somewhat  decumbent,  sligiitly 
rugulose,  containing  brownish  green  granula  gonin)a.  Dia^- 
cious.  Spores  8,  colourless,  oblong,  simple  or  1  septate, 
•on  to  OIG  X  -003  to  -OO-t  mm.  (Nyl.)  Paraphyses 
indistinct. 

Hab.  on  sub-Alpine  rocks.  Mount  Macedon.     Sterile. 

Previously  named  by  me  (Trans.  Lin.  Soc.  1890)  Stirjo- 
nema  ephebioides,  Wilson,  from  a  few  small  imperfectly 
developed  specimens.  The  lenticular  con-colorous  bodies 
then  noted  by  me  as  apparently  connected  with  the  plant, 
were  possibly  foreign  to  it. 

Genus  2.— Lichina,  Ag. 

Thallus  brownish  black,  fruticulose,  firm  ;  granula  gonima 
bluish ;  apothecia  terminal  in  sub-globose  open  thalline 
receptacles.  Spermatia  oblong.  Spores  8,  colourless,  ellijj- 
soid,  simple. 


The  Lichens  of  Victoria.  151 

1.    L.  pygmcea,  Lightfoot. 

Thallus  small  (i  inch  or  a  little  more),  branches  flattened 
towards  the  apices.  Spores  -022  to  029  X  -Oil  to  01 G  mm. 
'^y\.     Gelatina  hymenia  unaiiected  by  iodine. 

Hab.  on  maritime  rocks  washed  by  the  sea.  Rep.  Gov. 
Bot.  1854.     Doubtful  ;  probably  the  next  species. 

2.    L.  conjinis,  Ach. 

Similar  to  the  preceding,  but  smaller,  in  more  compact 
tufts,  and  with  terete  branches.  Spores  'OlOo  x  "Oil  mm. 
(Nyl.) 

Hab.  on  maritime  rocks  between  high  and  low  watei', 
Sandringham,  Barwon  Heads,  Lome,  Warrnambool. 


Tribe  2. — Collemei. 

Thallus  various  in  form,  membranaceous,  lobate  or  laciniate 
or  microphylline,  sometimes  fruticulose,  sometimes  granu- 
lose;  rigid  when  dry,  turgid  and  gelatinous  wlien  moist. 
Apothecia  lecanorine,  in  a  few  cases  biatorine,  in  still  fewer 
endocarpoid. 

Genus  1. — Synalissa,  D.R.  Nyl. 

Thallus  small,  of  various  forms,  incrusting,  submembran- 
aceous,  granulose  or  frnticulo.se.  Granula  gonima  in  globulai- 
cells.  Apothecia  innate,  lecanorine,  or  rarely  eudocarpoid. 
Spermatia  oblong. 

1.    S.  cancellata,  Wilson. 

Thallus  black  or  obscurely  olivaceous,  submembranaceous, 
cancellate,  minutely  atro-granuluse,  effuse  at  circumference 
and  encrusting  the  substratum.  Granula  gonima  light  green, 
contained  in  gelatinous  sacs  (inky  with  I),  1-5  in  each  ;  also 
moniliform  among  fine  elementary  filaments.  Apothecia 
minute  (to  '25  mm.),  prominent  in  the  thalline  granules, 
one  in  each,  at  first  endocarpoid,  at  length  I'ufescent, 
lecanorine,  elevated,  disk  concave  or  plane,  with  thalline 
margin  withdrawn.  Spores  colourless,  ellipsoid  or  ovoid, 
simple,  with  narrow  epispore,  'Ol  to  -012  X  •OO-i  to -OOG  mm. 
Paraphyses  slender  ;  thecal  cylindrical ;  gelatina  hymenia  I, 
vinous,  then  yellow. 


152      Proceedings  of  the  Ro//al  Societfj  of  Victoria. 

Hab.  on  sub-Alpine  rocks  and  moss,  Mt.  Maoedon. 
Previously  named  by  me  (Trans.  Lin.  Soe.  1890)  aS*.  micro- 
cocca,  Born,  et  Nyl. 


Genus  2.— Collema,  Ach.  Nyl. 

Thallus  very  various,  granula  gonima  moniliform,  no 
distinct  cortical  layer.  Apothecia  ruf'escent,  usually  leca- 
norine ;  hyjjothecium  distinctl}^  cellulose;  spores  eight, 
colourless,  commonly  multilocular,  rarel}'  simple. 

Sub-Genus  1. — Collema,  Spores  eUlptioid. 

1 .    G.  Iceve,  Ta3dor. 

Thallus  olivaceous,  under  surface  paler  or  cinerascent, 
smooth,  rotundo  lobate,  undulate.  Apothecia  rufous  or 
fusco-rufous,  plane,  at  length  convex,  with  a  thin  entire 
thalline  margin.  Spores  fusiformi  ellij)soid,  -013  X  '004  mm., 
o  to  5  septate,  and  also  longitudinally  divided.  Granula 
gonima  moniliform.     Gel.  hym.  blue  with  iodine. 

Hab.  among  mosses  on  gi-anitic  rocks,  M'Crae's  Island. 
Rep.  Gov.  Bot.  1854.     Traawool,  Beaconslield. 

Previously  named  liy  me  (Trans.  Lin.  Soc.  1890)  Leptogimn 
olivaceum,  Wilson. 

Form  gvanulatiLin,  Wilson. — Thallus  olivaceous  or  fusco- 
olivaceous,  here  and  there  plumbeous  :  beneath  paler,  firm, 
moderate  (one  inch  wide),  smooth,  very  often  near  centre  or 
wholly  obscurely  granulate. 

Hab.  on  bark  of  trees,  Wan-nainbool,  Gipp.sland,  Kew. 

¥ovn\  Jimbriatum,  Wilson.— Thallus  crisped  at  circumfer- 
ence, and  isidioso  fringed.     Sterile. 

Hab.  on  bark  of  trees,  Warrnambool. 

Form  IskUosum,  Wilson.— Thallus  plumbeo  coerulescent, 
here  and  there  olivaceous,  membranaceous,  thin,  plicato 
undulate,  more  or  less  covered  with  ccesious  or  obscurely 
plumbeous  isidia. 

Hab.  on  bark  of  trees,  Warrnambool,  Gippsland. 

2.    0.  plumheum,  Wilson. 

l^hallus  plumbeous,  small,  complicate,  membranaceous, 
rotundo-lobate,  lobes  undulate.  Apothecia  minute,  often 
ciowded,  rufous  bi'own,  sessile,  plane,  thalline  margin  entire. 


The  Lichens  of  Victoria.  153 

Spores  ellipsoideo-fusitbrni,  (Yl  x  OO-i  mm.,  5  septate,  and 
also  longitudimilly  divided.  Gran.  gon.  oblongo  globose, 
•()()3  to  OOo  mm.,  monilironii  :  yellow  with  iodine.  Gel.  liym. 
blue  with  iodine. 

Hab.  on  mosses  on  trees,  Warbuiton. 


3.    C.  jarviun,  Acb. 

'J'halliis  dark  fuscous  green  oi-  nigro-olivaceous,  membran- 
aceous, granulate,  lobate,  lobes  coniplicate,  often  undulate  ; 
blood-red  with  iodine.  Apotheeia  fuscous,  plane,  thalline 
margin  enti)-e.  S[)ores  ovoid  or  ellipsoid,  •018  to  •024'  x 
•OOIJ  to  •Oil  mm.,  3  septate,  irregularly  murali-locular. — B. 
V.  M.,  Vic.  Nat.,  Oct.  1887,  p.  88. 

4.    0.  ((train,  Wilson. 

Thailus  black,  moderate  (to  1^  inch),  circumference  lobate, 
eomplicato  squaniose,  tiiick,  cartilaginous,  granulato-corru- 
gate.  Apotheeia  black  or  dark  rufous  or  sometimes  pale, 
with  entire  thalline  margin,  attaining  1  mm.  diam.  Spores 
ovate  or  fusiforrni  ellipsoid,  acuminate  at  one  or  both  apices, 
•018  to  •026  X  •OOo  to  •OOG  mm.,  3  to  4  loculate.  Thecie 
clavate,  intensely  blue  vvith  iodine.  Paraphyses  slender, 
crowded. 

Hab.  on  calcareous  maritime  rocks,  Warrnambool. 


Sirji-GENiis   2.— Synkchoblastus,  s^jores  elongate. 
1.   S.  congestits,  Wilson. 

Thailus  black  or  atro-fuscous,  small  (to  1  inch)  cartilagin- 
ous, dittbi^mi-lobate,  lobes  i^otundate,  undulate,  circumference 
elevated,  incrassate,  arcuate.  Apotheeia  black  or  jmllid  or 
dark  red,  moderate  (2  mm.  diam.),  plane,  with  thickened 
margin,  at  length  convex,  imuiarginate.  Spores  cylindrical 
or  ellipsoideo  cylindrical,  sometimes  curved,  often  acuminate 
at  the  apices,  simple  or  1  septate,  containing  two  to  five 
locules,  •OIT  X  0035  mm.  Paraphj'ses  thick,  inarticulate. 
Gel.  hym.  blue  with  iodine,  thecse  intensely  blue.  Gran, 
gon.  conglomerated  into  sacs,  two  or  three  or  more  in  each, 
not  moniliform. 

Hab.  on  mosses,  &c.,  on  calcareous  maritime  rocks,  Warr- 
nambool. 


154     PivGecdin</.s  of  the  Royal  Socktij  of  Victoria. 

2.    S.  quadril  ocular  is,  Wilson. 

Thallu.s  fusco  olivaceous  or  iiigricaiit.  membranaceous, 
adhering,  lobate,  thicker  at  circumference,  crisped.  Apothecia 
moderate,  crowded,  black  or  rufous  black,  plane  or  somewhat 
concave,  thai  line  margin  entire.  Spoi'cs  cylindrical,  rounded 
at  each  apex,  somewhat  curved,  -02  to  03  X  003  to  -OOo 
mm.,  o  septate.  Paraphyses  slender,  inarticulate.  Gran, 
gon.  moniliform,  or  sometimes  conglou)erated  in  fours  into 
gelatinous  sacs. 

Hab.  on  mosses  on  sub-Alpine  granitic  rocks,  Mt.  Macedon. 

3.    S.  senecionis,  Wilson. 

Thallus  green  or  fuscous  green  or  olivaceous,  under  surface 
caerulean  green,  thin,  membranaceous,  smooth,  shining  or 
somewhat  shining,  or  sometimes  gi-anulato  rugulose,  moderate 
(2  to  3  inches),  rotundo  lobate,  lobes  imbiicate,  undulato 
crispate,  circtnnference  ascending.  Apothecia  I'ufous  or 
testaceo  rufous,  about  1  mm.  diam.,  often  crowded,  plane 
or  rather  convex,  thalline  margin  entire.  Spores  elongato 
fusiform,  straight  or  curved  or  .spirally  contorted,  "03  to  'Oo 
X  004  to  00(S  mm.,  3  to  9  septate.  Gel.  hym.  blue  witli 
iodine.  Gran.  gon.  oblong  or  reniform  (1  to  2  mm.  long),  or 
subglobose  (1  mm.  diam.) 

Hab.  on  bark  of  Senecio  bedfordii,  rarely  and  smaller  on 
bark  of  Prostanthei'a  lasiantlms,  Mt.  Macedon  ;  Lakes 
Entrance,  Gippsland.  When  young,  the  thallus  is  tense  and 
vivid  in  colour,  like  a  thin,  glistening  tilm  of  green  paint. 

4.    S.  leiicocarpas,  Taxdor. 

Thallus  1  to  3  inches  diam.,  loliaceo  membranaceous, 
smooth,  olivaceous,  lobes  rt)tundate,  undulate  and  plicate, 
margin  flexuo.se.  Apothecia  often  crowded,  albocarneous  ; 
disk  convex,  pruinose  ;  margin  entire,  at  length  concealed. 
Spores  elongato  fusiform,  oi'ten  acuminate  at  apices,  3  to  o 
septate,  -03  to  -Qh   x  '008  to  -01  mm. 

Hab.  on  trees,  Cromb.,  Journ.  Lin.  Soc,  XVII ;  Wilson  s 
Promontory,  Gov.  Bot.  Rep.,  1854;  Krplhbr.,  Vcrliandl. 
Zool.  Bot.'Gesell.s.,  in  Wien,  1880;  by  Curdie's  Creek,  Mt. 
Macedon,  Warburton,  Lome,  Glenmaggie,  Beaconsfield,  Mt. 
William,  Dandenong  Hills. 

Var.  1  pefrcois,  Wilson. — Obscurely  olivaceous  or  nigri- 
cant;  lobes  smaller,  somewhat  complicate.     Apothecia  small, 


Tlie  Lichens  of  Victoria.  155 

nigj-iennt,  seldom  carneous,  albo  pruiiiose,  disk  plane.  Spoi'es 
pluii-  (about  7)  septate,  -03  to  '04  x   •()()4'  to  -005  inm. 

Hab.  on  granitic  rocks  in  mountain  streams  in  Tallarook 
Ranges, 

Var.  2  minor,  Wilson. — Much  less  and  darker  than  the 
type ;  snbmonophyllous,  rotundate,  often  obscure!}"  furfur- 
aceo  granulose.  Apothecia  minute  and  much  crowded. 
Spores  as  in  type. 

Hab.  on  ti-ees  near  Lake  Wat  Wat,  Gippsland. 

5.   S.  glaucophthalmus,  Nyl. 

Thallus  olivaceo-fuscous,  -|  inch  or  more  diam.,  here  and 
there  fenestrato  dissected,  scrobiculate  and  often  granulifer- 
ous.  Apothecia  glaucous  lilac,  plane  or  somewhat  concave, 
thalline  receptacle  prominent,  margin  veiy  thin.  Spores  as 
in  .S.  nigrescens,  to  which  this  species  is  allied. 

Hab.  on  bark  of  trees  a.nd  bushes  ;  Leptogium  glauco- 
phthalmum,  B.  v.  M.,  Vic.  Nat,  Oct.  1887,  p.  89  ;  Warr- 
nambool,  Mordialloc,  Cunninghame,  Buninyong,  Lake  Wat 
Wat. 

0.  (S.  nigrescens,  Huds. 

Thalkis  black  green,  thinly  meml)riinaceous,  submono- 
phyllous,  (jrbiculai-,  depressed,  rotundato  lobate,  radiately 
rugoso  plicate.  Apothecia  obscurely  rufous,  plane,  crowded, 
thalline  margin  entire.  Spores  fusiformi  cylindrical,  often 
pluriseptate,   03 i  to  -042   x    00.5  mm. 

Hab.  on  trunks  of  trees,  Warrnanib')ol,  Mordialloc,  Cun- 
ninghame, Buninyong,  Metung. 

Genus  3. — Leptogiu.m,   Fries. 

1.  L.   hiloculare,  Wilson. 

Thallus  plumbeou.s,  membranaceous,  small  (|  inch)  lacin- 
iato  lobate,  lobes  sinuate  and  undulate.  Apothecia  pale 
rufous,  minute  (2- 7  mm.  diam.);  thalline  margin  prominent. 
Spores  fusiformi  ellipsoid,  bilocular,  015  x  "006  mm.  Gel. 
hym.  blue  with  iodine. 

Hab.  on  the  bark  of  a  tree,  Mt.  Macedon. 

2.    L.  sinuatum,  Huds. 

Thallus  plumbeo-fuscescent,  rotundato  lobate,  reticulato 
rugulose,  lobes  crowded,  imbricated,  margin  entire  or  crenate. 


\')(j      Proceedings  of  the   Royal  Society  of  Victoria. 

sub-erect.    Apotbecia  brown,  scattered,  small,  sessile,  concave, 
margin  smooth,  entire,  elevated.     8pores  oblongo  ellipsoid, 
attenuated  at  apices,  irregularly  murali  locular,  02  x  'OOS  mm. 
Hab.  on  mossy  rocks,  Mt.  Macedon,  Kilmore,  Lome. 


3.    L.  lacerum,  Acli.  var.  intcrriiediurn,  Ai-n. 

Thallus  plumbeous,  or  pallido  plumbeous,  or  fuscescenti 
plumbeous,  very  thin,  smooth  or  slightly  rugulose,  undulate 
and  plicate,  laciniato  lobate,  lobes  rotundate  or  lacerate  ; 
margin  crisped,  irregularly  crenate  or  spatulato  fimbriate, 
fimbria  often  repeatedly  branched.  Apothecia  pallido  rufous, 
not  frequent,  small  or  moderate,  margin  elevated.  Spores 
oblongo  ovoid,  narrow  at  one  or  both  apices,  murali  locular 
in  typically  3  series,   025  x    008  nun. 

Hab.  on  mossy  rocks  and  bushes  in  bed  of  stream,  Cobden, 
Mt.  Macedon,  Beaconsfield,  Lome. 

Aspect  intermediate,  between  lacerum  and  tronelloides. 

Var.  2.  inilvinaturn,  Hftm. — Thallus  dark  brown,  smaller, 
])ulvinate,  lobes  minute,  much  crowded,  denticulato  laciniate; 
sterile. 

Hab.  on  earth,  Kew  ;  rare. 


4.     L.    tremelloide-s,     L.     var.     azureum,    Sw.  =   Collema 
azureum.      lleport  Gov.  Bot.   1854. 

Thallus  plumbeo  glaucescent,  here  and  there  fuscescent, 
smooth,  lobate,  imbricate  and  crispate.  Apothecia  rufous, 
elevated,  margin  entire,  plumbeous  or  pallid.  Spores 
ellipsoid,  acuminate  at  apices,  5  septate  and  also  longitudin- 
ally divided,  -016  X    00(3  mm. 

Hab.  on  trees  {Golleriuh  azureum),  McCrae's  Island, 
Rep.  Gov.  Bot.  1854 ;  Cobden,  Lake  Elingamite,  Black 
Spur,  Mt.  Macedon,  \Varl)urton,  Lome,  Beaconsfield,  Lakes 
Entrance. 

Var.  2.  muscite(jenf<,  Wilson. — ^Darker  and  firmer  than  the 
type,  less  undulate,  ascending. 

Hab.  on  stems  of  mosses  on  trees,  Warburton,  Korum- 
bui'ra. 

Var.  3.  isidiosum,  Wilson.— Much  smaller  than  the  type, 
partially  covered  with  a  gianulose  isidium.  Apothecia 
small,  occasionally  isidiose  on  margin. 

flab,  on  mossy  bush,  Cunninghame. 


The  Lichens  of  Victoria.  157 

5.   L.  philorJienma,   Wilson. 

Thallus  more  or  less  obscui-ely  plumbeo  cinereous  oi- 
brown,  very  thin,  to  |  inch  wide,  sub-ascending,  lobate, 
plicato  undulate ;  margin  crenate,  sinuate  and  crisped. 
Apothecia  small  (1  to  I'o  mm.),  disk  more  or  less  obscurely 
rufous,  sometimes  black,  concave  or  plane  ;  thalline  margin 
entire,  thick,  rounded,  elevated,  at  length  thin,  equal. 
Spores  ovate,  3  septate,  and  also  longitudinally  divided, 
•01  o  X  -007  mm. 

Hab.  on  mosses  and  rocks  in  the  channels  of  stream^;, 
Curdie's  Creek,  Lome,  Tallarook. 

Named  by  Dr.  Knight  as  L.  dactylinum,  and  so  reported 
by  me  (Trans.  Lin.  Soc.  1890). 


6.    L.  victorianwin,  Wilson. 

Thallus  obscurely  plumbeous,  here  and  there  rufo-fuscous, 
under  surface  nearly  concolorous  or  sub-cinerascent,  large 
(3  inches  or  more  diam.),  more  or  less  confusedly  rugulose, 
rotundo  lobate,  lobes  undulate,  firm,  but  in  old  lobes  thick 
and  occasionally  fusco-furfuraceous,  as  if  deprived  of  cortex, 
sometimes  clothed  with  squamules.  Apothecia  moderate, 
sometimes  rather  large  I'ufous  oi- .  fusco-rufous,  thalline 
margin  sometimes  excluded,  more  generally  plicate  or 
granulate  or  briefly  laciniate.  Spores  ovoideo  fusiform, 
often  acuminate  at  the  apices,  typically  three  septate,  often 
with  the  central  locules  longitudinally  or  obliquely  divided, 
•013  to  "017  X  "OOo  to  •00()  mm.     Granula  goniraa  moniliform. 

Hab.  among  mosses,  on  trees  and  rocks  abundantly,  Mt. 
Macedon,  Black  Spur,  Cobden,  Sandiingham  (one  specimen), 
Warburton,  Korumburra. 

Allied  to  L.cklovomelum,  Sw.,  and  perhaps  a  variety  of  it. 


7.    L.  pliyUocarpum,  Pers.  var.  dwdaleum,  Flot. 

Thallus  fusco  i)lumbeous,  here  and  there  pallido  plumbeous, 
fii'm.  lobate,  lai'ge  (three  or  four  'n.  diameter),  longitudinally 
and  very  closely  undulato  rugose,  or  finely  and  acutely 
corrugate,  under  surface  paler.  Apothecia  dark  rufous, 
often  large,  thalline  margin  thick,  densely  corrugato  rugulose, 
spores  ellipsoid,  attenuate  at  each  apex,  five  septate,  and 
also  longitudinally  divided,  -03  to  034  x  012  to  -015  mm. 
(Nyl.) 


158      Proceedhu/fi  of  ihe  Royal  Society  of   Victoria. 

Hab.  on  trees  and  bushes,  Warrnanibool,  Lake  Victoria, 
Ounningliame,  Lake  Wat  Wat ;  abundant,  l)ut  rather  rare 
in  fruit. 

<S.    L.  pecten,  Wilson. 

TliaUus  minute,  very  thin,  squamulifonn,  plumbeous  or 
brown,  margin  digitato  crenate,  often  pulvinato  crowded. 
Apotheeia  large  for  the  size  of  the  plant  (I "5  mm.  diameter), 
rufescent,  concave,  with  a  thin,  pallid  margin,  often  immar- 
ginate.  Spores  ellipsoid,  three  septate  with  central  locules 
often  longitudinall}^  divided,  -016  to  -024?  x  'OOS  ram. 

Hab.  on  dead  or  old  bark  of  trees,  not  common,  Mordialloc, 


\).    L.  Burge.ssii,  Lightfoot. 

Thallus  plumbeous  or  brown,  laciniato  lobate,  complicate, 
lobes  variously  margined,  undulate  and  curled,  under  surface 
cinerascent  and  albido  tomentellose.  Apotheeia  dark  rufous, 
somewhat  large,  plane  or  concave,  margin  thin,  entire,  or 
sub-foliaceo-crenulate.  Spores  ellipsoid,  attenuate  at  each 
apex,  three  septate,  and  also  longitudinally  divided,  'OS  to 
■04  X   -012  to  015  mm. 

Hab.  on  bushes  and  trees  and  mossy  rocks,  Curdie's 
Creek,  Warrnambool,  Buninyong,  Lake  Wat  Wat,  Mount 
William  ;  not  common. 

10.    L.  injiexum,  Nyl. 

Thallus  plumbeous  or  plumbeo  cajrulescent,  membranace- 
ous, dilated,  two  to  three  inches  diameter,  smooth,  laciniato 
incised,  margin  intlexo  convolute,  broadly  sinuate  and 
crenulate ;  under  surface  pallescent,  very  thinh-  tomentel- 
lose, but  wide  at  margin.  Apotheeia  rufous,  plane  or  some- 
what concave,  rather  large,  appressed,  foliaceo  crenulate. 
Spores  ellipsoid,  attenuate  at  each  apex,  plurilocular,  -03  to 
•036  X  -013  to  017  mm.  (Nyl.) 

Hab.  on  rock  at  Waterfall,  Upper  Maffra. 

Var.  limhutum,  Wilson. — Thallus  orbicular  and  rosulate, 
margin  for  the  most  ]iart  densely  and  minutely  fimbriate. 

Hab.  on  trees  and  mossy  logs  in  sub-Alpine  localities, 
Black  Spur,  Warburton,  Mt.  Macedon. 

11.   L.  dcnticulatum,  Vic.  Nat.,  Oct.  1887,  B.  v.  M. 


The  Lichens  of  Vietoria.  \oO 

12.    L.  hi/p(>tmchi/)niiii,  Mull.  Arg. 

Thallus  about  4  ceutim.  wide,  lacinia^  liovizorital  or 
nscendiiig,  obovate,  obtusely  lobate,  mavgin  entire,  thinly 
coriaceous,  fusco  olivaceous,  both  surfaces  concolorous, 
smooth  above  or  shghtly  rugulose,  crowded  beneath  with 
polymorphous  prominences,  obovoid,  obtuse,  entire  or  ob- 
tusely lobate,  exasperate  or  verruculose  tomentellose.  Apo- 
thecia  2  mm.  diam.,  spores  fusiformi  ellipsoid,  5  septate, 
niultilocular,  -02.")  x  "Ol  mm.  Mull.  Lich.  Beit.  XII,  12, 
Katisb.  Flora. 

13.  L.  mi sf vale,  Hook  and  Tayl. 

Thallus  foliaceo  membranaceous,  thin,  blackish  olive, 
smooth,  lobes  ascending,  sub-imbricate,  somewhat  concave, 
rotundate,  undulate,  entire,  under  surface  paler,  sub-tomen- 
tose.  Apothecia  elevated,  black,  at  length  convex,  margin 
thin,  entire.     McC'iae's  Island,  Rep.  Gov.  Bot.  1854. 

14.  L.  rufjatum,  Hook  and  Tayl. 

Thallus  gelatinous  membranaceous,  3  inches  diam.,  fuscous 
green,  covered  with  close  longitudinal  plaits  ;  lobes  crowded, 
ascending,  oblongo  rotundate,  erenate,  somewhat  concave, 
with  minute  granulate  stipitate  isidia  expanding  into 
rlialline  lobes,  sterile. 

Hab.  on  trees,  McCrae's  Island,  Rep.  Gov.  Bot.  1.S54. 


FAMILY   II.— MYRIANGIACEI. 
GeiNUS  1 .— Myriangium,  Mnt.  and  Berk. 

Thallus  black,  noduloso  pulvinate,  cellulose,  unstratified. 
Apothecia  sublecanorine,  sphseroideo  celhdose.  Spores  8, 
colourless,  irregularly  septate. 

1.    M.  duriwi,  M.  and  B.  =  M.  dwrieui,  of  De  Bary. 

Thallus  black,  opaque,  small,  tuberculato  glon:ierate  or 
nodoso  confluent,  often  depresso  pulvinate.  Apothecia 
minute,  black,  slightl}"  impressed.  Spores  oblong  or  oblongo 
ovoid,  variousl}-  septate,  "(UT  to  "024  x  "007  to  '008  nmi. 

Hab.  on  bark  of  trees,  Mount  Macedon,  Sandringham, 
Koruraburra,  Kilmore. 


!(!()      Proceedhif/s  of  the  Royal  Society  of  Victoria. 

2.    M.  dolicJiosponirn,   Wilson. 

TluiUus  black,  opaque  or  slightly  shining,  small  (2  to 
5  mm.  wide  and  2  mm.  high),  unequal.  Apothecia  numerous, 
nearly  covering  the  thallns  and  concolorous  with  it, 
stipitate;  epithecia  subrufescent,  plane  or  concave,  to  1  mm. 
broad,  with  rotundo  obtuse  thai  line  margin  ;  stipe  some- 
times 1  mm.  long,  tapering  downwards.  TheciB  sphseroidal, 
dispersed  in  the  cellular  sidjstance  of  the  epithecium. 
Spores  cylindrical,  sim[)le  oi-  obsoletely  septate,  arcuate, 
someiivhat  acuminate  at  a])ices,  with  minute  guttse  arianged 
in  the  longitudinal  axis,  -04  x  OOG  mm.  Gran,  gonim. 
•002  to  -007  mm.  diam.,  often  conglomerate.  Texture  of 
thallus  fuscous,  cellular,  cells  angular,  OOS  to  '005  mm.  diauj. 
Cells  in  epithecium  spherical. 

Hab.  on  twigs  of  Hymenanthera  banksii,  Maffra. 

The  whole  ])lant  is  often  covered  with  the  scyphophoroid 
apothecia  standing  out  in  all  directions,  and  of  various  sizes 
and  stages  of  development.  The  epithecium  is  almost 
identical  in  texture  with  the  epithallus,  but  is  geneially 
concave  and  slightly  rufescent.  In  old  apothecia  it  is  worn 
into  cavities,  whicli  give  it  a  granulato  rugulose  appearance. 
Both  thallus  and  apothecia  contaiii  granula  gonima,  usually 
conglomerate.  When  a  dried  specimen  is  submerged  iri 
water,  there  arise  from  it  on  all  sides  stieams  of  minute  air 
bubbles  for  a  considerable  time,  showing  the  porous  nature 
of  the  plant.  It  does  not,  however,  appreciably  increase  in 
size  when  moistened  as  the  Collemacei  do. 


FAMILY   III.— LICHENACEI. 

Thallus  various  in  coloui-,  white,  whitish,  cinerascent, 
flavicant,  rufous,  fuscous,  veiy  rarely  nigricant,  and  various 
in  form,  filamentous,  foliaeeous,  sijuamose,  ci'ustaceous. 
pulverulent  or  evanescent.  The  gonidial  stratum  very 
geneially  of  ti'ue  gonidia.  Ajiothecia  vaiious  in  form, 
.stipitate,  lecanorine,  peltate,  patellulate,  lirellate  or  pyreno- 
carpous. 

Series  I. — Epiconiodei. 

Apothecia  with  the  spi)res  naked,  collected  into  a  s[)oral 
mass  on  the  surtace. 


The  Lichen.s  of  Victoria.  IGl 

Tribe  1. — Caliciei. 

Thallus  crustaceous,  granulose  or  obsolete,  yellow  or  flavo- 
virescent,  or  cinerascent,  or  whitish,  or  none.  Apothecia 
cnpuliform,  sessile  or  stipitafce. 

Genus  1. — Sphinctkina,  Fr.  pr.  p.  D.  N. 

Thallus  none.  Apothecia  parasitic  on  Pertusarice,  globoso 
turbinate,  shining,  black,  sessile  or  shortly  stipitate.  Spores 
8,  nigrescent,  simple. 

1.    S.  microceplMla,  Nyl. 

Apothecia  black,  globoso  turbinate,  briefly  stipitate,  nearly 
sessile,  capitula  small  (about  -1  mm.  broad),  spores  nigricant, 
fusifoiini  ellipsoid,  nearly  globose,  but  acuminate  at  apices, 
epispore  thick,  reddish,  -01  to  -012  x  'OO-i  to  008  mm. 

Hab.  on  some  pertussaria,  on  bark  of  Hymenanthera 
banksii,  Maffra. 

Form  tenella,  Wilson. —  Like  the  type,  but  with  a  smaller 
capitulum  and  longer  stipe  (to  o  mm.) 

Hab.  along  with  type,  Maff'ra. 

Genus   2. — Calicium,  Ach,  Nyl. 

Thallus  granulose,  powdery,  crustaceous,  squamulose,  or 
altogether  evanescent.  Apothecia  generally  black,  stipitate 
or  subsessile,  capitula  globose,  or  turbinate,  or  cupular. 
Spores  fuscous  or  nigricant.     Spermatia  short,  oblong. 

1.    C.  clirysocepJtalum,  Ach. 

Thallus  citrine  or  obsolete.  Apothecia  small  (G  mm. 
high),  black,  stipe  slender  (-OG  mm.  thick).  Capitulum 
small  (12  mm.  broad),  turbinate  ;  beneath  citrino  suffused. 
Sporal  mass  umber  brown  ;  spores  fuscous,  globose,  -OOS  to 
006  mm.  diam. 

Hab.  on  decorticated  decaying  eucalyptus,  near  river,  at 
Maffra,  Kilmore. 

YciY.  jilare,  Ach. — Stipe  longer  and  more  slender  (to  -8  x 
•4r  ram.) ;  capitulum  smaller  ;  sporal  mass  protruding  upwards 
to  a  great  height. 

Hab.  along  with  type,  Maffra. 

M 


162     Proceedings  of  the  liot/al  Society  of  Victoria. 

2.    G.  2)hoeocephalum,  Borr.  var.  phfjedrosporurii,  Wilson. 

Tballus  white,  or  whitish,  witli  pale  glauceseent  venucose 
congested  granules,  which  are  sometimes  dissolved  into 
citrine  soredia.  Apothecia  atro-fuscous,  with  slender  stipe 
(about  "2  mm.  high,  -1  mm.  thick),  the  upper  ]>art  citrino 
suffused  ;  capitulum  hemispherico-turbinate  or  sub-globose  ; 
margin  citrino  suffused  ;  sporal  mass  from  fulvous  to 
umbrine.  Spores  dilutely  nigrescent,  very  nearly  colourless, 
<lelimitated  by  a  dark  line  ;  form  variable,  globose  or 
ellipsoid,  simple,  nucleated,  diameter  -002  to  "004  mm. 

Hab.  on  decaying  decorticated  eucalyptus,  near  Kilmore. 

I  am  doubtful  whether  the  granules  of  the  tballus  belong 
to  this  liclien,  or  are  au  undeveloped  form  of  some  other. 
Perhaps  the  plant  is  of  a  new  species,  which  may  be  called 
C.  phmdrosporurti. 


3.    C.  nivewni;  Wilson. 

Thallus  snowy  white,  thick,  or  cinerascenti  albid,  thinner, 
effuse,  rimulose  with  convex  areolse.  Apothecia  minute 
(•5  to  '8  mm.  high),  stipe  slender  (-07  mm.  thick),  either  all 
whitish  or  partly  hyaline  and  partly  fuscous,  or  all  fuscous, 
or  ail  black,  sometimes  furcate.  Capitulum  hemispherico 
lenticular,  black,  about  "25  mm.  broad,  sometimes  divided 
into  two  or  three  or  more  lobes.  Spores  dilutely  nigrescent, 
fusiformi  ellipsoid,  or  oblong,  compressed,  simple,  ■004'  to 
•006  X  -002  to  0025  mm.,  paries  thick.  Gel.  hym.  with 
iodine  vinous  yellow. 

Hab.  on  dead  bark  of  living  eucalyptus,  Cunninghame, 
Maffra,  Beechworth. 

Perhaps  a  variety  of  0.  pusiolum,  Ach. 


4.    C.  Victories,  C.  Knight. 

Thallus  white  or  whitish,  or  cinerascent,  more  or  less 
marked,  effuse.  Apothecia  all  Ijlack,  -5  to  1  nun.  high, 
stipe  slender  (•]  mm.  thick)  and  a  little  thicker  at  the  base, 
(vapitulum  turbinate  lenticular  or  hemispherico  lenticular, 
■25  to  "5  mm.  broad.  Spores  fuscous  or  fuscescent,  fusiformi 
ellipsoid,  compressed,  simple,  -005  to  '008  x  "002  to  -003  mm., 
when  viewed  from  the  side  bacillar,  -0015  mm.  wide  ;  paries 
thick,  defined  by  a  dark  line  on  the  outside. 


The  Lirheu.'^  of  Victoria.  103 

Hab.  on  decaying  decorticated  eucalyptus,  Croydon,  Kew, 
Warrnarabool,  Wan-agnl,  Black  Spur,  Lakes  Entrance,  -Mt. 
William,  Tallarook,  Mt.  Macedon,  Beech  worth  ;  frequent. 

Allied  to  C.  jxirietinuiii.  Somewhat  variable.  Cjejitiiwiii 
reported  by  me  (Trans.  Lin.  Soc.  1890),  is  now  judged  by 
me  to  be  a  not  clearly  marked  form  of  C.  Victories. 

o.    C.  /MtrruJam,  Wilson. 

Thallus  white  or  whitish,  sub-determinate.  Apothecia  all 
black,  4  mm.  high,  stipe  slender  ('OS  mm.  thick).  Capituliwn 
lenticular,  -KJ  nnu.  broad.  Spores  dilutely  nigrescent, 
ellipsoid,  simple,  OOo  to -OOO  x  OOlo  to  •0()2o  mm.,  paries 
thin,  black. 

Hab.  on  decorticate  eucalyptus,  Maffra,  Mt.  ]\Iaced(3n, 
Beech  worth. 

The  thallus  looks  like  a  thin  coat  of  whitewash,  on  which 
the  densely  Ijlack  apothecia,  though  ver}^  minute,  ai'e  clearly 
visible.  The  outline  of  the  spores  is  remarkable  for  its 
blackness,  being  in  this  respect  like  G.  Vldovke. 

(J.    C.  coiitortum,  Wilson. 

Thallus  whitish,  very  thin.  Apothecia  all  black,  1  mm. 
high,  stipe  •!  mm.  thick,  contorted.  Capitulum  hemispherico 
lenticular.  Spores  dilutely  nigrescent,  fusiformi  ellipsoid, 
simple,  -004  x  OOl-t  to  002  mm. 

Hab.  on  decorticate  deca\'ing  eucalyptus.  Lakes  Entrance. 

Allied  to  G.  Victoria',  which  it  resembles  in  its  apothecia, 
but  the  capitula  are  smaller,  the  spores  also  are  smaller 
and  narrower,  and  dilferent  in  colour. 

7.    G.  (jracillimuTii,  Wilson. 

Thallus  indicated  by  a  whitish  s})ot.  Apothecia  all  black, 
small  ("8  mm.  high)  ;  stipe  very  slender  (06  mm.  thick)  ; 
capitulum  minute  (1  mm.  broad)  ;  tui-binato  lenticulaj-. 
Spores  more  or  less  dilutely  nigrescent,  ellipsoid  or  fusiforn)i 
ellipsoid,  uniseptate,  'OOS  to  OOl  x  -001  to' -002  mm.,  with  n 
locule  in  each  cell. 

Hab,  on  decaying  decorticated  Ader  a/r(ji/rophyllas,  Mt. 
Macedon, 

The  apothecia  are  extremely  minute,  being  visible  only 
under  a  powerful  lens. 

M  2 


1G4     Proceedings  of  the  Royal  Sociefi/  of  Victoria. 

8.    C.  deforrae,  Wilson. 

Tballus  cinerascent,  thin,  gmnulose.  Apothecia  black, 
detbi-med  by  thalliue  and  other  granules,  1  mm.  high,  or  a 
little  more;  stipe  "2  mm.  thick;  capitulum  turbinate 
lenticular,  to  '5  nnn.  broad.  Sporal  mass  black,  protruding, 
sometimes  extending  fai*  on  one  side.  Spores  nigrescent, 
fusifoi-mi  ellipsoid,  uniseptate,  septum  often  indistinct,  '006 
to  -008  X   -0025  to  -004  mm. 

Hal),  on  decaying  decorticated  eucalyptus.  Lakes  Entrance. 
The  apothecia  have  a  deformed  appearance,  unlike  the 
ordinary  neatness  of  the  genus.  Its  surface  seems  to  be 
glutinous,  readily  retaining  any  gianules  or  other  particles 
that  fall  on  it. 

[).    C.  roseo-alhidum,  Wilson. 

I'halius   rosy-whitish,  thick,  minutely  cancellate,  chr3^so- 

gonimic.     Apothecia  minute  ("7  mm.  iiigh),  all  black  ;  stipe 

slender  ('06  mm.  thick)  ;  ca[)ituluni  lenticular  ('2  mm.  broad). 

S[)ores  nigrescent,  oblong  or  oblongo  ellipsoid,  apices  rotund- 

te,  uniseptate,  '(K).")  or  more  x  "002  to  -OOS  mm. 

Hab.  on  decayed  decorticated  eucalj'ptus,  Maifra,  Kilmore. 
The  thallus  covers  a  good  part  of  the  tree,  and  when  bruised, 
it  becomes  a  deep  yellow. 

10.    C.  ca/pillave,  Wilson. 

Tliallus  white,  thin,  or  very  thin.     Apothecia  all  black, 

1  mm.  high ;  stipe  slender  (•]  mm.  thick)  ;  capitulum 
turbinato  lenticular,  "3  to  "4  mm.  broad.  Spores  nigrescent, 
oblong  or  ellipsoid,  uniseptate,  "COS   x  •0025  mm. 

Hab.  on  decaying  decorticated  eucalyptus,  Mt.  Macedon, 
Warburton,  Maffra. 

Perhaps  a  variety  of  0.  xnhtile.  Pers.,  of  which  I  reported 
it  a  variety  (Trans.  Lin.  Soc,  1890). 

IJ.    C.hiloculaTe,W\\HO\\. 

Tliallus  whitish  or  cinerascent,  thin.  A})othecia,  all  black, 
•8  to  ■!  mm.  high  ;  stipe,  '1  mm.  thick  ;  capitulum  lenticu- 
lar or  sub-turbinato  lenticular,  -.3  to  4  mm.  broad.  Spores 
iuscescent  or  fusco  nigrescent,  ellipsoid,  or  sub-fusiformi 
ellipsoid,  bilocular  or  obsoletely  bil  ocular  or  simple,  with 
septum    not    visible,    epispore    thickish,    "005    to   '008     x 

002  to  -0035  mm. 


Tlie  Lichens  of  Victor  la.  1G5 

Hab.  on  decaying  decorticated  eucalyptus,  Warrnanibool, 
Maffi-a,  Bright. 

Perha}).s  a  variety  of  C.  suhtile,  Pers.,  of  which  I  reported 
it  a  variety  (Trans.  Lin.  8oc.,  1890). 

12.  G.  obovatum,  Wilson, 

Thallus  cinerascent,  thin.  Apothecia  black,  to  8  or  I  mm. 
high;  stipe  to  'OS  or  "1  mm.  thick;  capituliim  obovate  or 
turbinate,  '2  to  '3  mm.  broad.  Spores  nigrescent,  ellipsoid, 
nniseptate,  each  cell,  containing  a  globular  locule,  septum 
not  always  visible,  005  to  -012   x  -003  to  OOi  mm. 

Hab.  on  eucalyptus  wood  in  mountain  regions,  Mt. 
JVJacedon. 

Distinct  by  the  obovate  capitulum. 

13.  G.  piperatum,  Wilson. 

Thallus  albido  cinerascent  or  cinereous,  thin.  Apothecia 
black,  sub-sessile,  "2  mm.  higli ;  stipe  thick  (1  mm.)  ; 
capitulum  lenticular,  disk  plane,  -25  mm.  broad.  Spores 
fusco  nigrescent,  oblong,  nniseptate,  each  cell  containing  a 
globular  locule,  -001^  to  008   x  "0025  to  -005  mm. 

Hab.  on  eucalyj^tus  wood,  both  trees  and  fences,  common, 
Mt.  Macedon,  Kilmore,  Beech  worth. 

14.  G.  nigrum,  Sch?er  var.  miiiiitum,  Knight. 

Thallus  obscurely  cinerascent,  granulose.  Apothecia  all 
black,  small  ("o  mm.  high);  stipe  thick  (-1  to  -12  mm.); 
capitulum  turbinate)  c_ylindrico],  disk  pruinose,  "3  to  7  mm. 
broad.  Spores  nigrescent,  ellipsoid,  nniseptate,  constricted 
in  the  middle,  each  cell  containing  a  globular  locule,  '004 
to  -012   X   -002  to  -OOG  mm. 

Hab.  on  the  horizontal  surface  of  decaj'ing  eucal3"ptus 
fences,  Kew,  Maffra,  Oakleigh. 

15.  G.  quercinurii,  Pers.  var.  bidbosum,  Wilson. 

Thallus  albido  cinerascent.  Apothecia  to  -1  mm.  high  ; 
stipe  to  "2  mm.  thick,  capitulum  glubose,  'o  mm.  diam., 
cinereo  pruinose  beneath.  Spores  fusco  nigrescent,  subfusi- 
formi  ellipsoid,  OOG  to  '012  x  003  to  005  mm.,  nniseptate, 
septum  often  indistinct,  cells  containing  each  a  nigrescent 
locule,  epis])oro  distinct,  rubescent. 


l(i(i      Proceedings  of  ihe  Royal  Society  of  Victoria. 

Hab.  on  decajing  decorticatpcl  eucalyptus,  Mt.  Macedon. 

Re[)orted  by  ine  (Trans.  Lm.  Soc.)  as  C.  hulbosum,  and 
perhaps  a  variety  of  C  quercinimi. 

Var.  2.  mic7-ocai-puin,  Wilson.— Thallus  cinereous.  Apo- 
tbecia  sraall,  '3  to  -4  mm.  high  ;  stipe  black,  -1  to  "2  mm. 
high,  -Oo  to  "1  mm.  thick  ;  capitulum  turbinate,  disk  flat,  "1 
to  -2  mm.  broad,  margin  cinerascent  or  albido  cinerascent. 
Spores  fuscescent,  1  septate,  paries  thick,  constricted  in 
middle  ;  apices  ratlier  acuminate,  "OOS  x    OOS  nnn. 

Hab.  on  decaying  eucalyptus  stump,  near  Tallarook. 

Var.  3.  Clarensis,  Wilson. — Thallus  whitish  or  cinerascent, 
of  medium  thickness.  Apothecia  black,  '8  mm.  high,  stipe 
•1  mm.  thick,  ca])itulum  '3  mm.  broad,  turbinato  lenticular, 
margin  whitish.  Spores  fuscous  or  fuscescent,  ellipsoid, 
narrow  at  apices,  often  constricted  in  middle,  uniseptate  or 
bilocular,  -00.5  to  -OOS  x  •002  to  -003.5  mm. 

Hab.  on  decaying  decorticated  eucalj'ptus,  Bright,  Beech- 
worth. 

16.    C.  cwrtmn,  Borr. 

Thallus  whitish,  thin  or  evanescent.  Apothecia  to  1  '8  mm. 
high,  but  often  much  less,  stipe  to  "2  mm.  thick,  capitulum 
turbinate,  to  'G  mm.  broad,  albo  suffused  beneath.  Sporal 
mass  black,  protruded  upwards.  Spores  nigricaiit,  ellipsoid, 
uniseptate,  OOo  to  -01  x    002  to  003  mm.  " 

Hab.  on  decaying  dec(n-ticated  eucalyptus  and  old  hard- 
wood fences,  frequent  and  abundant,  Lome,  Mt.  Macedon, 
Oakleigh,  Black  Spur,  Mafira,  Bright,  Mordialloc. 

17.    C.  tracheUintm,  Ach.  var.  elattospo7  aiH,  Wilson. 

Thallus  obscurely  cinerascent  or  albescent.  Apothecia 
very  various  in  size,  to  2  mm.  liigh  ;  stipe  at  the  base  '25 
mm.  thick;  capitulum  globose  or  turbinate,  to  -5  mm.  broad, 
rufous  at  margin  and  upper  part  of  stipe  and  even  the  disk. 
Spores  003  to  008  X  -002  to  004  mm. 

Hab.  on  decaying  decorticated  eucalj-j^tus  and  fences, 
Cobden,  Warburton,  VVarragul,  MafFra,  Lome,  Cunninghame. 

'J'he  dnnensions  of  the  spores  are  half  of  those  described 
by  Nylander.  This  is  in  Victoria  the  commonest  species^of 
this  genus,  and  often  grows  in  large  patches  on  the  trees, 
covering  man}^  square  feet  with  abundant  apothecia,  some- 
times making  the  tree  seem  as  thouL;h  clothed  with  short 
hair. 


Tlie  Liclu'UH  of  Victoria.  167 

Var.  2.  meiocarpiirit;  Wilson. — Thallus  whitish,  thin. 
Apothecia  small,  about  '8  niui.  high  ;  stipe  about  "1  mm. 
thick  ;  capitulum  turbinato  lenticular  "3  ram.  broad;  margin 
and  upper  ]iart  of  stipe  rutbus.  Spores  fuscescent,  ellipsoid, 
constricted  in  middle,  uniseptate,  with  minute  loculi  in  each 
'•ell,  -006  to  -007  X   003  mm. 

Hab.  on  decorticated  lio-htwood  tree,  Kilmore. 


18.    6'.  aiu'igerum,  Wilson. 

Thallus  white  or  whitish,  somewhat  thick.  Apothecia 
small,  stipe  black  2  to  '8  mm.  high,  -05  to  'l  ram.  thick  ; 
capitulura  wholly  covered  with  ilavescent  powder,  lenticular, 
■4  nmi.  broad.  Spores  nigrescent  or  fuscous,  ellipsoid, 
uniseptate,  rather  constricted  in  middle,  containing  a  locule 
in  each  cell,  -005  to  -007  x  -002  to  -004^  rara. 

Hab.  on  decajdng  eucalyptus  wood,  Mt.  Macedon. 

Possibly  only  a  variety  of  C.  roscidum. 


19.    G.  roscidum,  Flk.  var.  eucalypti,  Wilson. 

Thallus  cinerascent,  here  and  there  flavo  sorediose  and 
then  sterile.  Apothecia  to  l"3  rara.  high,  stipe  black,  '1  mm. 
thick ;  capitulum  turbinate,  beneath  more  or  less  flavo 
virescent,  to  '3  mm.  broad.  Spores  fuscescent  or  more  or 
less  dilutely  nigrescent,  defined  by  a  black  line,  ellipsoid, 
narrow  at  each  apex,  often  constricted  at  middle,  uniseptate, 
containing  a  paler  locule  in  each  cell,  •005  to  009  x  '003  to 
•005  nira. 

Hab.  on  dead  bark  and  decaying  wood  of  eucalypti. 
Beech  worth,  Mt.  Macedon. 


20.    C.  roscidulum,  Nyl. 

Thallus  white,  thick,  iiere  and  there  rufescent  (query 
alien  ?).  Apothecia  •O  mm.  high,  stipe  "l  mm.  thick ; 
capitulum  turbinato  lenticular,  '4  mm.  broad  ;  margin  and 
upper  part  of  stipe  golden  green.  Spores  fuscescent, 
ellipsoid,  constricted  at  middle,  uniseptate,  -003  to  '006  X 
•002  to  -003  ram. 

Hab.  on  decayed  eucalyptus  stump,  Kilmore. 

Probably  a  mo'e  variety  of  C.  roscidum. 


1G8     Proceedings  of  the  lioijal  Society  of  Victoria. 

21.    C.  hijperellum,  Ach. 

Thallus  flavo  viresceut,  grannlose  or  sub-leprose.  Apo- 
thecia  black  ;  capitulum  globoso-lentiform ;  stipe  l)]ack, 
elongate ;  sporal  mass  black  or  umber  black.  Spores 
nigrescent,  ellipsoid,  uniseptate,  'OOO  to  'OIC  x  "004  to 
•006  mm.  (Nyl.) 

Hab.  on  bark  of  trees.  B.  v.  M.,  Vic.  Nat,  Oct.  1877, 
p.  89. 

Var.  validius,  C.  Knight. — Thallus  yellow  or  sulplmreo 
cinerascent,  thickish,  veri-ucoso  unequal.  Apothecia  all 
black  ;  stipe  short  and  thick  (to  "5  mm.  high,  '8  mm.  thick)  ; 
capitulum  turbinato  lenticular,  disk  lecideine,  to  "5  mm. 
broad.  Spores  fuscous,  ellipsoid,  sub-acuminate  at  each 
apex,  somewhat  constricted  in  the  middle,  uniseptate,  "007 
to  -013  X  -003  to  -OOG  mm. 

Hab.  on  wood  and  decorticated  decaying  trunks  of 
eucalyptus,  MafFra,  Lakes  Entrance,  Bright,  Beechworth. 

Var.  2.  'pevhreve,  Wilson. — Thallus  flavo  virescent,  crust- 
aceous,  rugose.  Apothecia  very  short,  nearly  sessile. 
Sporal  mass  black,  protruded  horizontally  until  the  apothecia 
are  often  conjoined.  Spores  fusco  nigrescent,  ellij)soid,  otten 
narrov/er  at  each  apex,  somewhat  constricted  in  the  middle, 
uniseptate,  "007  to  "015  x  '003  to  •0075  mm. 

Hab.  on  decaying  eucalyi)tus  wood,  Maflra. 

22.    G.  tricolor,   Wilson. 

Thallus  sulphureous,  leproso  grannlose.  A})othecia  black, 
small  (to  "75  mm.  high)  ;  stipe  1  mm.  thick ;  capitulum 
turbinato  globose,  '2  to  "5  nnn.  broad,  margin  white.  Spores 
fuscous,  fusiformi  ellipsoid,  sub-acuminate  at  each  apex, 
somewhat  constricted  in  the  middle,  uniseptate,  with  a  fusco 
nigrescent  locule  in  each  cell,  -008  to  ^012  x  "003  to  -005  mm. 

Hab.  on  decaying  decorticated  eucalyptus,  Wariniambool. 
Lakes  Entrance. 

23.    C.  flavidum,  Wilson. 

Thallus  yellow  or  sulphureous,  crustaceous,  smooth,  nearly 
shining.  Apothecia  black,  but  with  mai-gin,  and  often  the 
stipe  tinged  with  the  thalline  colour,  1  mm.  high  ;  stipe 
5  mm.  thick,  tapering  downwards  ;  capitulum  globoso 
turbinate.      Sporal   mass   black,    protruded.      Spores    fusco 


The  Lichen:^  of  Victoria.  1G9 

nigrescent,    ovoid    or    snb-fusifornii    ellipsoid,    iiuiseptate, 
epispore   rubescent,    cells    fascescent,    containing    each    one 
or  two  nigrescent  locuies,  -OOH  to  -Ol   x  'OOS  to  •004. 
Hab.  on  decorticated  eucalyptus,  Lakes  Entrance. 


Genus  3. — Coniocybe,  Ach.  Nyl. 

Thallus  leprose  or  powdery,  eifuse  or  evanescent.  Apo- 
thecia  yellow  or  pale,  not  black,  stipitate,  excipulum  very 
open.  Spores  usually  spherical,  colourless  or  Havescent, 
forming  the  globoso  pulverulent  capitulum. 

1.  C.  citriocepkala,  Wilson. 

Thallus  white,  thin.  Apotliecia  minute ;  stipe  black, 
slender,  1  mm.  high,  06  thick,  often  less.  Capitulum  flavo 
virescent,  turbinate,  at  length  globose,  to  '2  mm.  diameter, 
becoming  fuscous  and  turbinate  when  stripped  of  the  sporal 
mass.  Spores  colourless  or  dilutely  flavid,  briefly  oblongo 
ellipsoid  or  spheroidal,  -02  to  -04  x    02  to  03  mm. 

Hab.  on  dead  wood  and  dead  bark  of  trees,  Lakes 
Entrance. 

2.  G.  oclivocepkala,  Wilson. 

Thallus  whitish,  often  with  green  leprose  granules. 
Apothecia  with  slender  fuscous  stipe,  "6  to  12  mm.  high, 
•06  to  -08  mn).  thick,  often  bifurcate  ;  capitulum  globose,  pale 
ochre,  '25  mm.  diameter.  Spores  colourless  or  dilutely 
fuscescent,  globose,  '002  to  'OOy  nun.  diameter,  containing  a 
central  locule. 

Hab.  on  decaying  decorticated  Aster  argyrophyllus,  Senecit 
bedfordii,  and  eucalyptus,  Mt.  Macedon  and  Korumburra. 

3.  C.  rliodocephala,  Wilson. 

Thallus  white  or  whitish  or  cinerascent  or  cinereous  or 
evanescent.  Apothecia  often  caspitoso  congested ;  stipe  scarlet 
or  hyaline,  at  length  fuscous  or  black  and  pruinose,  contorted 
and  compressed,  to  2  mm.  high,  ^2  mm.  thick,  sometimes 
furcate  or  two  partly  coalescent.  Capitulum  globose,  obscurely 
rufous  or  fuscous,  at  length  rose  or  flesh  colour,  pruinose, 
rarely  albid,  •4  to  ^8  mm.  diameter.     Spores  very  numerous, 


170      Froceedhif/s  of  the   Hoi/al  Society  of  Victoria. 

colourless,  ellipsoid  oi-  ovate  -OOo  to  'OOG  X  ■001 5  to  'OO-i  mm., 
bilocular  or  placodine  or  uuiseptate,  with  a  locale  in  each 
cell,  epispore  thick.     Paraphyses  numerous,  distinct. 

Hab.  on  dead  wood  or  hark  of  tree.  Lakes  Entrance. 

This  species  is  remarkable  for  the  form  of  its  spores.  All 
the  genus  hitherto  described  have  spherical  simple  spores. 

The  var.  rubens  reported  hy  me  (Trans.  Lin.  Soc),  having 
been  since  found  in  larger  quantity,  proves  to  be  scarcely 
more  than  a  juvenile  form. 

4,    G.  (jracilenta,  Ach.  var.  leucocephala,  Wilson. 

Tliallus  green,  conglomerato  leprose.  Apothecia  with 
stipe  long  and  tender  (lo  to  2"o  x  i  mm.),  fusco  nigricant, 
opacpie,  and  ca|)itnlum  small,  sporal  mass  white,  irregularly 
clothing  the  capitulum  and  the  upper  part  of  the  stipe. 
Spores  colourless,  minute,  sphieroidal,  "002  to  -003  mm. 
diameter. 

Hab.  on  earth,  Fernshawe,  Mrs.  Martin,  March  1891. 

Genus  -i.— Thachyija,  Fr.  pr.  p.  Ny], 

Tludlus  thin,  granulose  or  subleprose,  or  foreign.  Apo- 
thecia 'black,  sessile,  cupulitorni,  open,  sporal  mass  black. 
.S})ores  nigricant  or  fuscous  black,  ellipsoid  or  oblong,  uni- 
septate,  rarely  ])luriseptate. 

1.    T.  leauiorina,  Wilson. 

Tludlus  cinerascent,  verruculoso  leprose,  passim  verrucoso 
sorediate,  gonidia  moderate  and  abundant.  Apothecia  small 
(to  -5  mm.  broad),  crowded,  elevated  in  thalline  receptacles. 
Sporal  mass  black,  abundant,  often  much  protruded  and 
connecting  the  apothecia  Spores  fusco  nigricant  or  nigres- 
cent, or  nearly  colourless,  elli|)soid,  uniseptate,  Dl  to  '02 
X   -OOG  to  -01  nim. 

Hab.  on  old  eucalj'ptus  fences,  Cheltenham,  Yalla-y-poora, 
near  Streatham. 

This  plant  bears  at  first  sight  a  great  resemblance  to 
Lecanoiu  atra. 

2.    T.  viridilocularis,  Wilson. 

Thallus  obscurely  cinerascent.  Apothecia  black,  some- 
what  elevated,    '3   mm.  high,    -3   mm.    broad.     Sporal  mass 


The  Lichens  of  Victoria.  171 

abundant.  Spores  nigricant  or  viresceuti  nigricant,  irregu- 
larly ellipsoid,  uniseptate,  01  to  -02  x  -008  to  Ol  mm., 
with  one  or  rarely  two  loeules  in  each  cell. 

Hab.  on  sawn  eucalyptus  rails,  Kew. 

This  lichen  is  associated  with  Calicium  nigaruin,  var- 
iiunutam,  the  Gallciuvi  on  the  more  decayed  horizontal 
tace  of  the  squared  rail,  and  the  Trachylia  on  the  perpen- 
dicular lace.     The  spores  ai-e  tinged  bottle  green. 

8.    T.  emev(jens,  Wilson. 

Thrtllus  white  or  whitisli,  thin,  smooth,  somewhat  shining. 
Apothecia  seem  to  emerge  fi'om  among  the  fibres  of  the 
wood,  and  at  length  barely  stand  out  above  the  thallus, 
to  -o  mm.  broad.  Sj)ores  iusco  nigricant,  adhering  closely 
together,  ellipsoid,  about  DOo  x  'OOo  mm.,  but  very  various 
in  size,  uniseptate,  with  a  locule  in  each  cell. 

Hab.  on  eucalj^ptus  rails  in  sub-Alpine  regions,  Mount 
Macedon.  Found  also  on  Mount  Lofty,  in  South  Australia, 
and  Mount  Wellington,  in  Tasmania. 


4.    T.    V I ciori<  1,11(1,  Wilson. 

Thallus  cinerascent,  Ifhin,  effuse.  Apothecia  tj'pically 
sessile,  but  often  very  briefly  stipitate,  to  4  mm.,  high,  disk 
generally  sulphureo  pruinose,  to  "4  mm.  broad.  Spores 
fuscous,  oblongo  ellipsoid,  OOo  to  -OOG  X  "OOS  to  -OO^  mm., 
uniseptate,  with  a  nigrescent  locule  in  each  cell. 

Hab.  on  old  eucalyptus  rails.  The  fuscous  epispoie  readily 
rubs  off,  leaving  the  spore  nigrescent,  ellipsoid,  narrow  at 
each  apex,  and  smaller,  •004  to  •OO.')  x  "002  to  -003  mm. 

5.    T.  e.rigua,  Wilson. 

T.  exifjua,  Wilson  (Trans.  Lin.  Soc,  1S.90)  on  further 
examination  proves  not  to  be  a  Trachylia. 

TlilBE    2, — SPH.EROPHOREI. 

Thallus  fruticulose,  ramose  and  raraulose,  the  apices  sub- 
globoso-incrassate,  enclosing  the  apothecia,  which  are  nuclei- 
form,  enclosed,  ultimately  exposed  by  the  bursting  of  the 
thalline  coverino-. 


172      Proceedings  of  the  Royal  Society  of  Victoria. 

Genus  1. — Sph/Erophouon,  Pers. 

Thallus  casspitoso  fruticulose,  sniootb,  polished,  fragile. 
Apotliecia  in  the  apices  of  the  thallus,  receptacle  irregularly 
dehiscent.  Spores  nigricant  or  violaceo  nigricant,  spherical 
or  sub-globose,  covered  with  a-  black  powder. 

1.    &.  australe,  Laur.  =  S.  ceiunoides,  Hampe. 

Thallus  to  2  inches  long,  pallid,  ramose,  branches  com- 
pi'essed,  explanate,  often  distichously  ramulose,  beneath  albi- 
cant,  rugose.  Receptacle  "6  to  "12  mm.  broad,  lenticulari 
compressed,  external  margin  above  cristato  crenate.  Spores 
Oil  to  Olo  mm.  cliam. 

Hab.  Sealer's  Cove,  by  Dr.  F.  Mueller ;  Moe  ;  S.  cera- 
noides,  Hampe,  in  Linnsea  (1856),  XXVIII,  p.  217;  S. 
australe,  Miill.  Lich.  Beitr.,  XVII,  p.  1  ;  B.  v.  M.,  Vic.  Na.t, 
1887,  p.  89. 

Var.  proliferuin,  Wilson. — Thallus  cferuleo  pallid  on 
upper  surface,  convex,  smooth,  nearly  shining,  under  surface 
white,  fo.ssulato  canaliculate  or  scrobiculato  unequal,  to  3 
inches  long,  sub-yjinnatifid,  blanches  linear,  2  to  7  mm., 
broad,  variously  divided.  Apothecia  on  the  under  surface 
of  the  thallus,  the  margin  branching  and  proliferous  once  or 
twice. 

Hab.  on  the  trunks  of  large  trees  in  shady  woods,  Black 
Spur,   Warburton. 

The  plant  grows  horizontally  fi'om  the  tree,  then  droops 
downwards,  and  then  bends  gi-acefulh-  upwards,  displaying 
the  apothecia  on  the  under  side  of  the  thallus.  The  proli- 
ferous branches  grow,  one  or  more,  from  the  margin  of  the 
apothecium. 

2.    S.  cornpi'essum.  Ach. 

Thallus  pallido  albicant,  i-amose,  piano  compressed.  Apo- 
thecia obliquely  minute  on  the  ai)ices  ;  receptacle  lacero 
dehiscent,  or  discoid  and  open.  Spores  nigricant,  spherical, 
•007  to  01 1  mm.  diam.  (Nyl.) 

B.  V.  M.,  Vic.  Nat.,  Oct.  ]  887,  p.  89. 

8.    S.  coralhndes,  Pers. 

Krem}).,  in  Verhand.  Zool.  Botan.  Gesellsch.,  in  Wien, 
1880,  p.  329.  A  mistake  for  Stereocaulon  ramulosum, 
according  to  Prof.  J.  Mueller,  in  Ratishoii  Flora,  1887,  No. 
8. 


The  Lichens  of  Victoria.  173 

4.    «S\  fenerum,  Laur. 

Thallus  palo  or  whitish,  terete,  slender,  ver}'  much 
branched,  branches  tine  and  intricate.  Apothecia  small  ("I 
to  2  mm.)  on  the  jirimary  branches  :  thalline  receptacle 
persistent,  only  slightly  dilated.  Spores  nigricant,  or 
smeared  with  a  friable  nigricant  pigment.  Diam.  'OOT  to 
•008  mm.  (Nyl.) 

Ki-emp.,  in  Verhandl.  Zool.  Bot.  Gesells.,  in  Wien,  1880, 
p.  329.  According  to  Prof.  J.  Mueller  (in  Ratishoii  Flora, 
1887,  No.  8),  a  specimen  from  Mt.  Ellery  is  rightly  deter- 
mined, but  specimens  from  Black  Spur  and  Yarra  Yarra  are 
Gladina  aggregata. 

Series  2.— Cladodpzi. 

Thallus  generally  erect.  Apothecia  terminal  on  podetia, 
rarely  sessile  ;  biatorine,  rarely  lecanorine.  Spores  8, 
colourless,  usuallj-  oblong  and  simple,  sometimes  elongate 
and  septate.     Pai-aphyses  distinct. 

Tribe  3.— B^omycei. 

Thallus  horizontally  expanded,  crustaceous.  Apothecia 
pale  or  rufescent,  sessile  or  podetiiformi  stipitate.  Spores 
simple  or  septate. 

Genus  1. — Gomphillus,  Nyl. 

Thallus  very  thin,  consisting  of  gonidia  and  filaments 
irregularl}-  conglutinated.  Apothecia  stipitate,  small, 
corneous.  Spores  filiform,  multiseptate.  Paraphyses  in- 
distinct. 

1.    G.  hceomyceoides,  Wilson  =  Patelkiria  Wihoui, 
Mull.  Arg.* 

Thallus  cinereous  or  virescent,  effuse,  either  very  thin  and 
somewhat  shining,  or  rather  thicker  and  eroso  isidioso 
granulate.  Gonidia  various  in  size  and  form,  conglomerated 
into  gelatinous  globules.  Apothecia  of  a  tenacious  horny 
texture,  biatorine,  sometimes  margined  by  the  white  hypo- 
thecium,  scattered  or  conglomerated,  depresso  globose,  to 
1*5  mm.  diam.,  smooth,  rufo  fulvescent,  pale  when  5T)ung, 

*  Lich.  Beitr.,  in  Flora  1888,  No.  1435. 


174      Proceedings  of  tJte  Royal  Society  of  Victori'i. 

and  dark  in  age,  sub-sessile  or  stijiitate,  stipe  to  '5  mm.  higli 
and  -5  mm.  thick,  with  sometimes  two  or  three  capitiila^on 
one  stipe.  Spores,  eight  in  cylindrical  thecse,  acicuhui 
filiform,  about  -i-i  mm.  long,  ))luriseptate. 

Hab.  on  roots  and  trunks  of  trees,  upon  mosses  and  bark, 
and  jungermannias  and  lichens ;  also  on  the  earth  upon 
dead  leaves,  &;c.,  in  shady  mountain  forests,  Black  Spur, 
Mt.  Macedon,  Warragid. 

Genus  2. — B.eomycks,  Pers. 

Thallns  crustaceous,  powdery,  granulose  or  squamulose. 
Apothocia  biatorine,  sessile  or  stipitate. 

].   B.  rufii.'i,  i).  C. 

Thallus  albo  virescent  or  albido  glaucescent,  thin,  effuse, 
minutely  granulose  or  squamulose  or  leprose,  granules 
depressed  (K  yellow).  Apothecia  carneo  rufescent  or 
carneo  fuscescent,  somewhat  convex,  immarginate,  stipe 
moderate  or  very  short,  whitish.  Spores,  6  or  8,  oblongo 
ellipsoid,  simple. 

B.  v.  M.,  Vic.  Nat,  Oct.  1887,  p.  8!). 

2.    B.  fusco  caruca,  Wilson. 

Thallus  pallid,  granuloso  verrucose,  granules  sometimes 
depressed.  Apothecia  rufo  fuscescent,  quasi  pruinose,  1  to 
2  mm.  broad,  convex,  margined  by  the  hypothecium.  Stipe 
white,  nude,  short  (less  than  1  mm.  high).  K.  thai,  and 
apoth.  yellow,  then  blood  red.  Spores  ellipsoid,  simple. 
•008  to  -01  X   003  to  -005  mm. 

Hab.  on  clay  ground,  Kilmore. 

3.    B.  roseu.'i,  Pers. 

Thallus  whitish,  granulose,  eti'use  or  determinate.  Apo- 
thecia roseo  carneous,  or  albo  carneous,  nearly  globose,  about 
2  mm.  broad,  stipe  whiter  or  nearly  white,  subterete.  Spores 
six  or  eight,  fusiformi  oblong  or  fusiform,  simple,  "Oil  to  '02(1 
X  -0025  to  -003  (ISIyl.)     Paraphyses  slender. 

Hab.  on  bare  earth,  chieHv  clay.  B.  v.  M.,  Vic.  Nat, 
Oct.  1887,  p.  89. 

(See  note  on  next  species.) 


The  Lichoi.s  of  Victoria.  175 

4.    B.  fuujjoides,  Ach. 

Thallus  whitish,  granulose,  margin  of  granules  spreading, 
thin,  continuous.  Apothecia  roseo  carneous  or  albo  carneous, 
sub-globose,  or  globoso  elavate,  or  cliff ormi  clavate,  moderate 
or  large  (2  to  4  mm.  broad)  ;  stipe  long  (-i  to  8  mm.),  whiter 
or  nearly  white,  subterete.  Spores  oblong  or  fusiform, 
simple,  -Oil  to  •023  x  -0035  mm.     Paraphyses  slender. 

Halx  on  earth,  chiefly  clay,  in  mountain  regions,  Otway 
Ranges,  Black  Spur,  Warburton,  Mt.  Lookout,  (A.  F.  Wilson). 

Probably  a  variety  of  B.  roseu.^,  growing  in  a  warmer 
climate,  as  Tuckerman  suggests.  When  not  well  developed 
it  approaches  the  previous  species. 

5.   B.  heteromorjjltiis,  Nyl. 

Thallus  pallido  glaucescent  or  pallido  cinerascent,  verrucoso 
unequal,  forming  large  patches.  Apothecia  pale  carneous, 
or  carneo  fuscescent,  -5  to  1  mm.  broad,  margin  thick, 
undulate,  obtuse,  stipe  1  to  2'5  mm.  high,  variously 
compressed  or  plicate,  often  two  to  six  or  more  apotliecia 
on  one  stipe.  Spores  vety  transparent,  nearly  indistinct, 
ellipsoid,  simple,  01  x  "006  mm.    Thall.  and  Apoth.K.  +  C — . 

Hab.  on  cla}^  ground,  m<3sses,  dead  leaves,  &c.,  in 
mountain  regions.  Black  Spur,  Mt.  Macedon,  Warburton. 
Otway  Ranges,  Lilydale,  Mt.  Buffalo  (A.  F.  Wilson). 

6.   B.  squamarioides,  '!:iy\.  =  Kni;/htiella  leucoco  lya  = 
K.  squamarioides,  Mull.  Arg. 

Thallus  albo  or  albido  glaucescent,  subopacjue,  squamosa, 
sfjuamse  difformed,  about  5  mm.  broad,  affixed  (forming 
small  patches  about  an  inch  wide),  lobate  or  lobato  incised, 
plane  or  somewhat  depressed  in  the  centre,  concolorous 
beneath  or  whiter.  Apothecia  lurid  or  pale  lurid  or  lurido 
carneus,  "2  to  "3  mm.  broad,  biatorine,  plane,  margin  thickish, 
evanescent.     Spores  oblong  or  fusiformi  oblong,  uniseptate. 

Hab.  on  earth,  Mt.  William  (D.  Sullivan).  Mull.  Lich. 
Beitr.,  1888,  No.  13,  p.  8. 

7.    B.  FreRchlamis,  Mull.  Arg. 

Thallus  squamose  ;  squama3  Ctuspitose,  crowded,  broad, 
inciso  lobate  ;  lobes  ascending,  crenulate  or  entire,  olivaceous 
above,  white  beneath,  bearing  podetia  here  and  there  upon 


17G      Froceedingi^  of  the  Royal  Societu  of  Victoria. 

theii"  margins.  Pocletia  aVjoufc  2  mm.  high,  '60  mm.  thick, 
olivaceous,  thallino-corticate  and  sub-granuloso  asperulate, 
or  often  towards  the  apex  decorticate  and  somewhat  rose- 
coloured,  monocephalous.  Apothecia  about  equally  broad 
with  the  podetia ;  the  whole  of  the  apothecium  at  first 
rosello  fuscescent  and  very  widely  truncato  obconical  and 
plane,  but  soon  fuscous  and  convex.  Spores  not  fully 
evolved.     Prof.  J.  Mueller  in  Ratisbon  Flora. 

Genus  3. — Thysanothecium,  Berk,  and  Mont. 

Thallus  partly  horizontal,  granulose  or  squamose,  and 
partly  podetiiform,  often  expanding  at  the  apex,  variousl}'' 
divided.  Apothecia  thin,  ])ale  or  rufous,  darker  or  lighter, 
terminating  the  terete  podetium  or  covering  the  upper 
surface  of  the  frond-like  podetium.  Spores  small,  ellipsoid, 
simple. 

1.    T.  hyaliuum,  Tajdor. 

Thallus  pale  yellow  or  pale  lurid,  lobato  granulose  or 
squamose ;  podetia  various  in  size  (1  to  12  mm.  high,  -5  to 
2  mm.  thick),  sulcato  rugose,  sometimes  squamulose  below  ; 
apex  dilated  on  one  side  (J  to  10  mm.  broad).  Apothecia 
pallid  or  carneo-rufescent,  or  fusco  rufus,  forming  a  thin 
stratum  on  the  upper  surface  of  the  apex.  Spores  8  in  the 
thecEe,  ellipsoid,  simple,  -006  to  "008  x  -0035  (Nyl.) 

Hab.  on  earth  or  decayed  and  generally  burnt  wood, 
common  ;  Kew,  Box  Hill,  Youyangs,  Cobden,  Mordialloc, 
Cheltenham,  Oakleigh,  Ringwood,  Lilydale,  M.atfia. 

Form  squamulosum,  Wilson. — Thallus  yellow,  more  or 
less  sordid  or  lurid,  squamulose,  squamules  thick,  either 
depressed  and  lobate,  or  somewhat  ascending  podetiiform, 
swollen  upwards  and  briefly  ramose.  Apothecia  cephaloid, 
sessile  on  the  squamules  and  podetia,  minute,  crowded,  not 
fully  evolved. 

Hab.  on  poor  soil,  Trentham  (coll.  by  Mi-s.  Martin),  Kew, 
Sandringham.     Possibly  a  ncM^  species. 

Form  intortum,  Wilson.— Thallus  yellow,  squamulose, 
podetia  compressed  and  dilated,  lobulate,  apices  crispate  and 
intorted.  Apothecia  as  in  the  typical  form.  Spores  not  full}^ 
evolved. 

Hab.  on  decaying  wood,  Oakleigh.     Coll.  by  Mi-s.  Martin. 


The  Lichens  of  Victoria.  177 

•1.    T.  Hool-erl,  Berk,  et  Mnt. 

Thallus  lurido  iiaveseent,  iiinato  oraniforni,  effuse.  Podetia 
cinereo  tlavescent,  about  i  inch  liigh,  frondose,  stipitate, 
firm,  sti'iato  nervose,  stipe  subterete,  about  6  mm.  high, 
dilated  above  into  a  simple  or  lobed  frond.  Apothecia 
tbinly  but  equally  covering  the  one  side  of  the  frond,  rufus 
or  carneo-rufus  or  testaceo  carneous,  innnarginate.  Spores 
8,  ellipsoid,  -006  to  -007  x    0025  to  -008  mm". 

Hab.  on  earth,  near  sea,  Cheltenham. 


Art.  XIII. — On  a  New  Species  of  Leucosolenia  from  the 
neighbourhood  of  Fort  Phillip  Heads. 

By  Arthuk  Dendy,  D.  Sc. 

[Read  December  8,  1892.] 


The  species  here  described  was  collected  by  Mr.  J. 
Bracebridge  Wilson,  M.A.,  in  the  neighbourhood  of  Port 
Phillip  Heads,  bvit  unfortunately  too  late  for  it  to  be 
included  in  Part  I  of  the  Monograph  of  the  Victorian 
Sponges,  which  deals  with  the  group  (Homocoela)  to  which 
it  belongs. 

Leucosolenia  uteoides,  n.  sp. 

In  external  form  and  canal  system  tlie  sponge  very 
closely  resembles  Leucosolenia  stolonifer,  Dendy,*  belonging, 
like  tlie  latter,  to  the  section  of  the  genus  Leucosolenia  to 
which  I  have  proposed  to  apply  the  name  Simplicia.  The 
single  specimen  is  colonial,  consisting  of  about  one  hundred 
individuals  united  together  by  their  bases  only  and  rising 
vertically  upwards  side  by  side  so  as  to  form  a  compact 
colony.  The  spongorhiza  is  not  conspicuous,  being  repre- 
sented by  the  union  of  the  various  individuals  at  their 
bases.  From  the  basal  portions  of  the  individuals,  thus 
united,  arise  numerous  short,  slender,  downward-growing, 
tubular  processes,  which  apparently  serve,  as  in  L.  stolonifer, 
to  attach  the  colony  to  the  substratum.  The  fully  developed 
Ascon  individuals  attain  a  height  of  about  85  mm.  and  a 
diameter  of  about  2  5  mm.  Each  is  a  nearly  straight, 
slender,  cylindrical,  thin-walled  tube,  narrowing  slightly 
towards  the  naked,  terminal  osculum.  The  tubes  may 
branch,  especially  near  their  bases.  Under  a  lens  the  outer 
surface  of  each  tube  appears  very  slightly  hispid  and  also 
exhibits  that  longitudinal  striation,  due  to  the  presence  of 
large  oxeote  spicules,  which  is  so  characteristic  of  the  genus 

*  "  Monograph  of  the  Victorian  Sponges,"  Part  I,  p.  46,  Plate  I,  Fig.  2, 


On  a  Keiu  Species  of  Leucosolenia.  17.9 

Ute,  whence  the  specific  name  lUeoides.  The  wall  of  the 
tube  is  about  O'lo  mm.  thick,  the  mesoderm  being,  as  in 
L.  stolonifer,  very  strongly  developed  for  a  Homocoel 
sponge. 

The  skeleton  consists  of  quadriradiate  and  two  kinds  of 
oxeote  spicules.  The  quadrii-adiates  are  arranged  as  usual 
in  the  thickness  of  the  mesodei'm  towards  the  inside  of  the 
sponge- wall,  the  facial  rays  lying  parallel  to  the  gastral 
surface,  the  basal  ray  directed  away  from  the  osculum,  and 
the  apical  ray  projecting  into  the  gastral  cavity.  These 
spicules  are  markedly  sagittal,  the  oral  rays  being  widely 
extended  and  distinctl}^  recurved  towards  the  basal.  All 
three  facial  rays  are  long  and  slender,  but  the  basal  is  much 
more  so  than  the  orals  and  is  slightly  hastate  ;  all  three  are 
fairly  sharply  pointed.  In  an  average-sized  spicule  the  oral 
rays  measure  about  O'lSG  b}'  0-0082  mm.  (near  the  base) 
and  the  basal  about  0"31  by  O'OOG  mm.  (near  the  base),  but 
of  course  there  is  a  good  deal  (^f  variation,  and  I  have 
measured  the  basal  ray  up  to  ()42  mm.  in  length.  The 
apical  rays  are  very  strongly  developed ;  long,  slender  and 
sharply  pointed  ;  usually  more  or  less  crooked  and  varying 
greatly  in  length  ;  the  average  length  is  perhaps  about 
Oio  nmi.,  but  this  is  often  greatly  exceeded. 

The  oxeote  spicules  may  be  divided  into  two  classes  accord- 
ing to  their  shape,  size  and  position  in  the  sponge.  (1)  Very 
large  spindle-shaped  oxea,  completely  imbedded  in  the  outer 
portion  of  the  sponge  wall  and  arranged  parallel  to  the  long 
axis  of  the  sponge.  These  spicules  are  usually  straight 
and  symmetrically  fusiform,  very  thick  in  the  centre  and 
tapering  gradually  to  a  fine  point  at  each  end.  Fully  grown 
examples  measure  a  little  over  1  mm.  in  length  and  about 
()'()65  mm.  in  greatest  thickness  (in  the  centre).  They  are 
placed  pretty  close  together  side  by  side  in  a  single  layer. 
(2)  Much  smaller  oxea  projecting  from  between  the  large 
ones  and  abundantly  echinating  the  outer  surface  of  the 
sponge.  These  spicules  are  rather  slender,  often  slightly 
curved  or  even  crooked,  fairly  gradually  sharp-pointed  at 
each  end,  but  with  the  outer  end  often  bent  slightly  though 
sharply  to  one  side,  like  a  bayonet  ;  size  about  022  mm.  by 
0-008  mm.  The  colour  of  the  sponge  in  spirit  is  yellowish- 
white. 

As  already  pointed  out  this  species  is  nearly  related  to 
my  Leucosolenia  .stolonifer,  but  it  appears  to  be  even  more 

N  2 


180      Proceedings  of  the  Royal  Society  of  Victoria. 

nearly  related  to  Carter's  L.  (Laconoides*  with  which  it 
ao-recs  not  only  in  general  form  but  also  in  the  C/'te-likt' 
armour  of  huge  spindle-shaped  oxea.  In  L.  asconoido^, 
however,  there  appear  to  be  none  of  the  smaller  oxea  which 
so  abundantly  echinate  the  dermal  surface  of  our  species, 
while  the  large  oxea  are  nearly  twice  the  size  of  those  of 
L.  uteoides.  It  is  a  curious  fact  that  in  L.  asconoides, 
"more  or  less  of  the  arms"  of  the  quadriradiates  art- 
"  exserted  between  the  long  acerates,  so  as  to  give  this  part 
a  minutely  hispid  appearance.  At  first  sight  the  lattei"  look 
like  raortar-spicules  or  small  acerates,  but  although  they 
appear  to  serve  the  same  jjurpose,  they  are  not  so,  but  what 
I  have  stated."  f  In  view  of  this  verj-  definite  statement  it 
appears  tolerably  certain  that  L.  uteoides  is  specifically 
distinct  from  L.  asconoides. 

*  Vide,  "Monograph  of  Victorian  Sponges,"  Part  I,  p.  48. 

t  Carter,  "  Annals  and  Magazine  of  Natural  History,"  August  1886,  p.  13'>. 


Art.  XIV. — Tlte  Present  Pvsition  of  the  Snake-bite 
Controversy. 

By  James  W.  Barrett,  M.D.,  M.S.,  F.R.C.S.  Eng. 

Demonstrator  and  Examiner  in  Physiology  in  the  University  of  Melbourne. 
[Read  November  10,  1892.] 


Tlie  public  and  the  technical  press  have  of  late  been 
occupied  with  discussions  on  the  merit  or  demerit  of  the  so- 
called  strychnine  cure  for  snake-bite,  but  as  usual,  very  little 
definite  evidence  has  been  adduced.  I  have,  therefore, 
thought  it  advisable  to  bring  the  facts  of  the  case  under  the 
notice  of  the  members  of  this  Society,  so  that  the  position 
occupied  by  the  rival  disputants  may  be  rendered  perfectly 
clear.  Dr.  Mueller  of  Yackandandah,  it  seems,  has  satisfied 
himself  that  a  theory  respecting  the  action  of  snake  poison 
has  been  proved.  He  believes  that  strychnia  is  consequently 
indicated  as  a  remedy.  When,  however,  he  is  asked'[to 
substantiate  both  these  projiositions,  by  showing  that  the 
treatment  is  successful,  he  has  no  further  evidence  to  adduce 
than  the  report  of  cases  of  snake-bite,  real  or  supposed,  in 
which  medical  men  assert  that  patients  were  saved  from 
death  by  the  injection  of  strychnine.  Now,  it  is  obvious 
that  before  reports  of  such  cases  can  be  of  much  value,  it  is 
necessary  to  ascertain  the  percentage  of  individuals  who 
died  from  snake-bite  when  other  modes  of  treatment  were 
adopted.  In  other  words,  snake-bite  is  or  is  not  a  very  fatal 
affection. 

The  object  of  this  communication  is  to  endeavour  to  make 
answer  to  that  question.      In  investigating  it,  I  have  had 


182     Proceedings  of  ike  lioyal  Society  of  Victoria. 

extensive  recourse  to  tables,  furnished  to  me  by  the  ever 
obliging  Government  Statist,  Mr.  Hayter. 

Table  I,  which  follows,  shows  the  deaths  which  have  taken 
place  fi'oni  snake  and  insect  bite  (for  the  two  are  bracketed 
in  returns  together)  in  the  Australian  colonies  during  the 
decade  1881-1890.  In  accordance  with  the  foot-note 
appended  to  this  table,  I  have  rejected  from  further 
consideration  any  deaths  occurring  in  other  colonies  than 
Victoria,  New  South  Wales,  and  Queensland.  You  will 
further  note  that  of  the  total  1 25  deaths  which  occuri'ed  in 
these  three  colonies  in  the  period  mentioned,  at  least  5  or  6 
are  obviously  due  to  bites  of  other  animals  than  snakes. 
There  is  the  further  probability  tliat  some  of  the  deaths 
have  been  caused  by  the  enthusiastic  administration  of  alcohol 
to  persons  bitten  or  supposed  to  be  bitten.  However,  to  be 
well  within  the  mark,  I  assume  that  125  deaths  represent 
fatal  cases  of  snake-bite,  and  proceed  to  deal  with  them 
accordingly. 


Table   I. 

Deaths  from  Snake  and  Insect-bite   in  iite  Australian  Colonies, 
1881   tn   1890. 


Years. 

Victoria." 

N.  S. 

Queens 

South 

West 

Tasmaiiiat      Total. 

Wales. 

land. 

1681 

5 

5 

5 

1                 16 

1882 

5 

. 

3 

1 

1               10 

1883 

2 

4 

1                7 

1884 

3 

8 

•• 

11 

1885 

3 

3 

3 

.. 

1 

10 

1886 

4 

5 

u 

1887 

9: 

3 

8 

1 

21 

1888 

3 

0 

4 

12 

1889 

2 

4 

11 

1 

18 

1890 

2 

lU 

() 

1 

19 

Total 

38 

47 

40 

2 

2 

4        1     133 

ns 

1 

Note. — There  are  no  deaths  from  snake-bite  in  New  Zealand. 

In  Victoria,  in  1891,  there  were  .5  deatlis  from  snake-bite,  and  1  from  iguana-bite. 

*  In  other  years  than  1881  and  1882,  no  distinction  was  regularly  made  in  Victoria  between 

snake  and  insect-bites.     Two  of  the  deaths  in  the  former  yeai-,  and  1  in  the  latter,  were  from 

insect-bite. 

t  In  the  case  of  these  colonies,  it  i.s  not  certain  whether  there  were  any  deaths  in  several 
of  the  years,  as  the  cause  was  not  specifically  mentioned  in  the  list  of  causes  of  death, 
t  One  of  these  is  distinguished  as  "  vermin-bite  "  and  1  "  insect-bite." 


Present  Fosltlon  of  the  Snakc-hite  Controversy/.    183 

It  will  be  seen  that  in  this  period,  in  the  three  colonies, 
snakes  were  unable  to  kill  more  than  125  persons. 

In  order  to  determine  the  relative  frequency  of  death 
from  snake-bite,  I  next  append  a  table  showing  the  popula- 
tion (actual  and  average)  of  the  colonies  during  the  same 
period. 


Table    II. 
JlJeaa  Populations  of  the  Australasian  Colonies,  1881   to  1890. 


Yr,^ 

N.  S. 

Queens- 

Soutli 

West 

New 

Total  Aus- 

Wales. 

land. 

Australia. 

Australia. 

Zealand.* 

tralasia. 

1881 

868,942 

765,0151  226,522 

276,948 

29,516 

116,437 

492,887 

2,776,262 

1882 

889,720 

798,5401  237,611 

289,916 

30,389 

119,473 

509,308 

2,874,957 

1883: 

910,1301    838,15o|  267,865 

299,012 

31,233 

122,242 

529,292 

2,997,929 

1884 

932,6301    883,14o|  294,782 

308,648 

32,329 

125,352 

548,993 

3,125,879 

1885 

956,880,    927,275;  308,789 

313,102 

34,072 

128,160 

566,168 

3,234,416 

1886? 

981,860;    969,455  327,034 

311,254 

37,184 

130,441 

.582,306 

3,342,735 

1887    : 

1,016,750:1,004,835  346,545 

311,050 

41,699 

133,802 

596,373 

3,450,391 

1888 

1,054,9801,035,705  361,230 

312,253 

42,312 

137.167 

605,370 

3,549,017 

1889 

1,090,350  l,066,450j  374,240 

313,751 

43,053 

140,261 

611,716 

3,639,685 

1890^ 

1,118,500!  1,101,840;  385,805  |  316,425 

47,950 

143,733 

620,780 

3,734,685 

Average 

982,374 

939,041  313,042 

305,235 

36,973 

129,706 

566,319 

3,272,599 

Therefore,  the  proportion  of  fatal  cases  of  snake-bite  to  the 
average  number  of  persons  alive  during  the  period  is  shown 
by  the  following  table,  which  gives  :  — 


Table  III. 


The  Ratio  of  Deaths  from   SnaJce-hite   in    each    Goloiiy  during  the 
Decade  to  the  average  Population. 


Victoria 

New  South  Wales 
Queensland 
Avorase 


1  to  25,852 

1  to  19,980 

1  to  7,826 

1  to  17,886 


The  death-rate  from  snake-bite  in  Queensland  seams  very 
much  hioher  than  in  the  other  two  colonies.     The  following 


184      I'roceediiuj.s  of  (he  lioi/al  Sociefi/  of  Victoria. 

table    will,    however,    show    the    danger   of    drawing    rash 
conclusions  from  figures  : — 

Table    IV. 

Deaths  from    Violence  in  the  Australian  Colonies,   1881   to  1890. 


Vkars. 

Victoria. 

N.  S. 

Queens- 

South 

West 

New 

Total 

Wales. 

laud. 

Austialia. 

Australia. 

Zealautl. 

1881 

849 

906 

817 

288 

36 

to. 

459 

2,905 

1882 

811 

904 

489 

210 

58 

88 

505 

3,040 

1883 

908 

850 

896 

202 

59 

lOG 

494 

3,015 

1884 

799 

990 

50.) 

289 

51 

90 

548 

3,22(; 

1885 

816 

1,106 

492 

212 

45 

92 

517 

3,310 

1886 

942 

1,088 

496 

272 

67 

94 

571 

3,525 

1887 

l,C'2;-5 

1,148 

599 

2-9 

57   . 

112 

555 

3,728 

1888 

1,119 

1,140 

598 

284 

90 

118 

513 

3,807 

1889 

l,18fi 

1,110 

622 

203 

44 

144 

508 

3,822 

1890 

1,165 

1,168 

787 

238 

48 

138 

521 

4,005 

Total 

9.678 

10,400 

5,200 

2,277 

545 

1,087 

5,191 

34,378 

Table  V. 

l^he  Ratio  of  the    Total  Deaths  from  Snake-hite  ditring  the  Decade 

to  the   Total  Deaths  from    Violence. 

Victoria         -         -         .         - 
New  South  \V;ik's 
(i)ueenHlan(l   -         -         .         - 
Average 

The  following  table  shows  the  ratio  of  the  total  deaths 
from  violence  during  the  decade  to  the  average  population:  — 


1 

to 

254-7 

1 

to 

221-8 

1 

to 

180 

1 

to 

202 

Table  VI. 


Victoria 

New  South  Wales 
Qnecnshmd   - 
Average 


1  to  101-5 

1  to  90-8 

1  to  61-2 

1  to  84-3 


It  will  thus  be  seen  that  although  the  ratio  of  deaths  from 
snake-bite  to  the  average  population,  .and  also  to  the  total 
deaths  from  violence,  is  higher  in  Queensland  than  in  the 
other  two  colonies,  the  ratio  of  deaths  from  violence  to  the 
average  population  is  also  higher.  Consequently  it  is  unsafe 
to  infer,  from  the  evidence  furnished,  thttt  snake-bite  is 
necessarih'  a  more  fatal  affection  in  Queensland  than  in  the 
other  colonies. 


Present  Position  oj  the  Snukc-hite  Controversy.    185 

From  this  mass  of  figures  we  arrive  at  a  general  conclusion 
that  snake-bite  is  one  of  the  most  insignificant  causes  of 
death  in  our  midst.  For  example,  in  the  three  years  1887- 
88-89  more  persons  died  in  Victoria  from  hydatid  disease 
than  were  killed  by  snakes  in  Australia  during  the  decade. 
Anyone  who  cares  to  look  through  Mr.  Hayter's  tables  will 
find  that  the  snake-bite  contribution  is  a  veiy  sraaU  one. 

In  187G,  a  Committee  was  appointed  by  the  Medical 
Society  of  Victoria  which  experimented  in  a  methodical 
way.  The  Committee  consisted  of  Drs.  M'Crea  (Chairman), 
T.  M.  Girdlestone,  E.  Barker,  J.  E.  Neild,  A.  Bowen,  P. 
Smith,  J.  T.  Dempster,  and  Professor  J.  D.  Kirkland.  The 
particular  value  of  the  work  done  by  this  Committee  lay  in 
the  fact  that  it  found,  with  antidotes  then  in  use,  the 
recovery  of  a  dog  from  snake  virus  injected  hypodermically 
was  chiefly  a  matter  of  dosage.  IMone  of  the  dogs  used 
recovered  when  half  a  giain  of  fresh  liquid  poison  was 
injected.  They  further  found  that  tiger  snakes  3tt.  to  4  ft. 
long  injected  on  an  average  from  1  to  H  grains  of  liquid 
poison,  a  quantity  believed  by  analogy  to  be  bai'ely 
sufficient  to  kill  a  man.  One  grain  of  tiger  snake  venom, 
if  injected  fairly  into  the  skin,  would  be  approximately  a 
dangerous  dose.  It  is,  however,  quite  possible  that  a  snake 
driving  its  fangs  through  the  skin  finds  it  difiicult  to 
administer  the  full  dose,  if  the  snake  bites  through  clothing, 
tiie  chances  of  a  fatal  issue  are  diminished.  On  the  other 
hand,  in  the  case  of  some  of  the  Indian  snakes,  allied  in 
chai-acter  to  the  Austi'alian  black  and  tiger  snake,  the  dose 
of  poison  injected  amounts  to  from  10  to  13  grains.  Com- 
ment is  needless. 

Furthermore,  Dr.  M'Crea,  in  187G,  forwarded  a  circular  to 
a  number  of  medical  practitioners  asking  them  for  infoima- 
tion  on  the  suV)ject  of  snake-bite.  In  answer,  he  found  that 
253  cases  of  snake-bite  had  occurred  in  the  practice  of  a 
number  of  medical  practitioners,  and  that  of  tliese  only  25, 
or  10  per  cent.,  terminated  fatall}'.  Various  methods  of 
treatment  had  been  adopted. 

It  seems,  therefore,  that  fatal  results  from  snake-bite  are 
not  common,  and  can  scarcely  take  place  unless  the  condi- 
tions are  favourable  to  the  snake.  Nevertheless,  if  snake-bite 
were  resfwusible  for  onl}'  one  death  in  the  decade,  one  would 
hail  with  pleasure  the  remedy  which  Avould  obviate  the 
repetition  of  such  an  accident;  and  my  object  in  referring 
to  these  figures  is  not  to  under-rate  the  value  of  anj-  remedy. 


ISO      Proceedinrjs  of  the  Royal  Societi/  of  Victoria. 

but  to  «liow  the  difficulty  of  being  accurate  in  forming 
conclusions  respecting  its  value. 

These  iacts  are  so  well-known  that  I  must  apologise  for 
restating  them.  I  have  mentioned  them  in  outline  simply 
as  part  of  the  argument.  In  fact,  if  the  name  of  other 
remedies  used  in  the  past  be  excised  fiom  old  reports  in  the 
Journal,  and  the  word  strychnia  be  substituted,  the  descrip- 
tion would  parallel  the  present  accounts  of  the  efficacy  of 
strychnia. 

If,  then,  a  discoverer  of  a  snake- bite  antidote  has  to  refer 
to  mortality  tables  as  a  proof  of  its  success,  he  has  a  small 
margin  to  work  on.  He  is  dealing  with  a  disease  which  is 
not  usually  intractable. 

The  [)ublic  reports  of  cases  may  be  i-eferred  to  as  evidence 
of  its  value,  but  apart  from  preceding  facts  altogether,  1 
would  ask  anyone  who  is  inclined  to  attacli  any  value  to 
such  statements  to  think  for  a  moment  what  they  mean. 
Men,  women,  or  children  of  diffi3rent  physiological  resistance 
and  vigour  bitten,  or  supposed  to  be  bitten,  by  snakes  ot 
different  age,  biological  characters,  and  virus-producing 
capacity,  the  jnmctures  made  into  skins  of  ditferent  thickness 
and  in  different  parts  of  the  body — treatment  of  various 
kinds  adopted.  Are  there  here  not  enough  variables  to 
cause  grave  doubt  as  to  the  value  of  a  new  variable 
introduced  in  the  form  of  strychnia?  Again,  public  reports 
of  cases  have  been  held  to  prove  such  extraordinary  theories 
in  medical  history  that  one  may  be  pardoned  lor  receiving 
them  with  great  caution.  As  stated,  other  remedies  for 
snake-bite  have  been  similarly  commended  at  the  hands 
of  their  demonstrators  in  the  columns  of  the  Australian 
Medical  Journal. 

'J'here  is  one  method  by  which  the  value  of  strychnia  as  a 
remedy  may  be  settled,  viz.,  by  resort  to  experiments  on 
anim.ils  on  which  the  action  of  snake  poison  does  not  to  all 
appearances  differ  materially  from  that  in  the  case  of  man. 
From  this,  however,  Dr.  Mueller  dissents,  though  he  refers 
to  experiments  made  on  animals  in  support  of  his  theor3\ 

The  evidence  adduced  serves  to  show  that  there  is  no 
wai-rant  for  believing  strychnia  to  be  of  any  value  as  an 
antidote  foi-  snake-bite  ;  but  there  is  no  warrant  for  asserting 
that  it  is  valueless.  By  the  experimental  method  alone,  can 
the  vexed  question  be  settled. 


Art.  XV. — Sneezing:    Fdllacious  Observations. 
By  James  W.  Barrett,    M.D.,   M.S.,  F.R.C.S.  Eng. 

Demoustrator  and  Examiner  in  Physiology  in  the  University  of  Melbourne. 
fReacl  December  8,  1892.] 


In  the  last  edition  of  ''  Foster's  Physiology,"  there  occur 
the  following  passages  : — "  Coughing  consists  in  the  first 
place  of  a  deep  and  long-drawn  inspiration,  by  which  the 
lungs  are  well  filled  with  air.  This  is  followed  by  a  complete 
closure  of  the  glottis,  and  then  comes  the  sudden  forcible 
expiration,  in  the  midst  of  which  the  glottis  suddenly  opens, 
and  thus  a  blast  of  air  is  driven  through  the  upper  respira- 
tory passages.  The  afferent  impulses  of  this  reflex  act  are 
in  most  cases,  as  when  a  foreign  bod}^  is  lodged  in  the  larynx 
or  by  the  side  of  the  epiglottis,  conveyed  by  tlie  superior 
laryngeal  nerve.  But  the  movement  may  arise  from  stimuli 
applied  to  other  branches  of  tlie  vagus." 

"  In  sneezing,  the  genei'al  movement  is  essentially  the 
same  (as  in  coughing),  except  that  the  opening  from  the 
pharynx  into  the  mouth  is  closed  b}'  the  contraction  of  the 
anterior  pillars  of  the  fauces,  and  the  descent  of  the  soft 
palate,  so  that  the  force  of  the  blast  is  driven  entirely 
through  the  nose.  The  afferent  impulse  is  usually  given 
from  tiie  na.sal  branches  of  the  fifth."  When  sneezing, 
however,  is  produced  by  bright  light,  the  optic  nerve  would 
seem  to  be  the  afferent  nerve. 

In  Landois  and  Stirling,  sneezing  is  described  as  consisting 
"of  a  sudden  violent  expiratory  blast  through  the  nose  for 
the  removal  of  mucus  or  foreign  bodies  (the  mouth  being 
rarely  open),  after  a  simple  or  repeated  spasm-like  inspiration 
(the  glottis  remaining  open)." 

In  "  McKendrick's  Physiology,"  coughing  and  sneezing  are 
described  as  powerful  expirations,  in  which  the  air  is  driven 
through  the  oral  cavity  in  the  first,  and  through  the  nasal 
passages  in  the  second. 


188     Proceedinrjs  of  the  Royal  Society  of  Victoria. 

"Hermann's  Physiology"  contains  the  following : — "The 
expulsion  of  foreign  particles.  Such  explosive  expiration  is 
called  sneezing  when  the  nasal  cavities  are  concerned,  and 
coughing  when  the  irritant  is  in  the  larynx." 

Eacli  is  accompanied  b\-  a  noise  produced  by  the  sudden 
bursting  open  of  a  closed  aperture,  which  in  sneezing  is 
found  by  the  opposition  of  the  velum  palati  to  the  pharyn- 
geal wall,  and  in  coughing  by  the  opposed  vocal  cords. 

In  "  Carpenter's  Physiology  "  it  is  stated  "  the  difference 
between  coughing  and  sneezing  is  this,  that  in  the  latter  the 
communication  between  the  larynx  and  the  mouth  is  partly 
or  entirely  closed,  by  the  drawing  together  of  the  sides  of 
the  velum  palati  over  the  back  of  the  tongue,  so  that  tlie 
blast  of  air  is  directed  more  or  less  completeh'  through  the 
nose  in  such  a  way  as  to  cany  off  any  source  of  irritation 
there.  Of  the  purely  automatic  character  of  the  movement  of 
sneezing  there  can  be  no  cpiestion,  since  it  cannot  be 
imitated  voluntaril}-." 

In  "Kirk's  Handbook  of  Physiology  "  we  find  "the  same 
remarks  that  apply  to  coughing  are  exactly  applicable  to  the 
act  of  sneezing,  but  in  this  instance  the  blast  of  air  escaping 
from  the  lungs  is  dii-ected  by  an  instinctive  contraction  ot  the 
pillars  of  the  fauces,  and  descent  of  the  soft  palate,  chiefly 
through  the  nose,  and  any  offending  matter  is  expelled." 

In  "Huxley's  Elementary  Physiology  "  it  is  stated  "in 
sneezing,  the  cavity  of  the  mouth  is  described  as  being  shut 
off  from  the  iihiuynx  by  the  approximation  of  the  soft  palate 
and  the  base  of  the  tongue,  the  air  l)eing  forced  through  the 
nasal  passages." 

All  these  writers,  then,  are  agreed  in  describing  sneezing 
as  a  modified  respiratory  act.  in  which  air  is  blown  through 
the  nose,  and  most  of  them  assume  that  it  consequently 
serves  the  })urpose  of  driving  irritating  substances  from  the 
nose. 

On  the  other  hand,  in  one  of  the  most  recent  works  on 
the  diseases  of  the  nose  (Greville  MacDonald,  published 
1892),  one  finds  the  following  reference  to  sneezing: — 
"Again,  it  may  be  doubted  whether  the  physiological 
reflexes  can  be  considered  in  an}'  way  beneficial.  Sneezing, 
it  may  be  argued,  is  not  of  any  use  in  driving  irritating 
particles  from  the  nose,  seeing  that  it  consists  essentially 
in  a  closing  of  the  palate  during  spasmodic  expiration,  and 
thus  prevents  the  current  of  air  from  passing  through  the 
nose.       But  we  probably  find  the  most  accurate  explanation 


Sneezing:     Fallai-iou>i  Observations.  189 

of  the  phenomenon  in  the  following  considerations  : — On  the 
entrance  of  an  irritating  particle  into  the  nose,  the  [jriniary 
object  of  the  reflex  phenemonon  is  to  increase  the  flow  of 
mucus,  not  only  for  the  sake  of  interposing  some  non- 
irritating  substance  between  the  sensitive  membrane  and  the 
foreign  pai-ticle,  but  even  more  for  the  purpose  of  washing  it 
away.  This  increased  flow  is  produced  by  a  double 
mechanism.  In  the  first  place  there  is  a  supply  of  more 
blood,  and  the  stimulation  of  the  secreting  cells,  through 
nerve  influence  ;  and  in  the  second,  there  is  an  increase  of 
vascular  pressure  from  over-fllling  of  the  venous  sinuses,  as 
described  in  Chapter  I.  Now,  this  pressure  on  tiie  venous 
sinuses  must  be  enormously  increased  by  the  convulsive 
respiratory  act  comprised  in  sneezing.  This  latter  consists 
in  a  violent  contraction  of  the  diaphragm,  &c.,  together  with 
the  closing  of  the  glottis  and  the  post-nasal  space,  by 
contraction  of  the  velum  and  the  superior  strictors  and  of 
tlie  buccal  orifice  by  the  approximation  of  the  tongue 
firmly  to  the  teeth  and  hai'd  palate  ;  in  fact,  every  possible 
movement  is  thrown  into  action  to  prevent  the  exit  of  air 
from  the  larynx,  mouth,  and  nose.  What  is  the  immediate 
consequence  of  this  ?  Increase  of  the  intra-thoracic  pressure, 
which  necessarily  increases  the  intra-vascular  tension, 
especially  in  the  veins,  and  hence  in  the  venous  sinuses  of 
the  nose.  The  act  of  forcible  expiration,  with  all  the  outlets 
from  the  thorax  closed,  if  voluntarily  induced,  i.e.,  without 
tlie  preliminary  irritation  in  the  nose,  is  scarcely  operative 
in  producing  the  effect  described,  and  it  is  probably  only 
when  the  nerve  stimulation  is  excited  at  the  same  time,  and 
the  gland  cells  are  set  working,  that  this  increase  in  the 
venous  pres.sure  is  of  some  additional  assistance." 

Reviewing  these  conflicting  statements,  we  find  difference 
in  matters  of  fact,  and  necessarily  in  the  inferences  drawn 
from  them.  Of  the  inaccuracy  of  the  description  of  sneezing 
given  in  "  Foster's  Physiology"  and  the  other  works  referred 
to,  there  can  be  no  question.  The  process  seems  to  be 
similar  to  that  followed  in  coughing,  with  the  following 
amongst  other  distinctions: — (1)  That  it  is  entirely  involun- 
taril3^  (2)  That  it  is  caused  mainly  through  stimulation  of 
the  anterior  portion  of  the  nose.  Stimulation  of  the  posterior 
portion  of  the  nose  generally  results  in  coughing.  (3)  That 
the  forced  expiration  is,  if  anything,  more  mai-ked  than  in 
coughing.  (4)  That  the  air  in  persons  with  normal  palate 
(and    apart   from   voluntarj^  efforts   modifying   the   act)   is 


190      ProceediiKjs  of  the  Royal  Society  of  Victoria. 

driven  entirely  through  the  mouth  ;  that  is  to  say,  that  the 
palate  is  probably  pressed  firmly  back  against  the  pharynx 
so  as  to  completely  cut  oft  communication  with  the  nose. 
The  peculiar  noise  made  in  sneezing  is  probably  pi'oduced  by 
the  impact  of  the  imprisoned  air  on  the  back  of  the  hard 
palate,  combined  with  certain  modification  of  the  shape  of 
the  mouth  produced  by  movements  of  the  tongue  and  lips. 
In  coughing,  on  the  other  hand,  it  would  seem  that  the  com- 
munication between  the  nose  and  throat  is  not  necessarily 
cut  off,  and  that  the  air  sometimes  passes  through  the  nose 
as  well  as  the  mouth,  and  that  special  movements  of  the  lips 
and  tongue  are  certainly"  different,  if  not  absent  altogether. 
The  mouth  is  generally  opened  more  widely  in  coughing,  and 
the  noise  produced  by  a  cough  is  very  different  from  that 
produced  in  sneezing.  The  one  is  lar^'ngeal  in  the  main,  the 
other  is  chiefly  buccal. 

It  is  possible  that  the  glottis  has  nothing  to  <lo  with 
sneezing,  and  that  the  ob.struction  is  entirely  pharyngeal. 
If,  however,  there  is  a  closed  glottis,  it  is  probable  that  the 
mode  in  which  it  is  opened  in  the  two  cases  is  somewhat 
different.  Coughing  has,  at  all  events,  sometimes  a  definite 
object  to  serve.  It  serves  for  the  removal  of  irritating 
particles  from  the  air  passages,  and  it  is  quite  likely  that  the 
glottis  may  be  differently  disposed  in  sneezing.  Hence  the 
absence  of  glottic  noise  in  sneezing.  The  statement  that  the 
blast  of  air  in  sneezing  is  driven  through  the  nose  has 
originated,  I  think,  in  the  following  manner: — The  observa- 
tions have  been  necessarily  almost  entirely  personal,  and  as 
usual  the  introspective  method,  if  the  term  can  be  used  in 
this  sense,  has  again  proved  fallacious.  When  people  sneeze, 
they  feel  first  a  profound  irritation  in  the  anterior  part  of 
the  nose.  If  this  persists,  there  follow  .some  long  and  deep 
inspirations,  then  a  violent  expiratory  effort  with  po.ssible 
closure  of  the  glottis  or  some  part  of  the  pharynx ;  the 
obstruction  is  suddenly  overcome,  and  the  air  expelled 
through  the  mouth  with  the  characteristic  noise.  Usually 
there  follows  almost  immediately  a  gush  of  watery  fluid 
from  the  nose,  which  is  evidence  of  increased  secretion. 

Now,  putting  these  facts  together,  those  who  first  described 
the  process  of  sneezing,  confused  as  usual  inference  and  fact. 
They  knew  that  coughing,  at  all  events,  served  the  one 
purpose  of  removing  foreign  bodies  from  the  air  passages. 
They  inferred  justly  or  unjustly  that  sneezing  was  adapted 
to  remove  foreign  bodies  from  the  anterior  portion  of  tlie 


Sneezing:    Fallacious  Observations.  191 

nose  by  means  of  the  blast  of  air.  They  felt  the  irritation 
of  the  nose,  and  found  that  sneezing  was  usually  followed 
by  relief  Without  examining  carefully  the  act  of  sneezing, 
to  see  whether  the  air  did  or  did  not  go  through  the  nose, 
they  assumed  that  it  did,  hence  the  description.  It  is  of 
course  possible  that,  in  some  cases  where  observation  was 
made,  abnormal  conditions  of  the  palate  may  have  permitted 
portions  of  the  air  to  get  to  the  nose.  As  the  act  of  sneezing 
is  involuntary,  while  that  of  coughing  is  not,  it  is  impossible 
to  study  the  phenomena  of  the  former,  except  in  an  im- 
promptu and  largely  subjective  manner.  The  vocal  cords 
can  be  examined  with  the  laryngoscope  in  coughing,  but  not 
in  sneezing.  Objective  examination  in  sneezing  is  very 
limited,  by  reason  of  the  nature  of  the  act.  It  seems  to  me, 
however,  perfectly  clear  that  we  have  another  example  of  the 
manner  in  which  hypothesis  has  biassed  observers.  They 
have  unconsciously  endeavoured  to  make  the  facts  fit  the 
theor}'.  An  observation  once  made  and  stated  by  a  com- 
petent authority  has  probably  been  copied  from  one  work 
into  another,  until  of  late  years  the  great  importance  given 
to  phj^siological  respiratory  reflexes  by  physicians  has  caused 
the  matter  to  be  more  closely  investigated. 

Greville  MacDonald's  ingenious  theory  of  the  value  of 
sneezing,  physiologicall}',  may  or  may  not  be  accurate. 
The  fact,  however,  that  patients  suffering  from  eye  disease 
frequently  sneeze  when  exposed  to  a  strong  light,  indicates 
the  necessity  for  caution  before  assuming  that  sneezing  has 
any  value  whatever.  It  may  have  as  little  to  do  with 
normal  physiological  function  in  the  human  being  as  appa- 
rently has  the  patellar  reflex,  the  cremasteric  reflex,  or 
some  other  of  the  general  reflexes.  If  sneezing  is  essential 
to  the  removal  of  a  foreign  body  from  the  anterior  portion 
of  the  nose,  it  is  very  difficult  to  understand  why  coughing 
or  blowing  through  the  nose  would  not  be  equally  serviceable. 
As  Greville  MacDonald  justly  ob.serves,  "it  is  quite  certain 
that  sneezing  alone  cannot  produce  the  rush  of  fluid  from 
the.nose.  It  requires  a  local  determining  agent.  At  present^ 
it  seems  to  me  the  only  conclusion  that  can  be  safely  arrived 
at,  is  the  Agnostic  one."  Greville  MacDonald's  explanation 
is  plausible,  and  has  the  merit,  as  far  as  I  know,  of  standing 
alone. 

How  much  more  fallacious  observation  of  a  similar 
character  exists  in  all  departments  of  science,  it  is  impossible 
to   conjecture,    but   I    think   it   fairly  certain   that,   if  the 


102      Proceed iii(/s  of  the  Royal  Socieij/  of  Victoria. 

treatment  of  diseases  of  the  nose  had  not  become  organised 
into  a  special  de})avtment  of  medicine,  it  would  have  been 
assumed  that  the  significance  of  the  respiratory  reflex  was 
fully  understood.  My  object  in  drawing  attention  to  tin- 
matter  is — (1)  To  put  the  facts,  as  far  as  possible,  befoie 
members.  (2)  To  stimulate  observation,  which  from  the 
necessity  of  the  case  must  be  largely  ])ersonal.  (3)  To  give 
another  example  of  the  manner  in  which  good  observers  are 
biassed  by  the  teleological  assumption.  (4)  Of  the  manner  in 
which  such  fallacious  observations  lead  men  to  accept  explana- 
tions which  wrongly  coloui"  the  work  of  tho.se  who  have  to 
apply  them,  in  practical  life.  An  accurate  statement  of  facts 
with  regard  to  sneezing,  would  probably  have  stimulated 
inquiry  into  the  relation  between  nasal  disease  and  asthma, 
at  a  much  earlier  date  than  1871,  when  attention  was  first 
drawn  to  the  matter  by  Voltolini. 


Art.  XVI. — Physical    GonHtanU   of  Thallium. 

(With  Plate  XVII.) 

By  W.  HuEY  Steele,  M.A. 

[Read  November  10,  1892.] 

Being  in  possession  of  a  piece  of  thallium,  and  being 
unable  to  find  its  constants  in  the  ordinary  books  of  reference, 
[  determined  a  few  of  tliem  as  follows.  The  investigation 
was  conducted  in  the  Physical  Laboratory  of  the  Universit}^ 
of  Melbourne. 

(1)   Coefficient  of  Expansion. 

A  piece  of  the  thallium  was  drawn  into  a  wire  about 
foui'teen  inches  long,  the  ends  cut  off  square,  and  a  nick 
made  near  each  end.  It  was  put  into  a  glass  tube  through 
which  steam  could  be  passed  at  will  from  a  small  boiler. 
The  ends  of  the  tube  being  firmly  clamped,  micrometer 
microscopes  were  focussed  on  the  ends  of  the  wire.  These 
instruments,  supplied  by  the  Cambridge  Scientific  Instru- 
ment Co.,  read  to  xo^oo  ^^^^^^-  I'l^e  positions  of  the  ends  of 
the  wire  and  of  the  outer  and  inner  edges  of  each  nick  were 
observed,  the  observations  being  repeated  several  times. 
The  temperature  of  the  thallium  was  assumed  to  be  that 
indicated  by  a  thermometer  left  lying  beside  the  glass  tube 
all  night,  16-8°  C.  Steam  was  then  passed  along  the  tube 
till  it  was  fairly  dry,  and  after  about  fifteen  minutes,  the 
observations  of  the  positions  of  the  nicks  and  ends  of  the 
wire  were  repeated,  the  temperature  being  assumed  to  be 
1 00°  C.  The  gain  in  length  of  the  whole  wire  was  observed 
to  be  •02()1  inch,  between  the  outer  edges  of  the  nicks 
•()25o  inch,  and  between  inner  edges  •0255  inch.  On  re- 
placing the  glass  tube  by  a  scale,  the  length  of  the  wire  was 
found  to  be  13"83  inch,  and  the  distance  between  the  nicks 
13'69  inch.  Dividing  the  increase  in  length  by  the  rise  in 
temperature  (83-2°),  and  by  the  length  measured,  the  co- 
efficients come  out  0000227,  -0000224,  0000224,  giving  as 
mean  result  -0000225. 

o 


194     Proceedings  of  the  Royal  ^Society  of  Victoria. 

(2)  Specific  Resistance. 
I  at  first  made  several  determinations  of  the  resistance  of 
the  thallium,  in  the  form  of  short  thick  wires,  and  compared 
its  resistance  with  silvei-,  and  atterwards  with  lead,  and 
separately  determined  the  resistances  of  the  specimens  used 
in  comparison.  I  found  it  much  more  accurate  howevei' 
to  draw  the  thallium  into  a  finer  wire,  and  determine  its 
resistance  directly.  This  was  done  with  a  resistance  box, 
with  a  shunt  on  the  variable  arm.  It  was  measured  several 
times  at  slightly  difl:erent  temperatures,  as  sliown  in  the 
following  table  :  — 


t 

r 

•V 

R' 

R 

21 

•16 

5-80 

•1557 

•1551 

21-8 

•16 

6-55 

•1562 

•1.5.'51 

221 

•16 

C^87 

•1564 

•1551 

221 

•17 

1-95 

•1564 

•1551 

t  is  the  temperature  centigrade,  r  and  s  are  the  two 
resistances  in  parallel  which  balance  the  i-esistance  of  the 
thallium.  R^  is  -'^*^,  the  observed  resistance  of  the  thallium, 
and  R  is  the  resistance  at  20°  C,  reduced  by  the  coefiicient 
•0039  {vid.  inf.)  The  length  of  the  thallium  was  4614  cm., 
and  the  mean  value  of  diameter  measured  at  different  parts 
along  it  was  '0874  cm.,  the  mean  error  in  measuring  it  being 
•0005  cm.     The    specific   resistance   at  20°  C.  is   therefore 

TT  X  ^0437^  X    -1551  X  10»    _     20170 
46-14 

(3)  Variation  of  Resistanck  with  Temperature. 
To  determine  this,  the  thallium  was  made  into  a  small 
coil,  and  immersed  in  a  large  beaker  of  water,  with  a  ther- 
mometer about  the  middle  of  tlie  coil.  The  thallium  was 
connected  with  the  terminals  of  a  slide  metre  bridge  by 
means  of  two  stout  pieces  of  copper,  to  which  it  was  firmly 
bound,  and  whose  resistance  was  found  to  be  about  yAtt  of 
that  of  the  thallium.  The  resistance  of  the  thallium  was 
balanced  with  an  approximately  equal  resistance  of  German 
silver,  which  was  taken  as  an  arbitrary  unit  to  measure  the 
resistance  of  the  thallium  at  dift'erent  temperatures.  By 
means  of  a  verj''  sensitive  galvanometer,  the  slider  could  be 
adjusted  to  "1  mm.,  while  the  whole  change  in  position  for 
a  rise  of  80°  C.  was  about  70  mm.     In   reducing  the  bridge 


Physical  (Jondants  of  ThaMluvi. 


195 


readings  to  resist  ances,  correction  was  made  for  the  fact  that 
the  middle  point  of  the  wire  was  not  the  electrical  centre, 
which  was  JJ-.S  mm.  to  one  side,  and  the  resistances  were 
dirainislied  by  1  j)er  cent,  of  the  cold  resistance  of  the 
thallium  on  account  of  the  copper  connections.  Two  inde- 
pendent sets  of  observations  were  made,  and  from  each  the 
coefficient  at  20°  0.  was  calculated  by  the  method  of  least 
squares.  The  figures  fi-om  Avhich  the  calculations  were  made 
are  given  in  the  annexed  table  : — 


Sei 

1. 

Set 

2. 

t 

R 

t 

i 

i 

R 

17-8 

•987 

16-2 

i 

1-017 

30-5 

1-035 

32-1 

1-079 

41-4 

1-074 

48-2 

1143 

51-7 

1-102 

61-4 

1-196 

62-1 

1-154 

79-8 

1 

1-274 

80-8 

1-227 

99 

1-357 

98-8 

1-312 

85-1 
72-2 
67-5 

1-298 
1-246 
1-226 

The   values   of  the    coefficient   from   the  above  tables  are 
•00394  and  00400. 

Having  determined  these  values  by  means  of  the  slide 
bridge,  I  proceeded  to  verify  the  result  by  measuring  the 
resistances  with  a  resistance  box,  and  shunt  as  described 
above.  Two  independent  sets  of  observations  were  made  as 
before.  The  observed  values  are  oiven  in  the  followincj 
table  : — 


Set   4. 


f 

R 

t 

R 

181 

-2054 

]7-3 

•2134 

32 

■2162 

59-3 

•2490 

40-6 

-2228 

80-9 

•2679 

53 

-2329 

99-3 

•2812 

60-7 

•2381 

54 

•2456 

72-1 

-2188 

24^9 

•2232 

o  2 


I9G      Proceedings  of  the  Royal  Society  of  Victoria. 

The  values  of  the  coefficients  from  sets  3  and  4  are 
OOS.Sl  and  00391.  The  mean  of  these  four  gives,  as 
the  coefficient  at  20°  C,  -00392.  This  is  larger  than  the 
value  for  most  metals  other  than  iron. 


(4)   Thermo  Ei.ecthk;  Height. 

As  I  had  a  piece  of  pure  silver,  and  no  other  metal  pure, 
I  resolved  to  find  the  thermo  electric  height  of  thallium 
with  regard  to  silver,  and  assume  Professor  Tait's  result  for 
silver  in  order  to  obtain  the  absolute  value  for  thallium. 
Having  done  so,  it  was  found  that  the  thallium  line  thus 
determined,  crossed  Professor  Tait's  copper  line  at  about 
70°  C,  and  that  copper  was  therefore  an  exceptionally 
favourable  metal  with  which  to  compare  thallium.  I  there- 
fore obtained  ])ure  copper  and  compared  thallium  with  it, 
and  found  that  thallium  was  further  below  copper  than 
below  silver;  and  on  finall}"  fiyi'^g  coppei-  and  silver,  I  found 
the  lines  should  be  very  much  closer  together  than  the_y  are 
in  Professor  Tait's  diagram,  and  that  copper  should  be  above 
.silver  and  not  below  it.  I  therefore  purified  some  lead,  and 
constructed  a  diagram  of  my  own  ibr  the  four  metals — lead, 
thallium,  copper  and  silver.  To  obtain  pure  lead,  I  dissolved 
some  sheet  lead  in  nitric  acid,  and  precipitated  it  as  sulphate 
by  adding  dilute  sul])huric  acid.  The  sulphate  thus  obtained 
was  lieated  with  carbonate  of  soda  and  cream  of  tartar  in  a 
Hessian  crucible  in  an  injector  furnace,  and  lead  obtained 
which  was  assumed  pure,  tliough  it  contained  a  trace  of 
potassium.  I  used  an  astatic  low  resistance  galvanometer 
with  a  lamp  and  scale,  at  a  distance  of  about  four  feet,  the 
.scale  divisions  being  fortieths  of  an  inch.  The  resistance  of 
the  galvanometer  was  somewhat  less  than  an  ohm,  but  with 
the  leads  and  the  wires  of  the  thermo  electric  circuit,  .the 
resistance  was  a  little  over  an  ohm.  So  low  an  E.M.F.  as 
•00000 1  volt  or  U)0  absolute  units  gave  a  deflection  of  one 
scale  division.  This  appears  to  be  about  30  times  as 
sensitive  as  the  one  used  by  Professor  Tait  twenty  years 
ago.  To  determine  the  exact  value  of  a  scale  division,  the 
galvanometer  was  joined  in  series  with  an  ordinary  Daniell 
cell  and  various  high  resistances,  smd  immediately  after  or 
before  its  E.M.F.  compared  with  a  Latimer  Clark  cell,  by 
means   of    a    condenser    and    balliatic   galvanometer.      In 


Physical  Constants  of  Thallium. 


197 


examining  the  thermo  electric  power  of  two  metals,  I  twisted 
together  their  ends  and  coiled  the  joint  round  the  bulb  of  a 
thermometer,  immersing  the  whole  in  a  bath  of  olive  oil. 
The  other  junction  was  kept  in  a  large  beaker  of  cold  water 
with  a  thermometer  in  it,  which  was  observed  from  time  to 
time,  and  if  necessary,  correction  made  for  the  rise  of 
temperature.  Tliis  rise  was  never  more  than  a  degree,  the 
corresponding  correction  being  one  or  two  scale  divisions. 
The  relation  between  the  observed  values  of  temperature  and 
galvanometer  reading  is  parabolic,  and  if  we  express  the 
excess  of  the  temperature  of  the  hot  junction  over  the  cold 
by  t,  and  the  number  of  scale  divisions  by  s,  then  s  z=  a  t  -\- 
h  t^  is  the  connection  between  .s  and  t,  where  a  and  b  are 
constants  to  be  determined  preferably  by  the  method  of  least 
squares,  as  was  done  with  each  set  of  observations,  though 
the  operation  is  rather  laborious.  The  following  table  may 
be  taken  as  typical  of  the  accuracy  attained  : — 


T 

s 

•s 

(ob^-erved) 

[calcjilated] 

19 

0 

0 

76 

111 

119 

73 

107 

il2 

70 

102 

108 

76 

115 

119 

131-7 

206 

201 

ISO 

202 

199 

180-3 

219 

249 

200 

259 

260 

184 

249 

250 

179 

246 

247 

IGl 

232 

232 

In  this  case,  the  metals  being  thallium  and  lead,  the 
resistance  of  the  circuit  was  1-395  ohm.  The  equatioji  to 
the  parabola,  represented  by  the  first  and  second  columns, 
is  s  =  2-36^  -  00.512  t^  where  t  =  T  ~  19.  A  Daniell 
cell,  when  used  to  charge  a  condenser,  gave  a  throw  of 
276-3  sc.  clivs.;  a  Latimer  Clark  cell  gave  846-4- ;  the  E.M.F. 
of  the    Daniell   is   thus— 1-435  x  276-3  -^  346-4  =  1-144. 


19.S     Proceedinc/s  of  the  Royal  Society  of  Victovifx. 


The  same  Daniell,  when  connected  in  series  with  the  gal- 
vanometer,  gave  a  deflection   g  for  resistance   R.     If  the 


9 

R 

142-y 

12000 

176 

lUOOO 

195 

9000 

246 

7000 

287 

6000 

213 

8000 

current  through  the  galvanometer  be  kg,  then  -^  =  /c  g, 
or  /.•  =  ^^^.  Now  the  mean  value  of  Rg  is  1729000, 
therefore  Z;  =  ivoguoo-  ^^  ^  ^^®  ^^^®  electromotive  force  of  the 
thermal  circuit,  and  i'  its  resistance,  then  e  =  ksr,  or  the 
electromotive  foice  corresponding  to  one  scale  division  is 
^^1729000 '"  '^''^^t'  or  92-3  absolute  units  of  electromotive  force. 

Now  s  =  2S6t  —  0051 2f-,  or  if  we  measure  the  tem- 
perature from  the  neutral  point  of  the  two  metals,  s  = 
•00512/^.  If  the  temperatures  be  the  neutral  point,  and 
100°  above  or  below  it,  and  if  m  be  the  relative  Thomson 
effect,  then  512  x  92-3  =  5000  ?>i,  and  m  =  -94. 

To  find  the  neutral  point—  ^Jf  =  0,  i.e.,  2SQ  -  2  x 
•00512^  =  0,  t  =  230,  i.e.,  249°  C. ;  the  height  at  0'  C.  is 
therefore  249  X  '94  =  234.  The  thermo  electric  height  of 
thallium  above  lead  is  thus—  234  —  94 1,  t  here  being 
temperature  centigrade. 

Another  similar  but  independent  set  of  observations 
gave  as  the  height  198  —  -65  t,  the  mean  of  these  being 
217  — -79  /.  Of  the  four  metals,  each  pair  was  taken 
together,  and  the  following  results  obtained.  In  each  case 
the  higher  metal  is  the  first  : — 

-  217-     -79^ 

-  43  +  1-79^ 

-  252  +  i-oe^; 

-  12  +      10/ 

-  40  +  147 / 

-  200  +  1-35/ 

If  we  add  the  Thallium-Lead  to  the  Copper-Thallium,  we 
get  200  +  100  t,  which  agrees  fliirly  well  with  the  directly 


1. 

Thallium -Lead    - 

2. 
3. 
4. 
5. 

Copper-Thallium 
Copper-Lead 
Copper-Silver 
Silver-Thallium 

G. 

Silver-Lead 

Proc.  R  5.  Victoria  Plate  Ml 


^/- 

//  \y 

/ 

/ 

/         // / 

/       /      / 
/       /       f 

/ 

/ 

/'' 

/ 
/ 
/ 

/ 

/ 

/ 

f 

>< 

/ 

\ 

X 

(0 

\. 

Pb 

Tempe 

^afure 

Fhydcal  Cuiisfants  of  Thallium.  199 

observed  value  of  Copper- Lead.  The  results  with  the  silver 
are  not  very  consistent.  Various  diagrams  can  be  con- 
structed from  the  above  observations,  but  the  nearest  to  the 
true  one  will  ])robably  be  obtained  from  the  first,  third  and 
fourth,  from  which  the  figure  is  drawn  (see  Plate  XVII),  tlie 
dotted  lines  on  the  figure  being  Professor  Tait's  results  for 
silver  and  copper. 

I  have  been  for  some  months  engaged  on  a  series  of 
observations,  of  which  I  hope  to  give  an  account  to  this 
Society  shortly,  from  which  it  appears  that  thermo  electric 
values  cannot  be  absolutely  constant,  and  which  explains  the 
above  inconsistent  results  of  the  observations  on  these 
metals. 

Summary. 

The  constants  obtained  are  thus  : — 

(1)  Coefticient  of  expansion      -      "0000225 

(2)  Specific  resistance       -         -        20200  at  20°  C. 

(3)  Range    of    resistance    with 

temperature        -         -      -00392  at  20°  C. 

(4)  Thermo  electric  height        -       216—  '79^ 

The  thallium  was  obtained  from  Schuchardt,  and  Professor 
Masson  has  kindly  anal^^sed  it  and  supplied  the  following 
statement  of  his  results  :  — 

"  The  small  sample  of  thallium  wire  submitted  to  me, 
was  found  to  contain  as  impurities,  lead,  arsenic  and  copper. 
An  estimation  of  the  lead  showed  it  to  be  present  to  the 
extent  of  1-50  per  cent.  The  arsenic  and  copper  were 
present  in  too  small  amount  to  be  estimated  in  so  small  a 
sample.  A  direct  estimation  of  the  thallium  itself  sliowed 
the  wire  to  contain  9790  per  cent,  of  that  metal." 


Art.  XVII.  —  On  "  Confocal  Quadrics  of  MomenU  of 
Inertia "  'pertainiiig  to  all  Planes  in  Space,  and 
Loci  and  Envelopes  of  Straight  Lines  vjhose 
"  Moments   of   Inci-tia  "   are    Constant. 

By  Martin  Gardiner,  C.E. 

iRead  May  12,  1892.] 

Abstract. 

The  author  commences  by  solving  the  following  problem, 
by  the  Cartesian  co-ordinate  method  : — 

Problem. — Given  any  number  of  points  Pj,  P^,  P^,  .... 
in  space,  and  corresponding  numbers  «!,  a^,  «3,  .  .  .  ,  known 
in  signs  and  magnitudes  as  respective  multipliers  ;  to  hud 
the  Envelope  of  a  plane  L  L  X,  such  that,  in  every  position 
it  can  assume,  we  shall  have 

a^  .p\  +   a^.pl   +   (^3  .  pi  -\-   .  .  .  .   =   8, 

in  which  p\,pl,'pl,  .  .  .  .  ,  represent  the  squares  of  the 
pedals  from  the  points  Pj,  Po,  P3,  .  .  .  ,  to  the  plane  L  L  L, 
and  S  a  constant  entity  known  in  sign  and  magnitude. 

He  finds  the  equatloii  of  the  envelope  of  the  plane  L  L  L 
to  be  that  of  a  Quadric  whose  centre  is  coincident  witii 
the  meaTi-centre  of  the  given  points  for  the  multipliers 
«i,  a^,  a-i,  .  .  .  And  from  the  form  of  the  equation  arrived 
at  (which  is  given  abridged  and  expanded),  he  infers  that 
for  all  possible  values  of  the  entity  S,  the  corresponding 
Quadrics  are  Confocal  Quadrics. 

He  then  shows  by  a  purely  geometrical  method  (indepen- 
dent of  co-ordinates)  that  for  an}^  constant  value  of  *S',  the 


"  Oonfocal  Qiiadrics  of  Moments  of  Inertia. 


201 


envelo|)e  of  the  plane  L  L  L  is  a  Quadric  whose  centre  is 
coincident  with  the  mean  centre  of  the  points  jPj,  P-i,  P^, 

.  .   .  ,  and  their   respective    multipliers   «i,  a2,  a^, 

And  he  shows  that  tlie  quadrics  corresponding  to  all 
possible  values  of  the  entity  ^.S,  are  Confocal  Quadrics. 

In  order  to  amplify  his  Geometrical  Method,  he  proceeds 
to  give  a  full  and  complete  solution  to  the  particular  cases 
in  which  the  given  points  Pj,  Po,  Pg,  .  .  .  ,  are  all  in  one 
straight  line.  And  he  shows  that  it  depends  on  the  state  of 
the  data,  as  to  whether  the  Confocal  Quadrics  be  Ellipsoids  ; 
Hyperboloids  of  One  Sheet ;  Hyperboloids  of  Two  Sheets  ; 
Spheres ;   or  Paraboloids. 

He  then  directs  attention  to  the  Physical  Aspect  of  the 
problem,  which  he  enunciates  as  follows  : — 

Problem. — Given  any  masses  M^,  M2,  i/3,  .  .  .  ,  in  space, 
and  corresponding  units  a-^,a.2,a.i,  ■  ■  .  ,  known  in  signs  as 
their  respective  multipliers;  to  find  the  Envelope  of  a 
plane  L  L  L,  such  that  in  every  position  it  can  assume, 
we  shall  have  the  sum  of  the  Moments  of  Inertia  of  the 
masses  represented  by 

a,  .  ^  m,  (Pi  Lf    +    a.2 .  ^  m.  (P^  Lf   +    a,.^  m, .  (P3  Lf 
■+...   =   a  constant  S, 

in  which  97ii,  liiaj '>''*'3,  •  •  •  represent  molecules  of  the  masses 
i/i.jyo.Jfg,  .  .  .  ,  at  any  points  P^,  P2,  P3,  ...  in  those 
masses,  and  in  which  P^L,  PoZ,  P^L,  .  .  .  represent  the 
pedals  from  the  points  Pj,  Po,  P,,  .  .  .  ,  to  the  plane  LL  L. 

In  elucidation  of  this  aspect  of  the  problem,  he  recon- 
siders the  particular  cases,  in  which  he  now  replaces  the 
given  points  or  molecules  at  Pj,  P2,  P3,  ...  all  in  one 
line,  by  Spheres  whose  centres  are  all  in  one  straight  line. 
He  shows  that  the  results  arrived  at  previously,  apply 
when  masses  replace  mere  molecules  ;  and  that,  according 
to  analogous  states  of  the  data,  the  Confocal  Quadi-ics  will 
be  Ellipsoids,  Hyperboloids,  Spheres,  or  Paraboloids. 

He  establishes  the  limiting  values  for  the  constant  S,  and 
exposes  the  limiting  forms  of  the  Quadrics  in  minute  and 
full  detail.     And  he  corroborates  a  remarkable  theorem  of 


202     Proceedings  oj  the  Royal  Society  of  Victo7ia. 

Duhamel's,  as  to  the  existence  of  two  points,  for  each  of 
luhich  Poinsot's  "  Ellipsoid  of  stress "  is  a  Sphere.  He 
shows,  moreover,  these  two  points  to  belong  to  a  "Focal 
l-'onic"  of  the  family  of  Confocal  Quadrics. 

In  the  case  in  which  the  bodies  are  Spheres  situated  in 
any  manner  in  space,  he  gives  a  simple  and  effective 
metliod  of  finding  the  tliree  principal  axes  of  inertia. 

He  then  records  the  followino-  eioht  Theorems,  as  results 
01  his  investigations  : — 

Theorem  1. 

Given  any  raas.ses  i/j,  M^,  M^,  .  .  .  in  space,  and 
corresponding  numbers  a,,  a.2,  a^,  ...  of  known  signs 
as  multipliers.  If  a  plane  L  L  L  (otherwise  unrestricted) 
be  such  that  in  every  position  it  can  assume,  the  sum  of 
the  moments  of  inertia  of  the  entities  aj.  M^,  a^.  M^,  a^.  M^, 
.  .  .  ,  with  respect  to  it,  be  of  any  constant  magnitude  S, 
then  will  the  envelope  of  the  plane  be  a  determinable 
(piadric  Q,  whose  centre  is  coincident  with  the  mean  centre 
of  the  entities.     And  the  whole  system  of  quadrics  Qi,  Q.2,  Q^, 

corresponding    to    all    values  S,,  S2,  S^,  .  .  .  .  , 

of  S,  will  be  concentric,  coaxial,  and  confocal  quadrics.  And 
in  all  cases  in  which  the  multipliers  ai,  a.^,  .  ■  ■  are  all 
positive,  the  quadrics  will  be  Ellipsoids  and  Hyperboloids  of 
One  Sheet. 


Theorem  2. 

Given  any  masses  i/j,  ill 2,  M^,  ...  in  space,  and 
corresponding  numbers  a^,  a^,  0^3 ,  .  .  .  of  known  signs, 
as  multipliers.  The  envelope  of  all  planes  LLL  passing 
through  any  given  point  V  in  space,  and  such  that  the  sum 
of  the  moments  of  inertia  of  the  entities  a^.  M^,  cio.  M2,  (tj.  i/3, 

,  with  respect  to  them  severally,  is  of  any  constant 

magnitude  S,  will  be  a  determinable  quadric  cone  C, 
which  envelopes  a  determinable  quadric  Q  whose  centre  is 
coincident  with  the  mean  centre  0  of  the  entities.  And  the 
whole  family  of  such  cones  Cj,  G.^,  C3,  .  .  .  ,  corresponding 
to  all  values  8-^,  S2,  S-^,  .  .  .  ,  of  S,  will  be  coaxial  and 
confocal  cones  enveloping  coaxial  and  confocal  quadrics, 
whose  common  centre  is  the  mean  centre  0  of  the  entities 


"  ConfoccU  Quadrics  of  Moments  of  Inertia."      203 

«!.  Ml,  do.  M.2, And  if  the  point  V  be  at  infinity, 

unci  given  in  direction  by  means  of  a  vector  0  R  passing 
thi'ough  the  mean  centre  0  ;  then,  corresponding  to  various 
values  of  S,  the  envelopes  oi  L  L  L  consist  of  a  system  of 
confocal  cylinders  enveloping  the  quadrics,  dnd  having  as 
common  principal  axis  the  directing  vector  0  R. 

Now  Ml,  M.2,  M^,  .  .  .  being  masses,  and  a^,  a^,  a^, 
numbers  known  in  signs:  we  know  that  if  a  plane  ZX  2/ 
l)e  such  that  the  sum  of  the  moments  of  inertia  of  the 
entities  11^.  M^,  cu  M^,  a-^.  Mo^,  .  .  .  ,  with  respect  to  it  is  of 
a  constant  magnitude  S,  then  will  the  envelope  of  the  plane 
be  a  determinable  quadric  Q.  But  the  line  of  intersection 
/  I  of  any  two  mutnalh'  orthogonal  planes,  both  tangent 
to  the  quadric  Q,  is  obviously  such  that  the  sum  of  the 
moments  of  inertia  of  the  entities  with  respect  to  it  is 
represented  by  2..s'. 

We  can  easily  form  the  equations  of  tangent  planes 
to  tile  quadric  Q,  and  express  their  mutual  orthogonism  ; 
but  we  need  not  try  to  evolve  an  equation  of  a  surface 
which  could  be  the  envelope  of  all  the  lines  1 1  of  intersection 
of  the  pairs  of  mutuall}'  orthogonal  tangent  planes  to  Q. 
This  is  obvious : — for  if  we  suppose  j:)  to  be  any  point 
whatever  on  any  surface,  and  construct  a  Poinsot  Elli))soid 
having  such  point  as  centre,  we  perceive  that  the  lines  / 1 
tlirough  the  point  form  a  cone,  and  cannot  generally  ell  be 
tangents  at  one  point  to  any  other  surface.  However,  we 
j)roceed  to  find  the  Loci  and  Envelo})es  of  lines  ly  l^  which 
fulfil  the  conditions  as  to  equality  of  moments  of  inertia, 
and  respecting  which  other  conditions  are  imposed. 

1°. — With  respect  to  all  the  lines  li  iy  wliich  are  parallel 
to  any  fixed  straight  line  R  R  passing  through  the  mean 
centre  0,  which  is  also  the  centre  of  the  quadric  Qj. 

If  throLigh  0  we  draw  a  plane  normal  to  the  line  R  R, 
and  that  we  put  c^  c^  Cy  to  represent  the  conic  which  con- 
.-stitutes  its  trace  on  the  quadric  Qi  :  then,  from  a  well-known 
theorem,  we  perceive  that  the  pairs  of  mutually  orthogonal 
tangent  planes  whose  points  of  contact  lie  in  the  conic 
''iCiC,,give  us  all  the  lines  lilx  parallel  to  the  fixed  line 
RR,  and  that  they  constitute  a  Right  Circular  Cylinder 
liavinw  R  R  as  central  axis. 


204      Proceedings  of  the  Royal  ISociety  of  Victoria. 

2"". — With  respect  to  all  the  lines  /j  /j  situated  in  tangent 
planes  to  the  quadric  Qi . 

We  may  first  observe  that  if  Pj  Pj  1\  be  any  fixed  plane 
tangent  to  the  quadric  Q^,  and  that  we  project  the  quadric 
itself  orthogonally  by  means  of  othe)-  tangent  planes  upon 
J\  Pi  Pi,  then  will  the  ])rojection  be  a  conic  c^  q  c^  situated 
in  the  plane  PiP^P^,  which  is  obviousl}^  the  envelope  of 
all  the  lines  li  /^  in  the  plane. 

3°. —  With  respect  to  all  tlie  lines  l^  li  situated  in  any 
plane  B  BB  whatever. 

We  first  proceed  and  find  the  sum  s,^  of  the  moments  of 
inertia  of  the  entities  a^.i/i,  a.,.  M2,  a.^.  M^,,  .  .  .  ,  with 
respect  to  the  plane  BBS.  We  then  find  the  quadric  Q^ 
such  that  the  sum  of  the  moments  of  inertia  of  the  entities 
with  respect  to  any  of  its  tangent  planes  is  =  2.Si, — s^ .  Then, 
ol)viously,  the  orthogonal  projection  of  the  quadric  Q^  so 
found  (by  means  of  tangent  [)lanes  to  it)  upon  the  plane 
B BB  will  be  a  conic,  which  is  the  envelope  of  the  lines  l^  l^ 
situated  in  the  plane. 


The  foliowins^  is  an  obvious  deduction  : — 


Theorem  .'>. 

Given  any  masses  M^,  M2,  M-^,  ...  in  space,  and  corres- 
ponding numbers  a^,  a.^,  «3,  .  .  .  of  known  signs  as  multi- 
pliers ;  and  given  also  the  system  of  confocal  quadrics 
Qi,  Q'^'  Qz>  •  •  •  ■>  such  that  the  sum  of  the  moments  of 
inertia  of  the  entities  a^.  ili,,  a.-M.^,  a-^.  31-^,  .  .  .  ,  with 
respect  to  tangent  planes  to  the  quadrics  are  equals 
respectively  to  6'i,  s.,,  .^.g,  .  .  .  ;  then  the  orthogonal  pro- 
jections of  the  quadrics  on  any  given  plane  B  B  B  in  space, 
constitute  a  family  of  confocal  conies,  which  are  the 
respective  envelopes  of  straight  lines  l^li,  l.^l-i,  Izh,  •  .  •  ■  , 
situated  in  the  plane,  such  that  the  sum  of  the  moments  of 
inertia  of  the  entities  ftj.  i/j,  a._,.  M-2>  (-h- ^Z'  •  •  •  >  with 
]-espect  to  them,  are  determinable  constants.  And  if  the 
plane  P^  B^  B  be  parallel  to  either  one  of  the  two  systems  of 
parallel  circular  sections  of  the  conibcal  quadrics,  then  will 


"  Confocal  Quadrics  of  Moments  of  Inertia^''      2().> 

the  projections  of  the  ([uadiMcs  on  the  pLme  be  ;i  system  of 
concentric  circles. 

Note. — The  differences  of  the  moments  of  inertia  with 
respect  to  the  lines  ^j/j,  /., /o-  ^;5^3>  •  •  •  .  (tangents  to  the 
respective  conies)  on  the  plane  BBB  are  obvionsly  equals  to 
the  differences  of  the  moments  of  inertia  with  respect  to 
tangent  planes  to  the  qnadrics  Qi,  Q2,  Qs,  ■  •  ■ 

If  we  draw  planes  Pi  Pi  Pi,  P2P2P2)  ■  ■  ■  ,  tiirough  any 
diameter  D  D  of  any  one  Q  of  the  family  of  Confocal  quadrics. 
the  lines  / 1  situated  in  these  ])lanes  and  sucli  that  the  sum 
of  the  moments  of  inertia  of  the  entities  %.  J/i,  «o.  M.y,  '^3.  il/;j, 
....  with  respect  to  them,  several!}'-,  is  of  any  constant 
magnitude  2.s,  have  (as  alrendy  observed)  as  envelopes,  in 
the  ]ilanes,  determinable  conies.  And  we  know  that  those 
of  the  lines  II  which  are  parallel  to  D D  form  a  circular 
cylinder;  having  the  line  D  B  as  axis.  But  it  is  easy  to 
])erceive  that  it  is  onl}'  when  the  axis  D  D  is  normal  to  one 
of  the  circular  sections  of  the  quadric  Q  that  the  conies  cut 
D  D  in  the  one  and  same  point,  at  which  the  lines  1 1  form  a 
tangent  plane  to  all  the  conies.     Hence : — 

Theorem  4. 

Given  any  number  of  masses  i/i,  J/o' -^^3'  •  •  •  >  in  space, 
and  corresponding  nundjers  a^,  cio,  a^,  .  .  .  ,  of  known  signs 
as  multipliers  ;  if  a  straight  line  1 1  move  in  space  so  as  to  be 
always  in  contact  with  the  line  D  i)  of  a  diameter  of  any 
quadric  Q  (of  the  confocal  family)  normal  to  either  system  of 
its  circular  sections,  and  so  that  in  every  position  the  sum 
of  the  moments  of  inertia  of  the  entities  aj.  il/j,  «o.  i/o,  .  .  .  , 
with  respect  to  it,  is  of  any  constant  magnitude  2.s ;  then 
will  the  envelope  of  the  straight  line  Z  ^  be  a  determinable 
quadric  lu  of  revolution,  having  the  mean  centre  0  as  centre, 
and  the  fixed  line  B  B  as  axis.  And  all  such  quadrics 
'"1,  10-2,  iv-i,  .  .  .  ,  corresponding  to  all  possible  values 
2.S1,  2.S.2,  2.S3,  .  .  .  ,  of  the  constant  are  determinable 
quadrics  of  revolution,  having  the  mean  centre  0  as  common 
centre,  and  the  line  Z)  Z>  as  principal  axis. 

Theorem  5. 

The  Locus  of  a  straight  line  1 1  through  any  fixed  point  B' 
in  a  line  B  B  through  the  mean  centre  0  and  normal  to 


200      P roceedings  of  ike  Roydl  Hociely  of  Victoria. 

<!ircnlar  sections  of  the  confocal  (|uadiics  Qi,  Q2,  Qs,  ■  ■  •  , 
and  such  tliat  the  sum  of  the  moments  of  inertia  of  the 
entities  ((i.il/j,  a^.  Mo,  .  .  .  ,  with  respect  to  it,  is  of 
constant  magnitude  2.6',  is  a.  (juadric  cone  of  revolution. 
Jiaving  the  point  D^  as  vertex,  and  IJ  D  as  axis. 

We  know  that  the  locits  of  the  lines  1 1  of  intersection  of 
all  pairs  oi'  mutually  orthogonal  tangent  planes  to  any 
quadric,  cone  G  is  another  ([uadric,  cone  E  concyclic  with 
the  reciprocal  of  the  cone  C.  (8ee  Salmon's  "Geometry  of 
Three  Din:iensions,"  Art.  247).  And  if  C  be  a  cone,  such 
that  the  sum  of  the  moments  of  inertia  of  the  entities 
ttj.iVi,  «o.  J/o,  .  .  .  ,  with  respect  to  its  tangent  planes, 
severally,  be  equal  to  a  constant  .s,  we  know  that  the  sum 
of  the  moments  of  inertia  of  the  entities  with  respect  to  the 
lines  1 1,  severally,  must  be  equal  to  2.«.     Hence  we  have  : — 


Theorem  6. 

Given  any  masses  Mj,  Mo,  Mg,  .  .  .  ,  in  s{)ace  and 
corresponding  numbers  rtj,  a.,,  «;,,  .  .  .  ,  of  known  signs,  as 
multiplierh  ;  the  Locus  of  a  straight  line  II  passing  through 
any  given  point  Fin  space,  and  such  that  the  sum  of  the 
moments  of  inertia  of  the  entities  rq.  M^,  a-j-  Mo,  a-^.  M.^,  .... 
with  respect  to  it  — •  any  constant  2..s,  is  a  quadric  cone  JE, 
having  the  point  V  as  ver-tex,  and  concyclic  with  the 
reciprocal  of  the  cone  G,  having  V  as  veitex,  and  such  that 
the  sum  of  the  moments  of  inertia  of  the  entities  with 
respect  to  its  tangent  planes  =  s,  &c. 


Theorem  7. 

If  three  |)lanes,  always  mutually  oi'thogonal,  move  in 
space  so  as  to  continue  to  be  tangent  ])lanes  respectively  to 
any  three  of  the  confocal  quadrics  Q^,  Qo,  Q:i',  then  will  the 
Locus  of  their  common  point  of  intersection  be  a  Spltere,  whose 
centre  is  coincident  with  the  mean  centre  0  of  the  entities 
cij.  i/i,  ao.  Mo,  .  .  .  ,  which  is  also  the  centre  of  the  quadrics. 

Note. — This  Theorem,  which  is  an  obvious  deduction 
from  the  kinetic  properties  exposed,  was  arrived  at  by 
Salmon  by  means  of  a  formula  due  to  Chasles.  (See 
Sahnon's  "Geometry  of  Tlnee  Dimensions,"  Art.  172.) 


"  Confocal  Quadrics  of  Moments  of  Inertia."      207 

Theorem  8. 

If  two  planes  A  and  B  mutually  orthogonal,  be 
tangent  planes  respectively  to  any  two  qiiadrics  Q^,  Q2,  <»t' 
the  confocal  family  ;  tlien  will  the  other  pair  of  tangent 
planes  A^  and  B^  through  their  line  of  intersection  II,  to  the 
same  two  quadrics,  l)e  iinitually  orthogonal. 

This  is  an  obvious  deduction  from  the  kinetic  properties 
exposed. — The  planes  A  and  B  being  tangents  to  the  quadrics 
Qi  and  Q2,  the  moments  of  inertia  of  the  entities  d^.  i/j,  a^. 
M2,  ■  •  •  ,  with  respect  to  them  ai'e  constants  6?!  and  So ;  and 
the  sura  Sj  +  Sj  *^f  these  moments  of  inertia  is  equal  to  the 
moment  of  inertia  of  the  entities  with  respect  to  their  line 
of  intersection  1 1.  And  since  the  moment  of  inertia  with 
respect  to  the  line  II  m  equal  to  the  sum  of  the  moments  of 
inertia  with  respect  to  the  tangent  planes  A^  and  B^,  it 
follows  that  A^  and  B^  must  be  mutually  orthogonal. 

This  theorem  is  an  e.x:tension  to  confocal  quadrics  of  one 
pertaining  to  confocal  conies,  due  to  Admiral  De  Jonquieres 
of  the  French  Navy,  who  is  one  of  the  most  distinguished 
geometers  in  Europe.  (See  "Melanges  de  Geomdtrie  Pure," 
par  E.  De  Jonquieres.) 

Observations. 

The  family  of  confocal  quadrics  Qi,  Q2,  Q^,  •  •  •  ,  and 
the  properties  of  inertia  pertaining  to  them,  are  worth}' 
of  attention,  not  only  on  account  of  their  intimate  con- 
nection with  "  Wave  Surfaces,"  and  "  Surfaces  of  Elasticity," 
but  also  on  account  of  their  direct  applications  to  many 
important  problems.  (See  Salmon's  "  Geometry  of  Three 
Dimensions,"  Arts.  467,  480,  &c.) 

2°. — Some  interesting  properties  pertaining  to  confocal 
quadrics  can  be  deduced  by  application  of  the  numerous 
new  theorems  arrived  at  by  the  author,  and  published  in 
Vol.  X  of  the  "  Quarterly  Journal  of  Pure  and  Applied 
Mathematics,"  under  the  title — "  Properties  of  Quadrics 
having  Common  Intersection,  and  of  Quadrics  inscribed  in 
the  same  Developable." 

S". — Since  writing  the  present  paper,  the  author  has 
found  that  the  question  had  been  previously  considered  by 
the   late    Professor   Townsend,   of  the    Dublin    University. 


208     Proceedhi(]s  of  tite   lioiial  JSocieti/  of  Victoria. 

The  results  at  whicli  he  arrived  are  given  ivithout  any 
investigations  on  page  .'J  12  of  Williamson's  "Integral 
Calculus."  From  question  19,  as  there  enunciated,  it  would 
appear  that  Townsend  did  not  perceive  that  the  envelope  of 
the  plane  is  an  ellipsoid  ouly  when  the  prescribed  moment 
of  inertia  is  not  less  than  a  certain  determinable  magnitude  ; 
or  that  it  is  a  Hyperboloid  of  One  Sheet  for  all  values  less 
than  such  limiting  value.  Nor  does  it  a])pear  tliat  he 
considered  the  case  in  which  the  envelope  of  the  plane  is 
a  Hyperboloid  of  Two  Sheets,  or  any  limiting  values  of  the 
moment  of  inertia. 


Art.  XVIII. —  iVotes  on  a  Poisonous  Species  of  Homeria 
(H.  collina,  Vent. — var.  miniata),  found  at  Faseoe 
Vale,  causing  deaih  in  cattle  and  other  animals 
feediny  upon  it. 

By  D.  McAlptnk  and  P.  W.  Farmer,  M.B.,  Ch.  B. 

[Eead  November  4,  1892.] 


Introductory. 

The  sudden  death  of  a  nnmber  of  cattle  at  Pascoe  Vale, 
a  suburb  of  Melboui-ne,  about  the  middle  of  Septeinbei-, 
attracted  a  good  deal  of  attention,  and  from  various 
accompanying  circumstances,  there  were  grounds  for  believing 
that  the  herbage  had  .something  to  do  with  it.  Specimens 
of  the  supposed  poisonous  weed  were  sent  to  Baron  von 
Mueller,  Government  Botanist,  and  he  determined  it  to  be 
a  species  of  Homeria,  a  native  of  South  Africa.  He 
remarked  that  "in  their  native  country  occasionally  pasture 
animals  have  suffered  from  the.se  kinds  of  plants,  but  no 
poison  cases  have  hitherto  come  undei-  my  own  notice." 

Veterinary  surgeons  also  took  the  matter  up,  and  they 
decided  the  deaths  to  be  due  to  anthrax,  the  sudden  illness 
of  the  animals  and  the  subsequent  deaths  of  many  of  them 
giving  colour  to  this  supposition  ;  but  it  does  not  appear 
that  the  anthrax  bacillus  {Bucillu-s  anthracis,  Cohn)  was 
found  in  the  spleen  of  the  dead  animals,  which  is  always 
considerably  swollen  and  full  of  enormous  quantities  of  these 
bacilli,  when  death  is  due  to  that  cause.  But  skilled 
veterinary  surgeons  may  be  wrong,  and  the  plain,  common 
sense  farmer  may  be  right  in  some  instances,  even  where 
the  death  of  cattle  is  concerned,  and  here  was  evidently 
a  case  where  further  investigation  was  desirable.  Since  the 
Homeria  plant,  which  was  known  to  be  eaten  .by  the  cows 
which  died,  was  growing  luxuriantly  in  patches  extending 

P 


210      ProceexUiujs  of  the  Royal  iSociety  of  Victoria. 

over  several  acres  whei'e  the  cattle  v^as  ieediuiii",  and  since  it 
belonged  to  a  genus  of  plants  vv^ell  known  to  have  poisonous 
properties,  it  became  a  matter  of  great  importance  to 
determine  vv^hether  this  particular  plant  was  poisonous  or 
not  when  grown  in  this  Colony,  so  we  decided  to  submit  it 
to  the  test  of  experiment. 


History  of  Outbreak  of  Disease. 

A  local  dairyman  brought  ninety-five  head  of  cattle 
from  ])addocks  a  short  distance  to  tlie  north  of  Pascoe 
Vale,  and  put  them  on  the  land  overgrown  with  Homeria. 
Next  morning,  about  twelve  head  were  found  either 
sick  oi-  dying.  Another  dairyman  Ijrought  four  head 
from  Caullield,  putting  them  on  the  same  land,  and  next 
morning  two  were  dead  and  two  sick.  Several  others  lost 
cattle  in  the  same  paddock,  and  it  is  rej^orted  that  more 
than  twent}'  have  died  altogether.  It  is  Worthy  of  note 
that  the  cattle  reared  in  the  locality  have  escaped,  and  are 
in  excellent  health,  while  only  those  fresh  to  the  district 
have  succumbed.  The  ))lant  has  now  died  down,  and  no 
more  sickness  is  reported,  but  in  the  season,  when  the  fresh, 
green,  tall  leaves  of  the  Homeria  were  fully  developed, 
it  looked  quite  a  ten)pting  green  food.  The  owner  of  one  of 
the  cows  which  was  treated  and  recovered  assured  us  that  it 
would  not  now  eat  the  plant,  although  it  had  eaten  freely  of 
it  before. 

As  regards  the  presence  of  the  Horneria  in  the  locality,  it 
may  have  been  originally  a  garden  escape,  since  these 
flowers  are  cultivated  for  their  beauty,  but  although  several 
gardens  in  the  neighbourhood  were  visited,  no  ti'ace  of  it 
could  be  found.  It  has  also  been  suggested  that  the  plant 
maj'  have  ])een  introduced  along  with  tlie  oats  formerly 
sown  in  the  paddock,  for  it  may  be  multiplied  b>'  means  of 
small  Ijulbils  which  it  })roduces  in  great  abundance,  and 
which  might  easily  get  mixed  up  with  othei'  seed. 


References  to  Homeria  being  Poisonous. 

As  to  the  poisonous  nature  of  this  genus  of  plants,  there 
are  i-eferences  in  various  standard  works,  such  as  Le  Maout 
and  Decaisne's  "System  of  Botany,"  and  Redwood's  "Su[)ple- 
iiient  to  the  Phannacopwia;"  but  we  shall  cijntent  ourselves 


Poisonous  S/)e('les  of  Honhi'rhi  at  Pascoc    Vale.    211 

with  (Hioting  tVom  sncli  a  well-known  work  as  Bentley's 
"  Manual  of  Botany,"  otli  Ed,  1887.  At  pa<,^e  703,  lie 
■Hays: — "  Moraen  (Homevia).  Some  species  of  tliis  genus, 
more  especially  that  of  M.  eollhia,  and  of  other  iridaceous 
plants  known  under  the  name  of  "Tnlp"  at  the  Cape,  have 
poisonous  properties,  and  have  been  the  cause  of  fatal 
results  to  cattle  which  have  chanced  to  eat  it.  "  Tulp  "  is 
also  poisonous  to  human  beings."  lledwood  refers  to 
Homevia  colliiia  as  (Jape  Tulip,  and  as  a  plant  well  known 
to  almost  every  child  in  the  Colony  (Ca})e  of  Good  Hope). 


Reasons  for  Investkiatiox. 

Apart  altogether  from  the  pi-actical  imjjortance  of  the 
■subject,  there  were  two  main  reasons  Avhich  induced  us  to 
enter  upon  the  investigation. 

First,  the  poisonous  ])lants  introduced  into  Victoria  have 
not  yet  been  caretully  recorded,  and  therefore  any  one  to 
which  su.spicion  attached  was  worthy  of  being  enquiied 
into,  and  its  poisonous  properties,  if  present,  determined. 
In  Queensland,  a  work  has  been  prepared  by  F.  M.  Bailey, 
F.L.8.,  Colonial  Botanist,  and  P.  R.  (lordon,  Chief  Inspector 
of  Stock,  entitled  "  Plants  Re])uted  Poisonous  and  Injurious 
to  Stock,"  but  there  is  no  mention  in  it  of  this  one,  nor 
even  of  the  natural  order  to  which  it  belongs.  Also  in  New 
South  Wales,  the  Botanist  to  the  Department  of  Agriculture, 
Mr.  Turner,  has  a  paper  on  "The  Sui)[)().sed  Poisonous  Plants 
of  New  South  Wales  (both  Indigenous  and  Exotic),"  in 
A(j.  Gaz.  Vol.  11,  Part  3,  1891,  but  thei-e  is  no  reference  to 
this  plant  or  its  order.  Hence,  a  possible  new  poison  plant, 
as  ixv  as  these  Colonies  are  concerned,  deserved  to  Ijc 
satisfactorily  determined,  in  order  to  prevent  its  further 
distribution.  Such  a  determination  is  a  necessary  pi-e- 
liminary  stej)  to  its  eradication,  just  as  in  Western  Australia, 
where  certain  poisoned  land,  as  it  is  ealled,  can  only  Ije 
obtained  on  conditions  of  exterminating  the  poison  plant, 
which  is  only  dangerous  at  certain  seasons  of  the  yem-. 

Second,  as  the  cows  which  died  at  Pascoe  Vale  were  said 
by  skilled  veterinary  surgeons  to  have  died  from  anthrax, 
and  not  from  any  supposed  poisonous  weed,  this  became  a 
strong  additional  reason  for  sifting  the  matter  to  the  bottom, 
and  seeing  if,  after  all,  the  reputed  poisonous  weed  was 
simply  an  imagination  of  tlie  cattle  owners. 

P  2 


212      Proceedlii(/.s  of  the   Hoijal  Socieff/  of   Vic/oria. 

What  is  a  Poison  Plant? 

A  ])(>ison  ]>lant  beiug  one  that  poisons,  flie  first  thing  to 
(1;>  was  to  settle  that  point,  and  tlien  liave  the  plant 
analysed,  in  order  to  determine  the  poisonous  principle  or 
alkaloid.  Mr.  P.  Wilkinson,  of  the  Government  Analyst's 
Department,  has  made  an  extract  from  the  j)lant,  but  found 
no  alkaloid  present.  It  is  attempted  to  settle  the  former 
point  in  this  pa])er,  and  in  order  to  be  clear  as  to  what 
constitutes  a  poisoi.i,  we  shall  take  the  definition  as  given  in 
Gu}^  and  Ferrier's  "  Forensic  Medicine,"  Gth  Ed.,  1 888  : — 
"A  poison  is  any  substance  or  matter  (solid,  licjuid,  or 
gaseous)  which,  when  applied  to  tl^e  bod}*  outwardly  or  in 
any  way  introduced  intf>  it,  can  destroy  life  by  its  own 
inherent  qualities  without  acting  mechanically."  And  Dr. 
Neild's  definition  is : — "  A  poison  is  a  substance  which, 
taken  into  the  body,  is  fitted  to  injure  health."  So  if  thi& 
j)lant  can  be  proved  to  cause  the  death  of  animals  feeding 
upon  it,  it  will  deserve  the  name  of  a  poison  plant, 
irrespective  of  the  symptoms  which  it  pi-oduces. 

Experiments  on  Rabuits. 

Knowing  from  tlje  experiments  of  Professor  Halford  and 
others,  that  such  drugs  as  opium  and  belladonna  can  be 
given  in  very  large  doses  to  dogs  wnth  comparativelj-  little 
eftect,  the  first  difficulty  was  to  decide  upon  and  obtain 
suitab!,'  ninmals  for  experiment.  After  due  consideration, 
we  ri'si  !\(  <1  to  try  the  effects  of  the  herb  upon  herbivorous 
animals  such  as  rabbits,  which  Mr.  Wyatt,  of  Woodlands 
Station,  very  kindly  procured  and  sent  to  us.  Three  rabbits 
ariived  on  Saturday,  1st  October,  and  were  kept  for  a  week 
on  ordinary  diet.     They  were  all  in  good  health  and  lively. 

C)n  Saturday,  8th  October,  at  4  ]).m.,  two  were  placed  in 
a  separate  cage  and  fed  upon  the  Homer'to  plant,  the  other 
being  reserved  for  future  experiment.  Fresh  plants  were 
brought  from  Pascoe  Vale,  and  the  portion  growing  above 
ground,  similar  to  that  eaten  by  the  cattle,  was  moistened 
and  given  to  the  two  rabbits.  Nothing  else  was  in  the  cage, 
and  we  saw  them  eat  freely  of  the  plant. 

C)n  Monday  morning,  lOth  October,  both  were  dead,  and 
not  expecting  such  a  sudden  effect,  we  did  not  watch 
syui])toms  very  closely.  However,  the  question  of  symptoms 
was  a  secondary  one  at  tiiis  stage,  the  primary  object  being 


Poii^onoaii  Speck''^  of  Homer ia  at  Pascoe  Vale.   213 

to  determine  whether  feeding  upon  this  plant  would  cause 
death.  On  making  the  post-mortem,  we  found  the  mucous 
membrane  of  the  stomach  and  intestines  congested.  The 
rabbit  kept  for  control  was  lively  and  well  as  usual. 

On  Saturday,  loth  October,  the  third  rabbit  which  had 
been  fed  dui'ing  the  week  on  green  food  and  was  quite 
lively,  was  placed  upon  the  same  diet.  About  6  ]).m.  it  was 
given  the  freshly  cut  Homeria  plant,  which  it  readily  ate. 
On  Sunday  morning  it  was  drowsy,  eyes  half-closed  and 
distinctly  ill.  Towards  evening,  there  wei-e  distinct  traces 
of  scouring  in  the  cage.  At  11  p.m.  it  was  still  alive,  but 
on  Monday  morning,  17th  October,  it  died  at  7  o'clock. 

Post-mortem. — Externally  marked  evidence  of  scouring 
action  on  tail,  ^^c.  The  liver  congested,  kidneys  slightly 
congested.  Bladder  full,  which  was  also  observed  in  the 
other  two  rabbits.  Before  opening  the  stomach,  little  spots 
like  ulcers  could  be  seen  in  the  w^all,  and  on  opening  it  was 
found  to  contain  a  quantity  of  the  herb  and  some  mucus. 
The  contents  were  moister  than  in  the  other  two  rabbits, 
and  marked  corrosion  was  visible.  The  mucous  membrane 
was  completely  charred  in  places,  similar  in  fact  to  what 
would  have  been  expected  if  strong  sulphuric  acid  had  been 
administered.  When  these  black  spots  were  removed,  i-ound 
patches  of  inflammation  were  visible,  and  on  hi)lding  the 
stomach  up  to  the  light,  these  ])atches  w^ere  very  conspicuous, 
looking  like  little  ulcers.  Tlie  enti'-e  intestinal  trnct  was 
congested. 

Several  more  rabbits  were  obtained  from  the  same  quai-ter, 
and  on  Thursday,  2()th  October,  two  were  again  selected 
for  feeding  on  the  Homeria  plant.  This  was  given  to 
them  about  5  ]).m.,  along  with  water,  tiud  next  inorning, 
21st  October,  one  had  died.  On  examination,  the  stomach 
was  full  and  congested.  The  second  i-abbit  was  found  dead 
on  Saturday  morning,  22nd  October,  and  the  appearance  of 
the  stomach  was  similar  to  the  tirst,  only  the  peeling  off  of 
the  mucous  membi'ane  was  more  marked.  More  of  the  plant 
had  been  eaten  in  the  latter  case. 

In  these  experiments  live  healthy  and  lively  i-a])bits  were 
taken,  and  so  sure  as  they  were  fed  upon  the  Homeria.  plant, 
so  surely  did  they  die,  within  tw(»  days  at  the  most,  while 
other  rabbits  similarly  kept,  but  fed  on  oi'dinary  f<^od, 
remained  alive,  and  as  fresh  as'  when  first  received.  Pie- 
cautions  were  taken  to  exclude  all  disturbing  elements,  so 


21+      l-'roceediuf/s  of  tJie   Roi/al  Socicf//  of  Vidorvj.. 

tliat  the  one  point  of  ditference  was,  that  one  set  of  rabbits 
were  fed  iu  the  usual  way  and  HvcmI,  while  another  set  were 
fi'd  on  the  Homevla  plant  and  died. 

It  setnis,  therefore,  reasonable  to  conclude  that  the  eating 
of  this  plant  was  the  cause  of  death,  ;ind  tliat  it  is  possessed 
of  |>oisonoiis  properties  of  an  irritant  nature. 


Experiments  ox  Cows. 

It  was  considered  quite  satisfactory  to  test  the  eti'ects  of 
eating'  the  Hoinerhi  ])]ant  upon  i-abbits,  but  in  order  to 
settle  the  matter  even  for  cows,  the  Hon.  the  Minister  of 
Agriculture  (Mr.  Oraluun)  allowed  two  cows  to  be  purchased 
for  the  purpose,  orie  to  be  fed  ujK)n  the  plant,  and  the  other 
to  be  fed,  in  the  first  instance,  in  the  usual  way,  so  as  to 
show  by  way  of  contrast  the  effects  of  the  difi'erent  feeding. 
The  two  cows  wei'e  placed  in  se])arate  loose-boxes,  and  on 
the  evening  ot  October  14,  about  (J  p.m.,  one  was  given  half 
a  bftg  of  the  freshl}'  cut  Homcrin  ]dnnt  and  water,  while  the 
other  had  a  good  su[»|>!y  of  hay  and  straw.  She  ate  greedily 
of  the  plant  (although  not  specially  starved  for  tlie  occasion) 
while  we  were  ]>resent,  along  with  Mr.  W.  H.  Stephen, 
Acting  Chief  Ins|)ector  of  Stock,  and  Mr.  E.  Rivett, 
M.R.C.V.S.  On  the  following  day,  the  cow  feeding  upon 
the  Hoimrhi  plant  was  found  to  have  eaten  about  half 
the  quantity  given  her,  and  refused  to  have  an}-  more. 
On  the  Itith  sh(!  was  lying  down  sick,  and  on  the  17th  the 
same  ;  then  on  the  forenoon  of  the  18th  the  cow  was  killed, 
and  a  post-mortem  made  b}^  Mr.  Rivett.  This  cow  was  three 
days  and  a  half  under  treatment,  and  the  eating  of  the  ])lant 
had  ])n»duced  a  scouring  action,  along  with  general  weakness, 
and  a  very  percej^tible  trend)ling  at  the  loins. 

Tlie  second  co\a',  which  was  also  placed  in  a  loose-box  on 
the  evening  of  the  Uth.  October,  was  well  fed  on  tlie  15th 
and  IGth.  She  was  made  to  fast  on  the  17th,  in  order  to 
ensuie  active  feeding,  and  on  the  KSth,  about  1  p.m.,  was 
su[»plied  with  about  a  quarter  of  a  bag  o'i  Homer  hi,  together 
with  drinking  water.  She  ate  very  greedil}^  of  the  |)lant, 
and  seemed  to  reli.sh  it.  On  the  U)th  she  was  found  lying 
down,  unable  to  rise,  and  died  that  night.  The  examination 
of  the  animal  was  made  about  mid-day  on  Thursday,  October 
20,  by  Ml-.  Stephen,  in  the  presence  ()f  both  of  us.  Tiie  four 
stomachs  w'ere    carefull}'  examined,  and    in    the    lumen   or 


Poisonous  S2)ecics  of  Homer  hi  at  Fascoe    Vale.    215 

pauncli,  there  was  marked  congestion  at  the  cardiac  end, 
wiiile  the  mucous  membrane  peeled  off,  and  was  distinctly 
inflamed.  There  was  also  considerable  scouring  of  the 
animal  before  death.  The  stomachs  of  both  cows  were 
found  to  contain  a  fair  amount  c>f  food. 

The  evidence  derived  from  experimenting  upon  the  cows, 
supplements  that  obtained  from  the  feeding  of  the  rabbits. 


CON'CLUSIONS. 

To  sum  up,  as  far  as  these  experiments  go,  there  are 
decided  indications  that  the  Homcria  plant  has  poisonous 
properties,  capable  of  causing  the  death  of  cattle  and  other 
animals,  and  this  conclusion  is  based  upon  the  following 
grounds  : — 

(1)  This  plant  is  stated  to  be  poisonous  to  cattle  at  the 
Cape,  its  native  habitat,  by  Professor  MacOwan,  Govern- 
ment Botanist  there,  and  the  probabilities  are,  that  it  is  so 
in  Victoria. 

(2)  Several  healthy  and  lively  rabbits  were  fed  upon  this 
])lant,  and  with  abundance  of  material  they  invariably 
died,  while  rabbits  fed  in  the  usual  way  remained  quite 
healthy. 

(3)  A  cow  fed  U}K)n  this  plant  also  died,  and  the  symptoms 
indicated  poisoning. 

(4)  Cows  fed  in  the  paddock  where  this  Homeria  grew 
died,  while  those  in  adjoining  paddocks  where  the  plant 
did  not  exist,  were  unaffected. 

That  the  cows  ate  the  plant  was  shown,  not  only  by  the 
undigested  remains  found  in  the  stomach,  but  from  the 
characteristic  seed-like  bulbils  found  there,  as  well  as  in  the 
droppings. 


Government  Botanist's  Description  of  Plant. 

Baron  von  Mueller,  Government  Botanist,  has  kindly 
supplied  a  description  of  the  plant,  which  is  as  follows  : — 

Homcruc  colluia,  Vent. — var.  miniata. — A  native  of 
South  Africa.  Bulb  almost  spherical,  covered  closely  by 
a  coating  of  interwoven  fibres,  between  the  layers  of  which 
numerous  minute  readily  sprouting  bulbils  are  concealed. 
Whole     plant    to    3    feet    high,     but    usually    much    less, 


210      Proceedings  of  flu-   Royal  Society  of  Victoria. 

variable  also  in  more  or  less  robustness  or  slenderness,  (jften 
somewhat  branched.  Leaves  linear  to  1|  feet  long  to 
'6  inch  bi'oad,  but  frequently  of  much  less  size,  always 
channelled  and  gradually  much  narrowed  upwards  ;  grey- 
green  above,  dark  green  ber)eatli,  slightlj^  streaked,  small 
bulbils  also  formed  occasionally  in  the  axils  of  some  leaves. 
Inflorescence  fascicularly  compound  when  well  developed. 
Somewhat  paniculate,  the  supporting  lowest  floral  leaf  often 
much  elongated,  clasping  at  the  base.  Bracts  comparatively 
long,  nuich  pointed,  the  outer  green,  the  inner  smallei-, 
gradually  colourless,  and  very  tender.  Flower  stalks  to 
2  inches  long,  though  often  shorter.  Some  of  the  stalklets 
finally  to  1-^-  inches  long,  all  enclosed  in  longitudinally 
convolute  bracts.  Flowers  almost  horizontally  expanding, 
very  tender,  stem  shrivelling.  Tube  of  the  odyK  thinly 
cylindric,  pale-green,  darker,  six  streaked,  generally  about 
I  inch  long.  Lobes  of  the  calyx  three  (or  exceptionally  two), 
petal-like  lanceolar-ovate,  about  |  inch  long,  yellowish 
towards  the  base,  otherwise  almost  brick-coloured,  or  neai-ly 
orange- coloured.  Petals  similar  to  the  calyx-lobes,  but 
somewhat  nari-ower,  three  (or  eJvceptionally  two),  along  with 
the  calyx-lobes  tv.'isted  after  flowering,  finally  deciduous. 
Stamens  three  (seldom  two),  much  shorter  than  the  calyx- 
lobes  and  petals.  The  three  anthers  erect,  seated  on  the 
yellowish  narrow  staujinal  tube,  about  |  inch  long,  yellow, 
broad-linear,  blunt,  at  the  l)ase  minutely  bi-lobed,  bursting 
marginally.  Style  tilif(M'm,  about  as  long  as  the  stigmas. 
These,  as  well  as  the  anthers  opposite  to  the  calyx-lobes, 
three  (or  e.  ceptionally  two)  in  number;  hardl}'  extending 
beyond  the  anthers,  yellowish,  linear-cuneate,  with  numei'ous 
dilated  bi-lobed  crenulated  and  ciliolated  summit,  and  witli 
two  small  tender  inner  appendages.  Ovulary  quite  connate 
with  the  calyx-tube,  three-celled  (or  seldom  two-celled), 
cylindric  and  somewhat  angular.  Ovules  very  numerous, 
fixed  along  the  axis.  Fruit  dry,  trigonous  cylindric, 
dehiscent,  niany  seeded.  The  flowers  are  distinctly  smailei- 
than  those  of  Hoineria  rolJina,  their-  petals  and  calyx-lobes 
are  more  acute  and  of  a  ligliter  red  ;  also  less  venulated,  and 
the  staminal  tube  is  olabrous. 


Conclusion. 

It   will     be    seen    from    the   foregoing    description    what 
)nderful    powers  of   j)ro[)agation    this    plant  possesses   by 


Poiso}i<)us  Species  of  Hunierla  at  Pascoe   Vale.    217 

means  of  its  nuinerous  i-e[)ruducfcivo  seed-like  bulbils. 
It  can  easily  be  uiidersbood  liow  it  has  overspread  thr 
paddock  by  this  means  alone.  Its  showy  and  attractive 
flowers  likewise  rendered  it  an  object  of  interest  and  beauty 
to  the  numerous  wayfarers,  particularly  on  Sundays,  and  as 
handfuls  of  the  plant  were  taken  away,  it  would  thus  be 
spread  over  a  large  area,  and  carried  to  different  districts. 
It  is  known  in  other  places  besides  Pascoe  Vale,  but  now 
that  its  poisonous  properties  are  unmasked,  it  is  hoped  that 
this  brief  notice  of  it  may  lead  to  its  being  promptly 
destroyed  in  any  garden  or  cemetery  where  it  may  exist. 


Art.  XIX. — R<'i>urt  of  the  (Johvuut/^e  of  the  Rof/ai  Society 
of  Victoria,  consistiru/  of  Professucs  Kernot,  Lyle, 
and  Masson,  and  Messrs.  Ellkrv,  Love,  and  White, 
appointed  to  arrainje  for  the  rarrnliKj  oat  of  the 
Gravity  Survey  of  Australasia. 

To  THE  Royal  Soc^etv  of  Victoria. 

Gentlemen, — In  laying  befoi-e  you  tliis,  the  Second 
Annual  Report,  your  Committee  has  much  pleasure  in 
informing  you  that  the  work  of  the  Survey  has  now 
commenced.  The  penduhuns  and  other  apparatus  lent  by 
the  Royal  Society  of  London — of  whicli  a  description  is 
appended — -have  been  received,  and  ei'ected  in  a  cellar  at  the 
Observatory,  kindly  phiced  at  the  disposal  of  the  survey  hy 
tlie  Government  Astrononiei".  Tlie  observing  telescope  sent 
with  the  apparatus  ])rovt'S  to  be  somewhat  inconvenient, 
and  it  is  projXKsed  to  employ  a  different  arrangement.  The 
stand  for  tlie  air-])um])  was  badly  packed,  and  found  to  be 
broken  on  its  ariival ;  otherwise  the  instruments  were 
in  \'ery  fair  order.  It  is  ])roposed  to  devote  the  next  few 
months  to  a  careful  examination  of  the  effects  of  temperature 
and  i»ressure  on  the  times  of  oscillation  of  the  pendulums; 
such  an  investigation  being  rendered  especially  necessary 
by  the  very  considerable  changes  of  temi)erature  to  which 
the  instruments  may  possibly  be  exposed  in  tlie  course  even 
of  a  single  set  of  swings.  The  values  of  the  temperature 
and  pi-essui-e  coefficients  for  the  pendulums  numbered  (4)  and 
(1821)  were  worked  out  for  the  purposes  of  the  Indian 
Survey;  but  the  constants  of  the  third  pendulum,  numbered 
(11),  have  not  yet  been  determined.  General  Walker 
assumed  them  for  the  pui-pojjes  of  tlie  Greenwich  and  Kew 
observations  (lately  completed)  to  agree  with  those  of  the 
other  two  ;  but  your  Committee  is  of  opinion  that  the 
matter  requires  further  investigation. 

The  question  as  to  the  construction  of  a  new  pendulum 
has  received  a  o-ood  deal  ot  attention  fiom  your  Committee 


Ih'poii  oil   the  Grarlfi/  Stn-rr)/  of  Australasia.     21.4' 

during  the  past  year.  Fortunati'ly  the  Royal  Society  of 
London  has  forestalled  the  diseussion,  and  added  ])enduluni 
(11)  to  the  two  originally  asked  tor.  The  difticulty  and 
expense  attending  the  constructicni  of  a  new  pendukini  has 
thus  been  avoided. 

E.  F.  J.  Love,  Secretary. 


Appendix. 

De.scription  of  the  apparatus  to  be  enn)hjyed  in  tlie 
Gravity  Survey  of  Australasia,  by  E.  F.  J.  Love,  iM.A. 

In  drawing  up  a  description  of  tlie  a[)paratus,  we  may 
consider  se))arately,  [a)  the  pendulums,  {h)  the  clock,  (c)  the 
vaciunn  api)aratus  and  its  accessories. 

{a)  The  pen<luhims  are,  undoubtedly,  the  most  important 
portion  of  the  api>aratus.  The  three  which  it  is  pro))osed  to 
employ  are  all  constructed  of  the  same  materials,  and 
practically  identical,  both  in  form  and  dimension.  They 
aie  of  the  kind  known  as  'Invariable  Pendulum."  The 
form  is  a  flat  bar  of  ])late  brass  5  feet  '1  inches  long, 
Ui3  ijich  thick,  and  1 7  inches  broad,  for  a  distance  of 
40  inches  from  the  upper  end.  The  remaining  portion  of  the 
bar,  termed  the  "tail-piece,"  is  lenticular  in  section,  reduced 
to  a  breadth  of  (VJ  inch,  and  terminates  in  a  point.  Just 
above  the  tail-|)iece  is  a  tiat  circular  brass  bob,  G  inches  in 
diameter  and  J-.S  inches  thick,  which  is  fastened  to  the  bar 
l>y  solder  and  i-ivets.  The  knife-edge  is  a  prism  of  very  hard 
steel,  adjusted  per|)endicular  to  tlu^  plane  of  the  bar,  and 
attached  by  means  of  a  stout  T  head.     It  is  2  inches  long, 

0  2o  inch  in  height,  and  equilateral  in  secti<.>n,  save  that  the 
edge  on  which  the  (oscillations  are  performed  is  ground  to  an 
angle  of  120".  The  planes  on  which  the  pendulum  oscillates 
consist  of  two  pieces  of  polished  agate,  ground  true  and  set 
in  a  heavy  brass  frame  suppoi-ted  on  ver\-  massive  levelling 
screws.     Each  })endulum  has  its  own  set  of  planes. 

All  three   pendulums   are   about  70  years  old,  and  have 

1  teen  repeatedly  used  for  gravity  survey  work;  in  which 
the}^  have  given  such  consistent  results  as  to  warrant  the 
belief  that  the\'  have  reached  a  condition  of  api)roximate 
e(|uilibrium  as  regaixls  molecular  change.  For  a  statement 
of  their  history,  reference  may  be  made  to  the  "  Re])ort  of 
the  Great  Trigont)metrical  Survey  of  India,"  Vol.  V, 
Appendix  \^.  30. 


2:^0      r  rowed  lugs  of  the  Royal  Societi/  of  Victoria. 

With  the  pendulums  is  su])])]ied  :i  "  dummy  pendulum," 
of  identical  material  and  dimensions,  into  which  two  holes 
are  sunk  for  the  reception  of  the  bulbs  of  a  pair  of 
thermometei-s.  The  dummy  is  placed  in  the  same  vacuum 
chamber  with  the  pendulum  when  vibrating,  and  close  to  it. 
Their  temperatures  may  accordingly  be  assumed  as  identical, 
and  the  temperature  of  the  dummy,  as  given  by  the 
thermometers,  can  be  employed  for  deteriiiining  the  temper- 
ature corrections  to  be  applied  to  the  pendulum.  Tlie 
corrections  to  be  applied  to  the  thermometers  have  been 
determined  at  Kew. 

(b)  The  clock  employed  for  the  (jbservation  of  coincidences 
is  a  siderial  clock,  made  by  Shelton,  and  was  used  by  Sabine 
on  his  exjiedition  in  1822.  Its  mean  daily  rate  is  very 
constant,  but  it  is  subject  to  rather  considerable  horar}- 
fluctuations  of  rate.  The  clock  has  an  arrangement  which 
allows  of  its  being  re-wound  without  loss  of  driving  power 
during  the  winding. 

(c)  The  vacuum  apparatus  consists  of  a  cylinder  of  sheet 
copper,  half  closed  at  the  top  by  a  thick  brass  plate  for 
supporting  the  agate  planes,  and  closed  in  above  this  by 
a  glass  bell,  ground  to  lit  the  brass  plate  ;  it  is  closed  at  the 
bottom  by  a  metal  hemisphere.  It  has  one  glass  window 
about  lialf-way  up,  through  which  the  thermometers  are 
read,  and  four  others  in  the  plane  of  the  tail-piece  of  the 
pendulum.  Through  one  pair  the  coincidences  are  observed, 
the  other  pair  allowing  a  side  view  of  the  tail-piece,  which 
is  necessary  for  determining  the  amplitude  of  its  vibration. 
To  the  sill  of  the  back  window  is  attached  a  bi-ass  plate 
bearing  two  scales  at  right  angles  to  each  other  etched  on 
ground  glass,  and  with  well  blackened  divisions  for 
measuiing  this  amplitude. 

The  cylinder  is  supported  by  three  large  levelling  screws 
on  a  heavy  iron  girder,  which  is  itself  bolted  to  a  very 
massive  timber  framework  fastened  together  with  iron  bolts 
and  clam])s.  The  massiveness  of  the  cylinder  and  fram«^ 
render  it  quite  impossible  for  the  oscillations  of  the 
pendulum  to  be  communicated  to  the  supports. 

The  starting  and  stopping  of  the  pendulum  is  perf  )rnied 
by  means  of  levers  worked  from  outside  the  cylinder  by 
metal  rods  passing  through  stuffing  boxes,  and  cases  filled 
with  oil  to  pi-event  leakage  of  air. 


R(q>oi't  on  the.   Gnwilf/  Survey  of  AustniUma.     221 

A  tap  attached  to  the  side  of  the  cylinder  is  connected  by 
rubber  tubing  to  a  Siphon  barometer,  and  a  second  tap 
allows  of  the  attaclinient  of  an  air-pump  in  order  to  reduce 
the  pressure  to  any  desired  amount,  wliich  is  measured  on 
the  Siphon  barometer. 

As  the  tail-piece  is  only  a  little  way  above  the  ground, 
the  short  telescoj)e  with  diagonal  eye-piece  sent  with  the 
apparatus  is  highly  inconvenient.  It  is  proposed  to  view 
the  coincidences  by  means  of  a  long  telescope  of  considerable 
aperture,  iuclined  at  a  small  angle  to  the  ground,  and 
carrying  a  plane  mirror  in  front  of  the  object  glass,  so  as 
to  reflect  the  image  of  the  apparatus  in  a  nearly  vertical 
direction.  This  method  will  result  in  a  considerable  saving 
of  light,  and  a  much  m(»re  than  considerable  addition  to  the 
comfort  of  the  observei-.  Anyone  who  has  had  experience 
in  really  delicate  })hysical  work  will  understand  the 
importance  of  these  considerations  to  the  accurac}'^  of  the 
experiments. 

In  order  that  the  images  of  the  detached  and  ch^ck 
j)endulums  may  be  in  the  same  plane,  a  large  lens  is 
provided,  by  means  of  which  an  image  of  the  clock 
pendulum  is  thrown  on  the  ground  glass  scale  inside  the 
cylinder.  The  lens  is  mounted  on  a  brass  angle  piece, 
which  slides  on  a  brass  frame  attached  to  a  wooden  stand. 
The  stand  rests  by  means  of  three  levelling  screws  on  a 
]»lank  bolted  to  the  framework  which  supports  the  cylinder. 


AlJT,  XX. — RepoH  nf  the  Ci'eiaoiloiK  Cohomitlee  oj  the 
lioijal  Society  of  Vlctorhi,  ((ppolhted.  to  eiujiiire  into 
and  report  upon  "  CreDudiori  "  and  other  methods  of 
ditipo.siii;/  of  th.e  dead,  with,  pariit-vJii.r  re</ard  to 
hyyieve  <iiul  economy. 

To   THE    PUKSIDKNT  AMD    MkMBEKS  OF   THE   RoYAL  SOCIETY 

OF  Victoria. 

Your  Committee  lias  tlie  honour  to  lepoit  that  it  ha.s 
lield  two  preliminary  and  tliiee  general  meetings,  and 
has  considered  the  various  methods  pro]iosed  for  the  sanitary 
and  economic  disposal  of  the  dead.  Your  Conunittee  linds 
from  tiie  evidence  collected,  that  burial  now  entirely  fails  to 
satisfy  the  demands  of  hygiene.  There  aic  the  strongest 
reasons  for  concluding  that  giaveyards  have  been  in  the 
past,  and  are  now,  prolific  sources  of  deadly  disease,  not 
only  by  reason  of  mephitic  vapouis  arising  thence  into  thi; 
atmosphere,  but  also  b}'  ])ei'colation  of  putrid  liquid  matter 
in  water  diainage  to  considerable  distances.  Many  cases 
have  notoriously  occurre<l,  in  which  wells  have  been  demon- 
strabl}'  poisoned  in  this  manner  at  long  distances  fiom  the 
source  of  infection.  The  i-isk  of  this  is  immens(,'ly  aggravated 
as  ])Opulation  increases.  In  America,  Europe,  and  Victoria 
itself  the  towns  grow  and  surround  the  cemeteries,  which 
soon  become  full.  New  ones  ai'e  formed  further  away,  and 
the  land,  being  imperatively  requiied  by  the  living,  the 
bodies  are  unceremoniously  removed  from  the  old  giuveyards,- 
which  ai'e  generally  used  tor  building  blocks,  public  gardens, 
and  other  purposes.  The  removal  is  a  dangerous  process, 
the  distuibance  of  the  putrid,  poisonous  remains  having 
been  almost  certainly  the  cause  of  outbreaks  of  malignant 
disease  epidemics.  It  is  practically  impi:>ssible  to  find  a  site 
for  a  cemetery  anywhere  in  the  vicinity  of  towns,  such 
that  there  would  be  no  danger  to  healtii  to  the  living,  in 
which  the  air,  the  water,  and  the  earth  of  tiie  neiglibouriiood 
would  be  secure  from  the  dea;llv  containination. 


Report  of  the  Crem<Ulo)i  Committee.  223 

As  regards  Economy. — The  disposal  of  the  dead  by  l)aiiai 
is  already  an  oj)])ressive  charge  to  the  large  majority  of  the 
population  wherever  it  is  numerous.  Cemeteries  are  made 
further  and  further  away,  and  the  longer  conveyance  )nateri- 
ally  eniiances  the  expense,  and  must  continue  to  do  so  more 
and  more.  The  unavoidable  crowding  of  cemeteries  has  also 
had  the  efiect  of  destroying,  or  outraging,  the  reverential 
sentiment  which  fondly  regarded  burial  as  finally  providing 
foi-  the  permanent  and  undisturbed  repose  of  the  departed. 

After  being  first  filled  with  corpses  to  the  extent  of  from 
twelve  to  twenty-two  (seventy  according  to  the  Duke  of 
Westminster^ — Times,  December  9,  1889)  in  each  grave,  in 
nearly  all  old  cemeteries,  the  ground  is  similarly  used  over 
and  over  again  at  intervals  of  a  very  few  years  ;  and  the 
purchase  of  space  for  a  grave  or  vault,  supposed  at  the  time 
to  secure  ownership  in  perpetuity,  is  a  delusion  and  a  snare  ; 
as  a  mattei-  of  fact,  headstones  are  broken  up  for  road  metal 
.&c. ;  the  coliins  are  bui-ned,  and  the  bones  used  for  manure 
or  shot  down  as  rubbish.  No  respect  is  shown  for  tiie 
remains  of  the  dead,  or  for  the  feelings  of  their  living  represen- 
tatives. All  ideas  of  sanctity  and  reverence  are  violated. 
The  \ise  of  vaults  scarcely  delays  the  process.  Persons  who 
have  wealth  and  influence  may,  if  watchful,  be  able  to  delay 
the  sacrilege  during  their  lives,  but  the  next  generation  loses 
both  inclination  to  resist,  and  power  to  postpone  it. 

The  method  pursued  by  the  Parsees  is  nmch  less  objection- 
able hygienically  considered.  It  consists  in  simple  exposure 
on  the  top  of  a  tower  for  vultures  to  dismember  and  devour 
the  corpse.  This  does  not  engross  an  increasing  quantity  of 
land,  or  involve  the  desecration  of  being  dug  up  again  in  a 
few  years  to  make  loom  for  some  one  else,  and  ]ierha})s  oH 
being  shot  as  rubbish.  Still  less  does  it,  like  burial,  [joison 
the  eai-th,  air,  and  water,  to  the  destruction  of  the  living ; 
but  it  is  practised  by  but  a  small  section  of  the  populatimi 
of  India,  outside  of  which  it  has  no  advocates,  and  is  not 
likely  to  extend. 

Desiccation  has  been  leconnnended,  and  may  be  adapted 
to  a  very  dry  climate,  but  apparently  not  to  others.  In  the 
Catacombs  at  Malta,  Palermo,  and  some  other  places  it  has 
been  used  ;  but  the  results  are  such  as  to  disgust  strangei's, 
and  present  such  featui-es  of  irreverence  and  deseci-ation,  as 
to  preclude  its  wider  adoption.  It  may  be  possible  to  secure 
hygienic  I'esuJts  by  it,  but  there  seems  to  be  much  more  risk 


224     ProceedliKjs  of  iJie  Ji<>y((l  8<>cieiy  of  Victoria. 

of  the  coiitraiy.  A  movement  in  favour  of  desiccation  has 
occnned  in  America,  hut  your  Committee  has  no  leason  or 
wisli  to  think  that  it  has  any  chance  of  success. 

The  use  of  quicklime  has  been  successfully  tried  in  several 
instances,  where  large  numbers  killed  in  battle  had  to  be 
rapidly  disposed  of,  and  in  some  other  cases.  It  does  not, 
however,  appear  to  be  adapted  tor  general  use,  particularly 
where  lime  is  not  readily  and  cheaply  procurable. 

Another  method  has  been  suggested  of  disposing  of  the 
dead,  by  simply  immersing  them  in  a  bath  or  tank  of  fused 
alkali,  in  w4iich  they  entirely  disappear  without  leaving  any 
discoverable  residue.  The  cost  and  feasibility  ol  this  method 
would  depend  upon  the  abundance  and  accessibility  of  the 
material,  but  it  seems  questionable  whether  it  would  ever 
commend  itself  to  public  sentiment.  There  appears,  however, 
to  be  no  hygienic  objection  to  it. 

The  expedient,  which  seems  to  be  in  a  fair  way  to  super- 
sede burial,  is  Cremation— an  old  one  revived,  and  practised 
widely  to-day.  Cremation  is  general  in  Japan,  and  in  India, 
where  the  Government  has  successfully  introduced  improved 
incinerators  to  expedite  and  perfect  the  primitive  process  in 
use  by  the  Hindoos.  Cremation  is  the  simplest,  cheapest, 
and  most  hygienic  of  all  ;  it  can  be  easily  eflected  wherever 
there  are  combustibles,  and  it  appears  particularly  adapted 
for  use  in  cities,  being  lapid,  economical,  final  and  complete. 
The  residue  is  small,  innocuous,  and  easily  pieserved  in  urns, 
the  cost  of  which  is  trifling.  Cremation  is  becoming 
popular  in  Italy,  where  it  is  rapidly  extending.  Large 
numbers  are  now  cremated  in  Paris,  and  at  Gotha.  In 
England,  its  progress  has  been  even  more  rapid  than  any- 
where else,  except  Rome.  At  Milan,  679  cremations  have 
been  eflected  in  14  years,  but  only  227  in  the  first  7  years. 
At  Lodi,  38  in  13  years.  At  Rome,  where  the  practice  has 
grown  more  lapidly  than  at  any  other  place  in  Italy,  there 
have  been  297  cremations  in  7  years.  At  21  towns  in  Italy 
there  were  in  all  1468  cremations  in  the  14  years  ending 
with  1890.  At  Woking,  in  Suarey,  the  first  cremation  took 
place  in  1885,  and  the  numbers  since  cremated  there  yearly, 
are,  8,  10,  18,  28,  40,  54,  and  99  in  1891—253  in  all;  the 
increase  being  more  uniformly  progressive  than  even  at 
Rome,  which  began  with  15,  and  ended  with  90  in  7  years, 
and  had  fewer  in  188G  and  1887  than  in  1885.  The  Duke 
of  Bedford,  Lord  Bramwell,  and   Mr.  Wm.  Eassie,  were  all 


Report  of  the  Cremation  Committee.  225 

cremated  during  the  current  year,  and  Crematories  are 
being  established  at  Manchester,  Liverpool,  Ilford,  Darling- 
ton, and  elsewhere. 

The  great  advantages  of  Cremation  appet^r  to  be — Firstly, 
the  perfect  extinction,  with  the  corpse,  of  the  possibility  of 
communication  by  it  of  any  disease  to  the  living.  Secondly, 
its  economy.  The  cost  at  Paris  is  only  two  francs,  and  it  is 
less  in  Japan  and  India.  There  is  eveiy  reason  to  believe 
that  it  could  be  done  in  Melbourne  for  a  guinea  each  at 
most,  including  examinations,  memorial  urn,  &c.  Carriage 
must  sometimes  form  a  comparatively  important  item  in  the 
cost.  It  can,  however,  be  much  reduced,  as  portable  iron 
crematories  have  been  successfully  constructed  for  military 
purposes,  and  will  no  doubt  come  into  general  use.  Thirdh-, 
its  finality.  Cremation  will  abolish  at  once  all  the  shocking 
desecration  which  is  now  inseparable  from  the  burial  system. 
Fourthly,  the  innocuous  residual  ashes,  less  than  a  quart  in 
quantity,  can  be  preserved  in  an  urn  of  aesthetic  n;iaterial 
and  device,  and  deposited  either  in  a  public  institution  (or 
Columbarium),  or  confided  to  the  care  of  the  family  ;  with 
Fifthly,  the  satisfactory  certainty  to  all  concerned,  that  the 
body  itself  can  never  afterwards  be  subjected  to  disturbance, 
insult,  or  desecration,  or  cause  incalculable  harm  to  others. 

The  only  apparently  plausible  objection  that  has  ever  been 
urged  against  Cremation  is,  that  the  body  can  never  after- 
wards be  available  as  evidence  in  cases  of  murder,  particuarly 
by  poison.  A  case,  however,  occurred  at  Milan,  which  goes 
far  to  prove  that  the  risk  is  actually  greater  in  case  of  burial 
(see  Robinson,  "Cremation  and  Urn  Burial,"  [)p.  177-8). 
The  parents  of  a  deceased  child  obtained  all  the  certificates 
necessary  for  its  burial,  before  resolving  to  have  it  cremated. 
The  additional  certificates  however,  which  were  required  at 
the  Crematorium,  elicited  the  fact  that  the  child  had  been 
poisoned  accidentally  by  eating  sweetmeats  containing 
copper.  Your  Committee  would  strongly  recommend  that 
no  system  whatever  be  tolerated  whicli  does  not  provide 
amply  strict  examinations  to  obviate  the  possibility  of  such 
facts  passing  undetected. 

An  Act,  legalising  Cremation  under  conditions,  has 
lately  been  passed  by  the  South  Australian  Legislature  at 
Adelaide. 

Lastly,  the  legal  aspect  of  the  question  remains  to  be 
considered. 

Q 


220     Froceedinc/s  of  the  Royal  Society  of  Victoria. 

Sir  Jas.  F.  Stephen's  judgment  in  the  case  of  Dr.  Price, 
ill  1874,  set  at  rest  the  question  of  the  legality  of  Cremation 
ill  England,  and  decided  that  there  was  then  no  law  against 
it  there,  so  long  as  no  nuisance  was  caused.  Of  course  no 
system  of  disposing  of  the  dead  should  be  tolerated,  unless 
all  that  can  be  called  a  nuisance  is  absolutely  prevented.  The 
objection  to  burial  is  that  it  produces  evils  far  worse  than 
nuisances.  Since  the  judgment  in  question,  the  Cremation 
Society  of  England,  though  previously  deterred  by  the 
discountenance  of  the  Home  Secretary,  proceeded  at  once  to 
cremate,  and  has  continued  to  do  so  since.  The  same  view 
appears  to  have  been  officially  taken  here,  in  the  Metro- 
politan General  Cemetery  Bill,  which  was  introduced  by  the 
Government  in  the  Legislative  Assembly  in  1891,  but  made 
no  further  progress.  The  existence  of  this  Bill  implies  that 
no  legal  objection  to  Cremation  could  be  discovered.  It 
provides  "  for  the  establishment  and  management  of  a 
Metropolitan  General  Cemetery "  at  Frankston,  with  nine 
managers ;  two  to  be  appointed  by  the  Government,  and 
seven  to  be  elected  by  the  Councils  of  eighteen  city  and 
surburban  corporations.  £20,000  was  to  be  granted  from 
the  consolidated  revenue  to  start  with,  and  the  corporations 
were  to  contribute  .€2500  a  year,  until  the  fees  to  be 
charged  should  amount  to  a  sufftcient  sum  to  defray 
expenses.  The  cemetery  consists  of  8008  acres,  worth 
,€15,000;  distance  from  Melbourne  20  miles.  It  is  11 1 
miles  round,  and  the  cost  of  fencing  it  has  been  estimated  at 
X^24,000.  More  thousands  are  required  for  a  short  branch 
railway.  The  Bill  provides  that  the  managers  may  make 
regulations,  to  be  approved  by  the  Governor  in  Council, 
prescribing  fees  for  burials,  &c.,  and  also  for  cremations. 
Section  71  provides  that  any  one  may  direct  by  Will  or 
otherwise,  that  his  body  shall  be  cremated,  and  that  his 
executors  or  others  may  carry  his  direction  into  effect,  in 
the  cemetery,  under-regulations  to  be  made  under  Section  77. 
The  admission  that  Cremation  is  not  illegal  is  something, 
and  the  attempt  to  legalise  it  is  more.  But  cremation  at  a 
distance  of  20  miles  is  useless.  There  is  ample  proof  that  its 
proper  performance  within  a  city  admits  of  no  reasonable 
objection.  Persons  living  next  door  would  not  even  know 
that  it  was  in  progress,  and  in  itself  it  is  essentially  purifying 
as  well  as  innocuous. 

Hj^giene  demands  the  reduction  to  a  minimum  of  the 
time  and  distance  between  the  death  of  the  body  and  its 


Report  of  the  Cremation  Committee.  227 

iinal  disposal.  One  weighty  objection  to  burial  is,  that  it 
must  be  as  far  from  the  city  as  convenient,  notwithstanding 
the  cruel  inconvenience  and  expense  to  the  mourning 
relatives  in  the  performance  of  their  sacred  duties.  Their 
strong  claims  to  sympathy  and  consideration  appear  to  have 
been  wholly  ignored  in  the  Frankston  scheme.  But  in 
Melbourne  now,  hundreds  of  pious  mourners  visit  the  graves 
of  their  departed  relatives  weekly,  and  even  more  frequently, 
to  plant  and  carefully  tend  flowers  around  them.  They 
would  be  cruelly  debarred  from  performing  this  pious  duty 
by  the  extra  cost  and  time  involved  in  frequent  journeys, 
even  by  railway,  of  52  miles.  Cremation  would  abolish  this 
difficulty  entirely.  Instead  ot  having  to  neglect  these  duties 
altogether,  or  to  travel,  say  weekly  or  daily  to  Frankston  to 
fulfil  them,  they  would  have  the  actual  pure  ashes  them- 
selves, in  an  elegant  urn  or  other  receptacle,  in  either  the 
mortuary  chapel,  or  family  household,  where  the^y  could 
fulfil  their  cares  and  soothe  their  feelings  by  daily  viewing 
them,  and  decking  them  with  fresh  flowers. 

As  regards  economy,  compare  a  central  City  Crematory 
and  Mortuary  Chapel,  costing  perhaps  £2000  or  £3000,  and 
2s.  6d.  or  8s.  worth  of  fuel,  and  a  fee  of  a  guinea,  with  a 
Cemetery  26  miles  oft',  costing  for  land  £15,000,  fencing 
£24,000,  and  several  thousands  more  for  a  branch  railway  to 
it.  But  these  are  of  minor  importance  concerning  the  state 
contribution  only.  The  snlient  point  is,  what  will  be  the 
charges  for  each  funeral  to  bereaved  mourners — the  people  ? 
The  deaths  in  Melbourne  may  now  be  taken  at  10,000 
yearly  (10,412  in  1889,  and  9,207  in  1890,  Hayter),  i.e.,  25 
to  28  daily.  oPlO  is  sureh'  a  low  average  for  ordinar}' 
funerals  now,  and  transport  is  always  and  necessarily,  a 
formidable  extra ;  and  however  performed,  the  26  miles 
cannot  but  add  largely  to  the  expense,  falling  upon  the 
unfortunate  moarners  in  the  shape  of  undertakers'  bills,  thus 
augmented  by  at  least  25  or  30  per  cent. 

The  fees,  also,  of  unknown  amount,  would  also  fall  upon 
them,  and  to  provide  the  projected  embellishments  upon  the 
scale  hinted  at,  the  fees  must  be  anything  but  light.  Even 
supposing  that  the  increase  altogether  might  not  exceed 
50  per  cent.,  £15  for  each  funeral,  multiplied  by  10,000, 
would  be  at  least  £150,000  to  be  paid  yearly  by  the  i:>eo2:)le, 
beside  the  contribution  of  the  state.  Cremation  would  per- 
form the  whole  service  for  probably  £1  Is.  each,  or  £11,000 
a  year,  in  a  few  crematories  costing  perhaps  £2000  each 

Q  2 


228     Proceedings  of  the  Royal  Society  of  Victoria. 

Your  Committee  for  all  these  reasons  confidently  recom- 
mends Cremation  as  incomparably  the  best  solution  of 
every  difficulty,  particularly  on  hygienic,  sentimental,  and 
economical  grounds. 

Your  Committee,  however,  also  feels  called  upon  to 
remark  here  that  not  only  in  the  Bill  in  question,  but  also 
in  the  Cemeteries  Act  of  1890  (No.  1072,  now  in  force), 
some  of  the  most  important  facts  which  should  be  kept  in 
view  in  disposing  of  the  dead  seem  to  have  been  entirely 
ignored.  Its  framers  appear  to  rely,  with  most  mistaken 
confidence,  upon  hermetically  closed  coffins  and  cemented 
vaults  to  prevent  the  escape  of  the  poisonous  gases  generated 
in  decomposition.  This  is  a  fallac3^  Siicli  escape  cannot  he 
prevented.  Your  Committee  cannot  do  better  than  repeat 
the  decisive  testimony  of  Sir  John  Simon,  the  eminent 
Sanitarian  (quoted  in  the  Duke  of  Westminster's  letter  to 
the  "  Times,"  dated  December  9,  1889) : — 

"  The  leaden  coffin  soouer  or  later  yields,  and  gives  vent  to  its  fatal 
contents.  The  most  successful  attempt  at  hermetical  enclosure  does  not 
reach  beyond  ijostponemeut  of  the  effusion  through  the  atmosphere  of  the 
products  of  decomposition.  Overcrowding  the  dead  causes  the  soil  to  be 
saturated  and  supersaturated  with  decomposing  animal  matter,  polluting  the 
water-springs  and  vitiating  the  air  ;  and  it  is  by  the  air,  vitiated  by  organic 
matter  undergoing  decomposition,  that  epidemicsand  infectious  diseases  most 
readily  diffuse  their  poison  and  multiply  their  victims." 

Your  Committee  has  made  its  deliberate  recommendation 
upon  the  evidence  before  it.  It  is  perhaps  scarcely 
necessary  to  say  that  that  recommendation  does  not  include 
that  those  who  prefer  burial  should  not  be  as  free  as  the 
advocates  of  cremation  to  do  what  they  prefer.  At  the  same 
time,  it  seems  clear  that  both  the  j^^to^^c  advantages  of 
cremation,  and  the  ])ublic  dangers  of  burial,  are  infinitely 
more  important  and  practical  than  any  lorivate  predilections 
either  way. 


(Signed) 


Llp:wellyn  D.  Bevan,  D.D.,  Member. 

J.  Talbot  Brett,  M.D., 

D.  A.  Gkesswell,  M.D., 

William  C.  Kernot,  M.A., 

William  Lynch, 

Orme  Masson,  M.A., 

William  L.  Mullen,  M.D., 

James  Edward  Neild,  M.D.,     „ 

G.  A.  Syme,  M.D., 

H.  K.  RusDEN,  Hon.  Secretary. 


Art.  XXI. — Report  of  the  Port  Pliillip  Biological  SuvveAj 
Committee,  1892. 


Your  Committee  regret  that  tliere  are  no  results  to  rej)ort 
as  received  from  the  specialists  in  Eui'ope  to  whom  material 
was  forwarded,  though  we  have  information  that  the  work 
is  in  hand.  During  the  course  of  next  year,  Professor 
Spencer,  who  is  on  the  Committee,  is  visiting  Europe,  and 
hopes  to  arrange  for  an  early  publication  of  the  descriptions. 

Professor  Tate,  of  Adelaide,  has  returned  the  specimens  of 
the  Nudibranch  Mollusca,  as  he  finds  himself  unable,  through 
pressure  of  other  duties,  to  undertake  their  determination. 

Dr.  Dendy  has  continued  liis  studies  on  the  Sponges,  and 
has  worked  out  the  classification  and  much  of  the  detailed 
anatomy  of  the  Calcarea  Heterocoela.  He  is  publishing  a 
Synopsis  of  this  work  in  the  Proceedings  of  our  Society,  and 
anatomical  accounts  in  the  European  journals. 

Your  Committee  have  incurred  no  expenses  during  the 
year. 

A.   H.  S.  Lucas,  Hon.  Secretary. 
Nov.  8,  1892. 


MEETINGS    OF    THE    EOYAL    SOCIETY. 

1892. 


[N.B. — The  remarks  and  speeches  in  the  discussions  are 
taken  down  verbatim  by  a  shorthand  writer,  and 
afterwards  written  out  at  length  with  a  tyjjewritei', 
for  reference  and  reproduction,  if  required  ;  and  there- 
fore, more  is  seldom  given  herein  than  an  indication  of 
their  general  drift.  If  any  person  should  wisli  to  refei 
to  the  verbatim  report,  he  can  api)ly  to  the  Secretary 
to  the  Society,  who  will  give  him  an  opportunity  of 
perusing  and  copjdng  it,  or  if  he  resides  at  a  distance, 
so  much  as  he  requires  will,  upon  payment  of  the  cost 
of  reproducing  it,  be  forwarded  to  his  address.] 


ANNUAL     MEETING. 

Thursday,  March  lOth. 

The  President  (Professor  Kernot)  was  in  the  chair. 

The  minutes  of  the  last  meeting  were  read  and  confirmed. 

Election  of  Office-bearers  and  Members  of  Council. 

The  following  Office-bearers  and  Members  of  Council  were 
duly  elected: — President — Professor  W.  C.  Kernot,  M.A.,  C.E. 
Vice-Presidents— E.  J.  White,  F.R.A.S.,  and  H.  K.  Kusden, 
F.R.G.S.  Hon.  Treasurer— C.  R.  Blackett,  F.C.S.  Hon. 
Librarian — A.  Dendy,  D.Sc.  Hon.  Secretaries — Professor  W. 
Baldwin  Spencer,  M.A.,  and  A.  Sutherland,  M.A.  Members 
of  Council— J.  E.  Neild,  M.D.,  C.  A.  'J^opp,  M.A.,  LL.B., 
Pj-ofessor  Laurie,  LL.D.,  R.  L.  J.  Ellery,  F.R.S.,  G.  S.  Griffiths, 
F.R.G.S.,  Professor  Orme  Masson,  M.A.,  D.Sc,  H.  Moors. 
Rev.  E.  H.  Sugden,  B.A.,  B.Sc. 

The  President  referred  to  the  services  rendered  in  past 
years  to  the  Society  by  the  retiring  Librarian,  Dr.  Neild. 


Proceedings  of  the  Royal  Society  of  Victoria.      231 
Annual  Report. 

The  following  Report  and  Balance  Sheet  were  taken  as 
read,  and  on  the  motion  of  Mr.  Ellery,  they  were  adopted:— 

The  Council  of  the  Royal  Society  herewith  presents  to  the 
Members  of  the  Society  the  Annual  Report  and  Balance 
Sheet  for  the  year  1891.  The  following  meetings  were  held, 
and  papers  read  during  the  session  : — 

On  the  1 2th  March,  at  the  Ordinary  Meeting  held  after 
the  Annual  General  Meeting,  T.  S.  Hall,  M.A ,  "  On  a  New 
Species  of  Dictyonema  ;"  A.  Dendy,  D.Sc,  "  A  Preliminary 
Account  of  Synute  pidchella,  a  New  Genus  and  Species  of 
Calcareous  Sponge;"  T.  S.  Hall,  M.A..  and  G.  B.  Pritchard, 
",The  Geoloo-y  of  the  Southern  Portion  of  the  Moorabool 
Valley." 

On  the  2nd  April,  R  Etheridge,  Jun.,  F.G.S.,  and  A.  Smith 
Woodward,  "On  the  Occurrence  of  the  Genus  Belonostomus 
in  the  Rolling  Downs  Formation  of  Central  Queensland." 

On  the  nth  June,  Professor  W.  Baldwin  Spencer, 
"  On  the  Anatomy  of  Geratella  fmca ;"  A.  Dendy,  D.Sc, 
"Additional  Observations  on  the  Victorian  Land  Planarians;" 
A.  H.  S.  Lucas,  M.A.,  B.Sc,  "  On  a  New  Species  of  Fresh 
Water  Fish  from  Lake  Nigothoruk,  Mount  Wellington, 
Victoria;"  Professor  W.  Baldwin  Spencer,  "Land  Planarians 
from  Lord  Howe  Island." 

'  ■  On  the  9th  July,  A.  Dendy,  D.Sc,  "Description  of  a  New 
Species  of  Land  Nemertean  {Geonemertes  australiensis) ;" 
R.  L.  J.  Ellery,  F.R.S.,  "The  Present  State  of  the  Inter- 
national Photographic  Charting  of  the  Heavens." 

On  the  13th  August,  A.  Dendy,  D.Sc,  "On  the  Mode  of 
Reproduction  of  Feripattis  leuckartii ;"  A.  Dendy,  D.Sc, 
"  Short  Descriptions  of  New  Land  Planarians;"  Professor 
W.  C.  Kernot,  "  Notes  on  the  recent  Flood  on  the  Yarra." 

On  the  10th  September,  G.  B.  Pritchard,  "On  a  New 
Species  of  Graptolitidas  ; "  A.  Dendy,  D.Sc,  "  On  the 
Presence  of  Ciliated  Pits  in  Australian  Land  Planarians." 

On  the  8th  October,  A.  H.  S.  Lucas,  M.A.,  "  Notes  on  the 
Distribution  of  Victorian  Frogs;"  R.  L.  J.  Ellery,  F.R.S , 
"  Notes  on  the  Magnetic  Shoal  near  Bezout  Island,  North 
West  Australia." 


232      Pfoceedings  of  the  Royal  Society  of  Victoria. 

On  the  12th  November,  G.  S.  Giiffiths,  F.R.G.S.,  "The 
Geology  of  Barwon  Heads  ;"  A.  Dend_y,  D.Sc,  "  Description 
of  some  Land  Planarians  from  Queensland." 

On  the  lOtli  December,  Professor  W.  Baldwin  Spencer, 
"  Preliminary  Notice  of  Victorian  Earth-worms.  Part  I— 
'J'he  Genera  Megascolides  and  Cryptodrilus  ;"  H.  H. 
Anderson  and  J.  Shephaixl,  "  Notes  on  Victorian  Rotifers  ;" 
Professor  W.  Baldwin  Spencer,  "  Note  on  the  Ha.bits  of 
ijeradotus  forstei-i." 

The  following  Members  and  Associates  were  elected 
during  the  year : — Members,  A.  Dudley  Dol)son,  J.  W. 
Barrett,  M.D. ;  Country  Members,  John  Desmond,  John 
Dawson  ;  Associates,  W.  J.  Strettle,  W.  L.  Mullen,  M.D., 
Miss  Agnes  Ross  Murphy. 

Your  Council  regrets  to  ]ia,ve  to  record  the  loss  b}^  death 
of  the  following  Members  of  the  Society  : — Hon.  J.  G.  Beaney, 
J.  P.  Bear,  Henry  Edwards,  Johnson  Hick.';,  G.  LeFevre, 
M.D.,  John  Wall,  Hon.  Sir  Wm.  McLeay.  the  Right  Rev. 
Charles  Perry,  D.D. 

During  the  course  of  the  year,  your  Council  received  witli 
great  regret  the  resignation  of  A.  W.  Howitt.  Though  this 
was  nece-^sitated  by  pressure  of  official  duties,  it  is  hoped 
that  Mr.  Howitt  ma}'  before  long  tind  himself  again  able  to 
take  an  active  part  in  the  work  of  the  Society.  A.  Dendy, 
r).Sc.,  was  elected  by  the  Societ}'  to  fill  the  vacancy  thus 
created. 

During  the  course  of  the  year,  also,  J.  Cosmo  Newbery, 
B.Sc,  who  was  leaving  for  England  on  a  scientific  commission, 
resigned  the  Vice-Presidency  of  the  Society,  and  H.  K. 
Rusden  was  unanimously  elected  to  fill  the  vacant  office. 
From  1870-77,  and  again  Irom  1886-91,  Mi'.  Rusden  had 
been  closely  identified  with  the  Society  in  his  position  as 
one  of  the  Hon.  Secretaries,  and  in  nominating  him  for  the 
post  of  Vice-President,  the  Council  expressed  its  warm 
appreciation  of  the  services  which,  in  the  capacity  of 
Seci-etary,  he  had  rendered  to  the  Society,  in  the  work 
carried  on  by  which  he  had  taken  an  important  and  active 
share. 

The  Librarian  reports  the  addition  to  the  Library  during 
the  year  of  1076  publications.  It  may  be  noted  that  a 
Manuscript  Catalogue  of  the  Library  has  now  been  drawn 
up  rendering  it  more  available  for  reference.     The  Assistant 


Proceedings  of  the  Royal  Society  of  Victoria.     233 

Librarian  is  in  constant  attendance  to  afford  assistance  to 
those  desirous  of  consulting  the  volumes.  Your  Council 
has  had  under  consideration  the  exchanges  which  are  made 
with  other  Societies  and  hopes  to  be  able,  in  the  future,  to 
add  considerably  to  the  value  of  these  in  consequence  of  the 
increased  amount  of  publications  which  now  emanate  from 
the  Society. 

The  most  important  publication  of  the  year  has  been  that 
of  Part  I  of  Dr.  Dendy's  "  Monograph  of  the  Victorian 
Sponges." 

The  Committee  appointed  by  the  Council  to  distribute 
the  collections  of  animals  obtained  from  Port  Phillip 
experiences  very  great  difficult}'-  in  obtaining  any  informa- 
tion from  specialists  at  home  to  whom  the  various 
collections  have  been  sent  for  investigation.  It  hopes, 
however,  to  obtain  reports  during  the  coming  year,  and  is 
endeavouring  to  hasten  the  work,  thougYi  this  must 
naturally  take  a  long  period  of  time  in  completion.  The 
Council  desires  to  place  on  record  its  high  appreciation 
of  the  labours  of  J.  Bracebridge  Wilson,  Esq.,  M.A.,  by 
whom  the  collections  have  almost  entirely  been  made. 

As  will  be  seen  from  the  list  of  papers  read  before  the 
Society  during  the  past  year,  a  considerable  numlier  of 
original  scientific  investigations  have  been  carried  on  by 
Members.  Your  Council  trusts  that  the  amount  of  work 
recorded  in  its  publications  may  increase  year  by  year. 


234     Proceedings  oj  the  Royal  Society  of  Victoria. 


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Proceedimjs  of  the  Roijal  Society  of  Victoria.     235 


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•236     Proceedings  of  the  Royal  Society  of  Victoria. 

Professor  Spencee,  in  accordance  with  notice  of  motion, 
moved  the  repeal  of  Rules  52  to  58  inclusive.  He  did  not 
think  the  Society  was  large  enough  to  be  broken  up  into 
Sections,  and  further,  such  breaking  up  would  affect  the 
Society's  maintenance  and  welfare  as  a  whole.  In  his 
opinion,  Sections  could  not  be  carried  on  with  any  benefit  to 
the  Societ}'-,  as  a  number  of  their  member.s  interested  in  any 
{^articular  work  set  themselves  apart  and  formed  what  was 
practically  an  independent  Society,  and  the  Council  which 
must  be  the  central  authority  lost  control  over  the  actions  of 
members.  Their  experience  of  the  Sections  which  had 
already  been  in  existence  was  such  as  to  create  a  feeling 
against  their  continuance,  and  in  favour  of  merging  all  into 
one  Society  working  together  as  one  body. 

Mr.  Ellery  seconded  the  motion.  Until  he  hkd  had 
})ractical  experience  of  the  working  of  Sections  he  had 
favoured  their  establishment,  but  he  was  now  of  the  opinion 
that  they  tended  to  lessen  interest  in  tlie  doings  of  the 
parent  Society  and  to  reduce  the  attendance.  He  thought 
it  would  be  wise  to  abandon  the  Sections,  at  all  events  until 
the  Society  became  very  much  stronger.  The  Royal  Society 
of  London  had  no  Sections.  Should  the  motion  be  carried, 
they  sliould  not  interfere  with  existing  Sections  until  the 
lap.se  of  a  certain  period. 

Mr.  White  considered  that  if  the  Sections  had  been  kept 
under  control  no  trouble  would  have  been  (occasioned.  As 
the  Rules  provided  that  the  Council  "  may "  prevent  the 
formation  of  Sections,  there  was  no  need  for  the  resolution. 

Mr.  Blackett  referred  to  the  merging  of  the  Microscopical 
Society  into  the  Royal  Society,  and  probably  it  might  be  con- 
sidered by  members  of  the  former  body,  which  was  afterwards 
carried  on  as  a  Section  of  the  Royal  Society,  that  they  had 
been  somewhat  unfairl}^  dealt  with.  He  agreed  with  others 
that  the  Sections  had  been  somewhat  unsuccessful. 

Mr.  Sutherland  supported  Mr.  White's  view.  There  was 
no  necessity  for  the  motion,  the  object  of  which  was  merely 
to  get  rid  of  Section  G.  It  could  not  be  expected  that 
[teople  would  attend  the  ordinary  meetings  of  the  Society 
and  listen  to  some  abstruse  paper  on  a  subject  they  knew 
nothing  of,  while  they  waited  for  the  paper  to  be  read  in 
which  they  were  interested.  Royal  Societies  were  not 
favourable  fields  for  special  papers  in  any  particular  branch, 
although  many  people  were  good  enough  to  attend  meetings 


Proceedings  of  the   Rof/cd  Society  of  Victoria.      237 

and  suffer  the  infliction  of  hearing  papers  read  in  which  they 
had  not  the  slightest  interest,  and  so  provide  audiences  for 
the  readers.  If  it  was  intended  merely  to  get  rid  of  Section 
G,  then  Section  G  had  no  desire  to  remain  if  it  was  not 
wanted,  and  it  had  already  taken  steps  to  withdraw.  The 
Engineering  and  Physical  Sections  had  had  meetings  which 
were  rendered  more  uleasant  than  any  mixed  meetings  could 
be.  It  should  be  borne  in  mind  that  the  Sections  acted  as 
feeders  to  the  general  Society. 

Dr.  Dendy  considered  that  the  more  the  Sections  were 
increased  so  were  the  expenses.  The  Council  practically 
lost  all  control  over  the  Sections. 

Professor  Spencer  said  that  his  motion  was  general,  and 
did  not  applj^  to  any  particular  Section.  They  had  given 
Sections  a  fair  trial,  and  he  did  not  think  they  had  been  a 
real  success. 

Professor  Orme  Masson  said  that  if  the  motion  were 
carried,  it  did  not  follow  that  Section  G  could  not  remain. 
He  would  urge  that  the  motion  be  carried,  for  the  reason 
that  it  would  not  be  an  arbitrary  abolition  of  any  existing 
Section.  It  would  simply  be  a  certificate  from  the  Society 
that  it  did  not  care  to  create  new  Sections. 

Mr.  Ellery  drew  attention  to  Rule  54,  which  provided 
that  meetings  should  be  for  scientific  objects  only.  If  that 
Rule  had  been  kept  in  view,  all  difficulty  might  have  been 
avoided. 

Mr.  Jaeger  hoped  that  the  abolition  of  the  Rules  would 
not  preclude  the  reading  of  any  papers  on  Art. 

Mr.  Ellery. — Certainly  not.  The  formation  of  Section  G 
was  the  result  of  many  years'  discussion.  It  was  urged  that 
Art  and  Literature  should  be  moie  thought  of  in  the  Society 
than  they  were. 

Mr.  RuSDEN  thought  it  was  hard  to  suppose  that  the 
Rules  were  framed  without  consideration  and  wisdom. 
Rule  30  provided  against  anything  objectionable  occurring 
in  any  Section,  and  that  Rule  had  been  overlooked.  If  it 
had  not  been  overlooked,  no  paper  would  have  been  read 
without  being  submitted  to  the  Council. 

Mr.  White  agreed  with  Mr.  Ellery  that  the  Society  was 
not  strong  enough  for  Sections.  As  the  Council  had  the 
right  of  vetoing  Sections,  wh}^  not  leave  the  matter  to  it. 


238     Proceedings  of  the  Royal  Society  of  Victoria. 

The  President's  opinion  was  that  the  passing  of  the 
motion  would  not  abolish  existing  Sections,  but  would  take 
away  the  power  of  establishing  new  Sections  in  the  future. 

Professor   Orme    Masson   said    that  the    passing   of   the 

motion   would    show   that  the  Societ}-  was  against  having 

Sections,  and  the  result  would   in   all  probability  be  that 
Section  G  would  abolish  itself 

Mr.  Sutherland  said  that  Section  G  did  not  intend  to 
abolish  itself,   but  to  sheer  off  and  have  an  independent 

existence. 

Mr.  Blackett  mentioned  the  fact  that  no  minute  books 
had  been  kept  by  the  Sections. 

Mr.  White  moved  "  that  in  the  present  state  of  the 
Society  the  formation  of  Sections  was  not  advantageous  to 
it,  and  that  in  the  future  no  Sections  be  allowed  " 

Mr.  RusDEN. — The  Society  should  request  the  Council 
not  to  appoint  Sections. 

Dr.  Jamieson  was  opposed  to  the  formation  of  Sections* 
as  the  more  numerous  they  were,  the  more  they  tended  to 
impoverish  the  Society.  It  was  not  necessary,  in  his  opinion, 
to  alter  the  Rules,  as  in  accordance  with  the  sentiments 
expressed  that  evening  the  Sections  would  be  abolished. 

Professor  Laurie  could  not  understand  why  it  was 
necessary  to  legislate  for  the  future. 

Professor  Spencer's  motion,  on  being  put,  was  carried. 

Mr.  Albert  Swanson  and  Mr.  J.  B.  L.  Mackay  wei-e 
elected  as  Members. 

Mr.  G.  B.  Pritchard  and  Mr.  L.  J.  Balfour  were  nomi- 
nated as  Associates. 

The  Librarian's  Report  showed  that  203  publications  had 
l)een  added  to  the  Library  since  last  meeting. 

Professor  Spencer  gave  a  summary  of  his  paper  on 
"  Victorian  Earthworms.    Part  IL    The  genus  Perichseta." 

The  President  wished  to  know  if  Professor  Spencer 
agreed  with  Darwin's  theory  of  the  production  of  the 
humus. 

Professor  Spencer  thought  that  the  work  done  bj^  earth- 
worms in  other  countries  was  largely  done  by  ants  here. 


Proceedings  of  the  Royal  Society  of  Victoria.     239 

Thursday,  May  I2fh. 

The  President  (Professor  Kernot)  in  the  chair. 

The  minutes  of  the  last  meeting  were  read  and  confirmed. 

Mr,  Wilsmore,  Associate,  signed  the  Roll. 

Mr.  W.  H.  Archer  and  Dr.  J.  W.  Barrett  wei-e  elected  as 
new  Members  of  Council,  from  which  Di'.  J.  E.  Neild  had 
retired. 

Mr.  R.  J.  A.  Barnard,  M.A.,  and  Mr.  G.  B.  Pritchard,  were 
elected  as  Associates. 

The  Librarian's  Report  showed  that  191  new  publications 
had  been  added  to  the  Library  since  the  last  meeting. 

Mr.  Martin  Gardiner's  paper  on  "  Con  focal  Quadrics, 
&c.,"  was  taken  as  read. 

Dr.  Dendy  read  a  paper  entitled  "Further  Notes  on  the 
Oviparit}'  of  the  larger  Victorian  Peripatus,  generally  known 
as  P.  leuckartii." 

Mr.  Alexander  Sutherland  read  a  paper  on  "  Tiie 
Responsibility  of  Criminals." 

Mr.  Archer  said  that  they  had  heard  Mr.  Sutherland's 
very  able  and  comprehensive  address,  which  was  however 
full  of  contestable  points.  Mr.  Sutherland's  conclusion 
appeared  to  be,  that  while  it  was  a  gross  act  of  injustice  to 
]:)unish  a  man  for  what  he  could  not  help  doing,  yet  he  must 
be  punished  because  it  would  not  be  right  to  stick  to 
abstract  justice.  The  word  "  respon.sible "  was  used  a 
number  of  times  in  the  paper,  but  if  a  man  could  not  help 
himself,  how  could  he  be  held  to  be  responsible  ?  Although 
Mr.  Sutherland  had  stated  that  the  incorrigible  boy  at  school 
had  no  more  right  to  be  punished  than  a  Maori  should  be 
for  having  a  brown  skin,  yet  he  afterwards  said  that  the  boy 
ought  to  be  punished  because  it  would  assist  him  in  forming 
liis  character.  Did  not  that  indicate  on  the  part  of  the 
Aouth  a  sense  of  right  and  a  power  of  self-control  ?  As  to 
lunatics  in  asylums,  it  was  known  that  a  large  proportion 
had  what  were  called  lucid  intervals,  during  which  their 
medical  attendants  would  reason  with  them  and  treat  them 
as  intelligent  men.  That  was  a  proof  that  they  recognised 
in  their  patients  a  power  of  distinguishing  right  from  wrong, 
and  of  exercising  a  control  over  their  actions.  They  could 
determine  to  do  what  they  saw  was  right. 


240      Proceedingti  of  the  Royal  Society  of  Victoria. 

Piofessor  Laurie  coDsidered  there  were  some  points  in 
the  paper  which  must  inevitably  give  rise  to  difi'erence  of 
opinion,  and  he  was  rather  sorry  it  had  raised  the  question 
of  free-will.  Discarding  free-will,  and  holding  any  doctrine 
that  might  be  preferred,  it  was  not  necessary  to  go  to  the 
opposite  extreme  and  to  say  that  every  person  who  fell 
into  crime  was  to  be  regarded,  not  as  an  object  of  blame,  but 
solely  of  pity.  The  fact  was,  that  persons  who  did  wrong 
frequently  blamed  themselves,  and  admitted  that  they 
deserved  punishment.  That  showed  that  they  could  not 
wholly  do  away  with  the  sense  of  blame  for  wrong  done. 
Mr.  Sutherland's  great  point  should  not  be  lost  sight  of 
namely,  that  the  aim  of  government  in  inflicting  punish- 
ment was  to  deter  crime.  The  aim  of  government  should  be 
to  suppress  crime,  and  it  was  justified  in  doing  all  it  could 
to  attain  that  object.  Punishment,  or  the  tear  of  punish- 
ment, was  a  great  factor  in  suppressing  crime— in  preventing 
a  person  who  contemplated  committing  crime  from  falling 
into  it.  At  least,  that  should  be  the  aim  of  punishment. 
He  was  entirely  in  accord  with  Mr.  Sutherland,  that  govern- 
ment was  entitled  to  say  how  it  was  best  to  inflict  punish- 
ment, but  at  the  same  time  he  did  not  see  any  reason  why 
the  moral  aspect  of  crime,  even  from  the  deterrent  view, 
should  be  omitted.  It  was  quite  true  that  people  were 
swayed  by  fear  of  punishment,  but  when  the  State  laid 
down  certain  rules,  the  violation  of  which  would  lead  to 
punishment,  it  did  not  hold  out  a  threat  to  deter  people 
from  doing  what  they  would  otherwise  like  to  do.  To 
many  people,  the  State  was  an  embodied  conscience. 
People  did  not  come  into  the  world  knowing  what  was 
right  or  wrong,  nor  did  they  evolve  propositions  as  to  right 
or  wrong  from  their  peisonality  ;  they  were  taught  by 
others,  and  in  a  great  measure  by  the  laws  of  the  State,  as 
to  what  was  right  or  wrong.  A  great  many  peo]jle  regarded 
the  laws  which  the  State  had  laid  down  with  regard  to 
crime  as  laws  which,  from  a  moral  point  of  view,  should  not 
be  transgressed,  so  that  the  deterrent  powei'  of  the  State 
appeared  to  him  to  act,  not  only  from  the  fear  of  punish- 
ment, but  also  because  of  a  moral  standard  which  the 
majority  of  people  adopted,  and  which  they  were  not  likely 
to  transgress.  It  should  be  borne  in  mind,  that  the  punish- 
ment inflicted  by  the  State  for  crime,  so  far  from  reforming 
the  criminal,  had  the  opposite  eflect,  and  it  was  a  question 
worthv  of  consideration  whether  the  State  should  not  adopt 


PvoceediiKjs  of  the  Royal  Society  of  Victoria.      241 

some  form  of  restriction  adopted  to  the  various  needs  of 
those  with  v/hom  it  had  to  deal.  There  were,  undoubtedly, 
persons  in  whom  hereditary  traits  were  very  strong,  who 
were  borne  almost  irresistibly  towards  crime,  and  ibr  the 
security  of  society  the  State  might  deal  with  tliose  persoiis 
in  a  ditferent  way  from  that  in  whicli  others  were  treate*]. 
Still,  with  regard  to  a  great  number,  it  should  be  borne  in 
mind  that  tlie  humanitarian  spirit  of  the  age  demanded  that 
the  possibility  of  reformation  should  be  kept  in  view.  The 
trend  of  modern  thought  was  away  from  the  old  idea  of 
retribution  ;  but  he  thought  they  should  recognise  the 
elements  of  truth,  even  in  that  old  theory  of  retribution, 
and  the  elements  of  trutli  were,  that  when  something  was 
committed  wliich  was  regarded  as  criminal — acts  of  violence 
or  fraud — there  was  a  healthy  feeling  of  resentment.  He 
did  not  think  that  feeling  should  be  suppressed,  though  it 
should  be  woven  in  with  the  other  ideas  of  reformation. 
It  should  also  be  borne  in  mind  that,  when  men  were 
punished,  they  very  frequently  felt  that  they  deserved  it. 
While  he  thoroughly  agreed  with  Mr.  Sutherland  that  the 
State  was  perfectly  warranted,  for  th<^  suppression  of  crime 
and  the  protection  of  society,  in  adhering  to  any  punish- 
ments which  had  been  adopted  to  bring  about  this  aim,  he 
did  not  think,  on  consideration  of  the  whole  question,  that 
the  moral  aspect  could  be  entirely  left  out. 

Dr.  Jamikson  agreed  with  the  conclusion  arrived  at  in  the 
jmper,  that  yjunishment  must  be  inflicted  without  yielding 
to  any  particular  philosophical  opinions  one  might  hold 
about  free-will.  The  crux  of  the  question  related  to  capital 
punishment.  If  that  punishment  was  to  be  applied  mainly 
for  deterrent  purposes,  and  for  its  influence  on  others  for 
reformative  purposes,  was  it  not  absurdly  unffur  that  it 
should  be  inflicted  on  a  person  who  was  held  not  to  be  fairly 
responsible  for  what  he  did.  It  was  a  question  how  far 
capital  punishment  was  allowable  at  all  on  any  supposition 
of  necessitarianism.  A  School  of  Criminal  Anthropologists 
was  in  existence,  who  were  prepared  to  carry  out  their  ideas 
to  perfectly  logical  conclusions — especially  what  was  called 
the  Italian  School.  Lambrosa  went  so  far  as  to  lay  down  as 
-a  principle  that  there  are  instinctive  criminals,  and  that  it 
was  as  absurd  to  punish  such  a  person  as  it  would  be  to 
punish  a  person  for  having  small-pox  ;  that  there  existed  no 
right  in  any  sense  of  the  word  to  punish  such  a  person  for 
retributive  or  deterrent  purposes.     He  thought  the  common 

R 


242      Proceedings  of  the  Royal  Society  of  Victoria. 

sense  of  most  people  would  incline  tbeni  to  object  to  that 
view,  as  being  an  extreme  and  dangeious  doctrine.  It  was 
based  very  much  on  the  assertion  or  opinion,  that  the 
criminal  was  a  criminal  in  virtue  of  a  certain  defective  con- 
struction of  his  brain  ;  that  the  proof  of  that  consisted  in  the 
fact  that,  given  a  certain  number  of  persons  who  were  known 
to  be  criminals,  it  would  be  found  that  they  had  smaller 
heads  than  the  average  person  ;  that  they  had  a  less  facial 
angle  ;  that  the  top  part  of  their  face  and  head  sloped  back- 
wards ;  that  their  head,  on  the  average,  was  a  little  wider 
transversely — being  wide  relatively  to  its  length  ;  that  such 
])ersons,  on  the  average,  had  badly  shaped  ears  ;  that  the 
ridge  over  the  eyes  projected  ;  that  in  very  many  of  them 
there  was  a  secondary  ridge  above  the  eye-brows,  which  was 
very  prominent ;  that  in  a  considerable  number  of  them  the 
lower  jaw  was  largely  developed,  and  projected  in  front  of 
the  under  jaw.  Whilst  that  generalisation  might  be  freely 
admitted,  it  was  dangerous  to  apply  the  test  to  individuals. 
There  had  been  notorious  criminals  in  existence  wdiose  heads 
were  full  sized,  whose  facial  angle  was  good,  whose  ears  were 
good  and  whose  jaws  were  in  proper  position,  and  it  would 
be  altogether  unsafe  to  base  a  man's  responsibility  for  certain 
acts  he  had  committed  o!i  the  size  or  shape  of  his  head.  An 
attempt  was  undoubtedly  being  made  to  popularise  that 
doctrine  at  the  present  time,  and  to  say  that  given  a  man 
who  exhibited  a  certain  shape  of  head,  who  had  imperfectly 
formed  ears,  who  had  prominent  ridges  over  his  eyes,  and  so 
on,  and  given  the  further  fact  that  he  had  committed  a  crime, 
that  that  man  should  be  held  to  be  irresponsible,  and  should 
not  suffer  the  consequences  of  his  crime.  The  conclusion 
seems  to  be,  that  the  criminal  type  of  person  represented  a 
degenerated  type — a  development  of  the  lower  type,  with 
brains  constituted  similarly  to  those  of  the  lower  race,  and 
therefoi-e  .should  not  be  held  responsible  for  doing  any  wrong 
act.  He  thought  it  would  be  extremely  dangerous  to  allow 
a  doctrine  of  that  kind  to  be  carried  to  its  logical  conclusion. 
Popular  opinion  was  easily  led  astia}^  in  a  similar  matter, 
namely,  the  influence  on  the  brain  of  disease  or  of  injuries  to 
the  head,  and  the  probability  that  thereby  a  man  would  be 
rendered  irresponsible.  It  was  perfectly  true  that  disease  of 
the  brain  Avas  almost  certain  to  interfere  with  a  man's 
intellectual  capacity,  and  would  m.ake  him  incapable  ol 
controlling  himself  That  general  principle  might  be  admit- 
ted ;  but  it  was  dangerous  and  absurd  to  infer  that,  when  a 


ProGeedings  of  the  Royal  Societij  of  Victoria.     243 

man  was  suffering  from  any  kind  of  disease,  he  was  t(3  be  held 
to  be  irresponsible.  It  was  known,  that  there  were  structural 
diseases  of  the  brain  that  did  not  necessarily  involve  the 
mental  faculties.  That  the  mental  faculties  were  affected  to 
a  considerable  extent  was  well  known,  but  it  was  absurd  to 
infer  iri'esponsibility,  and  to  declare  that  a  man  wjis  not  to 
be  punished  for  his  wrongful  acts.  Especially  did  it  become 
dangerous  and  absurd  when  it  was  attempted  to  apply  the 
doctrine  to  individuals.  It  was  extremely  dangerous  to  apply 
the  doctrine  of  anthropology  to  criminal  cases.  In  spite  of 
i-ecent  discoveries,  we  did  not  know  half  as  much  as  we 
would  like  to  know  about  the  functions  of  the  brain,  or  as 
much  as  we  hope  some  day  to  know.  Physiologists  know 
that  there  are  parts  of  the  brain  that  are  well  defined,  that 
there  are  parts  that  control  movements,  that  there  are  parts 
which  deal  with  hearing,  speech,  and  sight,  but  when  it 
came  to  the  question  of  learning  what  particular  part  of  the 
brain  is  exclusively  concerned  with  the  mental  faculties, 
they  would  have  to  depend  on  what  was  little  more  than 
inference.  It  was  probable  that  the  front  part  of  the  brain 
is  that  which  is  concerned  with  the  intellectual  and  moral 
faculties.  There  might  be  extensive  injury  to  that  front 
part  of  the  brain — parts  of  it  might  be  lost  or  degener- 
ated as  the  result  of  disease — but  he  did  not  know  that  it 
always  followed  that  the  mental  or  moral  faculties  were 
deterioiaied  in  proportion  to  the  amount  of  brain  lost.  The 
average  unscientific  person  was  apt  to  apply  to  individual 
cases,  appai-ently  logically,  but  still  dangerously  and  unfairly, 
the  scientific  generalisation,  which  was  fair  enough  as  regards 
a  particular  disease.  As  to  the  measure  of  responsibility  of 
people  unmistakably  insane,  that  was  a  difficult  question, 
with  which  he  was  not  then  prepared  to  deal. 

Dr.  Mullen  said  it  was  absolutely  neces.sary  that  lawyers 
should  act  on  precedents.  It  was  far  more  important  that 
the  law  should  be  as  certain  as  it  conld  be,  than  that  it 
should  be  just.  There  were  many  points  in  it  that  were 
unjust,  but  the  prerogative  of  mercy  conld  be  exercised  by 
the  Crown.  Of  course  the  law  was  the  true  embodiment  of 
everything  that  was  excellent,  and  it  did  not  deal  with 
motives.  Although  the  motive  for  an  act  might  influence 
the  amount  of  pvuiishment  inflicted,  it  could  not  influence 
the  responsibility  of  the  ])erson  who  had  done  wrong.  The 
responsibility  of  criminals  was  mixed  up  with  the  most 
important  question  of  the  protection  of  society.     If  a  man 

11  2 


244     Proceedhu/s  of  the  Royal  Society  of  Victoria. 

had  stolen  a  £5  note,  he  would  join  in  the  cry  against  the. 
wrongdoer,  because  others  might  steal  a  £5  note  from  hiii). 
He  unhesitatingly  said  that  the  modei'n  idea  of  the  i-esponsi- 
bilit}^  of  criminals,  and  consequent  punishment,  was  not  the 
best  wa}^  to  protect  society.  In  the  beginning  of  this 
century  nearly'  all  tlie  felons  were  hanged,  and  th*; 
adventurous  spirits  were  drafted  off  to  the  wtirs.  The 
consequence  was,  to  use  a  vulgar  expression,  that  the  breed 
was  stopped.  Now,  however,  owing  to  the  manner  in 
which  felons  were  treated  (imprisonment  for  a  short  period), 
there  was  being  developed  a  race  as  devoid  of  moral  sens*' 
as  a  child  born  blind  was  of  sight ;  it  was  nothing  less  than 
propagation  of  the  species.  We  were  developing  a  typical 
immoral  race ;  and  if  anyone  were  to  go  to  the  East  end  of 
London  and  pick  out  ten  children  of  from  8  to  JO  years  of 
age,  he  could  predict  with  tolerable  certainty"  what  would 
become  of  nine  of  them,  partly  by  heredity  and  partly  by 
surroundings.  The  question  to  be  considered  was  not  so 
nmch  their  responsibility,  because  the  law  made  them 
responsible,  but  how  were  they  going  to  protect  society 
from  criminals.  He  would  not  enter  into  the  discussion  of 
such  questions  as  free-will,  because  they  were  not  practical. 
As  to  the  responsibility  of  the  insane,  that  was  a  question 
to  which  it  was  impossible  to  give  an  answer.  He  might  as 
well  ask  the  President  if  a  bridge  over  the  Yaria  were 
injured,  what  strain  it  would  bear.  The  President  would 
want  to  know  what  bridge  it  was,  the  particular  kind,  the 
size  and  other  things  which  to  an  ordinary  layman  would 
look  absurd,  and  experts  would  differ  as  to  the  amount  ot 
strain.  He  might  as  well  ask  Mr.  Sutherland  how  long  it 
would  take  for  a  boy  to  construe  a  chapter  of  Csesar.  He 
would  want  to  know  all  about  the  boy  and  his  class,  and 
what  chapter  was  referred  to.  When  they  came  to  sjjeak 
of  lunatics,  each  case  must  be  considered  singly.  The  law- 
did  not  recognise  that  there  were  dozens  of  forms  of 
insanity  which  ran  into  one  another.  Originally,  there 
was  but  one  class  of  insanity.  Then  the  law  took  note  of 
delusions,  but  it  stopped  there  ;  a  mad  man  was  either  a 
maniac  or  delusionally  insane.  Those  laws  were  laid  down 
by  very  worthy  gentlemen,  who  knew  nothing  but  what 
they  found  by  metaphysical  reasoning  in  the  four  corners 
of  their  rooms.  Mr.  Sutherland  had  laid  down  rules  f()r  the 
guidance  of  lunatic  asylums ;  the  Act  of  Parliament  did 
not    allow    the    medical    officers    to    punish    any    lunatic. 


F roceediiujs  of  the  Royal  Society  of  Victoria.      24o 

Lunatics,  knowing  that  they  could  not  ba  punished,  did 
not  commit  crimes.  He  knew  the  case  of  one  inmate  who 
would  say  to  the  doctor  at  times,  "if  you  don't  lock  me  up, 
I  will  do  sometlring."  The  older  warders  would  say,  "I  think 
you  had  better  lock  him  up."  Such  men  knew  when  the 
impulse  to  do  some  violent  act  was  upon  them,  and  knew 
it  could  not  be  resisted.  Then  in  puerperal  mania,  he  had 
known  a  woman  to  say,  "  doctor,  if  3'ou  don't  take  the  child 
away,  1  will  kill  it ;  if  the  servant  goes  out,  I  will  kill  it." 
The  woman  was  perfectly  aware  of  the  illegality  of 
the  act,  but  could  not  help  herself,  but  even  the  presence 
of  the  servant  girl  would  deter  her.  That  was  a  strong 
case,  as  it  appealed  to  the  sentiments.  In  ordei-  to  show 
that  the  punishment  of  a  criminal,  sane  or  insane, 
would  not  deter  others,  he  referred  to  the  case  of  a 
woman  in  Brunswick  who,  on  the  same  night  that  Deeming 
was  condemned  to  death,  deliberately,  as  far  as  a  lunatic 
could  act  deliberately,  went  into  a  house  and  exploded 
dynamite,  and  when  she  was  about  to  be  arrested  she  tried 
to  blow  herself  up.  The  lawyers  would  say  she  was  partially 
insane,  whatever  that  might  mean  ;  she  certainly  had  a 
sufficiency  of  reason  to  know  right  from  wrong,  but  the 
})unishment  of  the  other  criminal  had  no  deterrent  effect  on 
her.  Mr.  Paisden  had  alluded  to  the  fact,  tliat  lunatics  and 
criminals  received  the  same  punishment,  viz.,  incarceration. 
He  supposed  there  was  no  sane  man  living  who  would  care 
tA)  Ije  locked  up  as  a.  patient  in  an  asylum,  but  it  was  certain 
that  that  punishment  had  never  prevented  a  man  from 
going  insane.  As  to  the  punishment  of  lunatics,  some  were 
responsible  and  some  were  not,  but  that  was  a  matter  on 
which,  as  to  justice,  they  should  go  to  experts.  As  to  the 
protection  of  Society,  he  would  say  they  were  responsible. 
In  the  present  state  of  the  law,  the  judges  get  out  of  the 
difficulty  as  best  they  can.  In  a  case  of  puerperal  mania,  in 
which  the  patient  admitted  to  the  doctor  that  she  knew  she 
was  doing  wrong,  the  judge  said  to  the  jury — "  Gentlemen, 
she  said  she  knew  she  wa,s  doing  wrong,  but  it  is  for  you  to 
say  whether  she  meant  what  she  was  saying."  That  was 
the  way  in  which  he  got  out  of  the  difficulty.  The  question 
of  sudden  impulse  is  a  dangerous  one  to  deal  with.  The 
question  ought  to  be — "Has  this  man  got  brain  disease; 
iias  he  got  a  certain  kind  of  brain  disease  affecting  certain 
functions  1  Yes."  The  lawyers  did  not  take  it  that  way. 
They  seized  upon  the  metaphysical  point  of  right  or  wrong. 


246     Froceedings  of  the  Royal  Society  of  Victoria. 

The  law  said  that,  if  the  man  had  a  sufficient  idea  of  right 
or  wrong,  he  was  responsible.  The  Executive  sometimes 
stepped  in  and,  as  he  thought,  wiongly  pardoned  the  man 
who  was  a  danger  to  society.  In  one  of  our  asylums  there 
was  a  terrible  case  of  homicidal  mania,  and  in  his  opinion, 
that  man  should  be  removed  simply  for  the  protection  ot 
society. 

Mr.  RusDEN  called  attention  to  an  article  in  the  Forum, 
on  the  Elmira  Prison  in  America.  The  conclusion  it  drew 
was  that,  wliile  the  population  was  increasing  three  per  cent, 
the  criminality  was  increasing  fift}'  per  cent.,  and  the  writer 
attributed  that  result  to  the  humanitarian  treatment  of 
criminals. 

After  a  few  words  in  reply  by  Mr.  Sutherland,  on  the 
motion  of  C'olonel  Goldstein,  seconded  by  Professor  Orme 
Masson,  the  discussion  v^as  adjourned  to  the  next  meeting. 


Thursday,  June  9th. 

The  President  (Professor  Kernot)  occupied  the  chair. 

The  minutes  of  the  preceding  meeting  were  I'ead  and 
confirmed. 

Mr.  J.  B.  Pritchard,  an  Associate,  signed  the  Roll  and  was 
introduced  to  the  meeting. 

The  Librarian  reported  that  1 14  publications  had  been 
received  since  the  last  meeting. 

Mr.  Sutherland  read  a  paper  "  On  the  Nest  and  Eggs  of 
the  Victoria  Rifle  Bird "  (Ptilorhis  victorise),  by  Mr.  D. 
Le  Souef 

The  discussion  on  "  The  Responsibilty  of  Criminals  "  was 
resumed. 

Colonel  Goldstein  said  that,  in  continuing  the  discussion 
of  M)\  Alex.  Sutherland's  paper  on  "  The  Resjjonsibilit}'^  of 
C'riminals"  i-ead  at  ti)e  last  meeting,  it  would  be  necessary 
to  state  a  few  of  the  difficulties  that  occurred  to  the  unscien- 
tific public  mind,  which  tended  to  prevent  the  formation  of 
just  opinions  on  the  subject.  One  difficulty  was  the  great 
divergence  ajjparent  between  tlie  views  advanced  by  those 


Proceedings  of  the  Royal  Society  of  Victoria.     247 

scientific  and  professional  gentlemen  to  whom  the  public 
looked  for  advice.  Qui  legal  luminaries  held  the  view 
that  all  men  were  responsible  for  their  crimes,  except 
in  the  case  of  acute  mania,  when  it  coiild  be  proved 
that  the  accused  did  not  know  the  difference  between 
right  and  wrong ;  while  the  medical  fratei-nity,  though 
by  no  means  unanimously,  urged  irresponsibility  when- 
ever there  was  any  form  of  brain  disease,  no  matter 
how  slight,  or  even  where  it  was  only  suspected.  It 
was  as  well,  perhaps,  for  the  general  well-being,  that 
society  was  still  willing  to  accept  the  guidance  of 
our  legal,  rather  than  of  our  medical,  friends.  But  as 
theories  that  were  flung  broadcast  among  the  people  were 
certain  to  obtain  some  adherents,  and  as  such  theories, 
if  largely  appealing  to  our  sj-mpathies,  might  lead  to 
dangerous  changes  in  public  opinion,  it  was  well  that 
scientific  societies  should  meet  and  discuss  all  such  theories, 
for  the  benefit  of  the  public,  and  in  order  that  our  laws 
might  be  wisely  ordained;  law  being  the  ciystalHsation  of 
public  opinion.  Another  difliculty  arose  from  contusion  in 
the  terms  used,  and  their  true  significance.  For  instance, 
"retributive  punishment"  was  frequently  described  as 
"  revenge,"  which  of  course  was  quite  incorrect ;  yet  many 
writers,  Mr.  Sutherland  also,  used  the  phrase  in  this  sense. 
Then  there  was  the  extremely  narrow  view  advanced  by 
some  writei's  that,  earl}'  in  the  world's  history,  punishment 
was  retributive  only  ;  that  later,  it  was  sought  to  be  made 
deterrent;  and  that  as  it  had  failed  to  be  deterrent,  it  should 
therefore  seek  to  be  reformatory  only.  He  alluded  to  this 
as  a  naiTow  view,  because  he  hoped  to  indicate  presently 
that  punishment  should  partake  of  all  three  qualities — 
that  it  should  be  retributive,  deterrent  and  reformatory. 
There  was  another  difficulty  that  must  frequently  occui 
to  the  unscientific  mind,  how  to  reconcile  such  statements 
as  Mr.  Sutherland's,  that  "the  treatment  of  criminals  was 
not  a  matter  of  abstract  justice,  but  of  pure  policy.  It  was 
not  concerned  with  ethics,  but  with  the  preservation  of  law 
and  order."  S])encer  laid  down  as  the  fundamental  law  of 
human  justice,  "that  each  individual  ought  to  receive  the 
benefits  and  the  evils  of  his  own  nature  and  consequent 
conduct ;  neither  being  prevented  from  having  whatever 
good  his  actions  normally  bring  to  him,  nor  allowed  to 
shoulder  off  on  to  other  persons  whatever  ill  was  brought 
to    him    by    his    actions."       The    question    of    the    respon- 


24S     Froceedinr/s  of  the  Royal  Society  of   Victoria. 

aibility  of  the  criminal  was  a  large  one,  and  was  to  be 
looked  at  critically  from  so  many  points  of  view,  that 
it  was  only  by  long  and  patient  stud}'  we  could  hope 
to  solve  it,  and  to  measure  out  punishment  to  the 
criminal,  with  the  view  to  the  gradual  reduction  of 
crime.  So  much  was  this  felt,  that  it  had  justlj'  been 
considered  necessary  to  examine  the  criminal  class  in  a 
thoi-oughly  scientific  manner.  Hence  the  new  Science  of 
"  Criminal  Anthropology,"  which  had  led  to  the  holding  of 
International  Congresses,  the  first  of  which  had  been 
held  at  Berne  in  1885,  the  second  in  1889  at  Paris, 
and  the  third  was  to  be  held  this  year  at  Brussels. 
There  had  also  been  established  a  few  years  back  the 
International  Criminological  Association.  An  enormous 
amount  of  useful  work  had  been  done,  but  the  new  Science 
was  only  in  its  infancy.  INumerous  articles  and  books  on 
this  subject  had  been  [)ublished  in  Europe  and  America,  so 
that  the  tirsfc  results  of  enquiry  were  within  reach.  Have- 
lock  Ellis,  in  1890,  had  published  a  book  called  "The 
Criminal,"  which  was  a  valuable  compilation  of  the  opinion 
of  those  who  had  taken  a  lending  part  in  the  work.  His 
(Colonel  Goldstein's)  attention  had  been  directed  to  three 
articles  in  the  "  International  Journal  of  Ethics,"  which 
fairly  represented  some  of  the  views  lield.  One  was  "  The 
Theory  of  Punishment,"  by  the  Kev.  Hastings  Rashdall ; 
another  "  The  Prevention  (jf  Crime,"  by  Dr.  Tonnies  ;  and 
the  third  was  a  discussion  on  these  by  Professor  James 
Seth,  of  Dalhousie  College,  According  to  Professor  Seth, 
the  new  Science  of  Criminology  was  founded  on  the 
theory  that  crime  was  a  pathological  phenomenon,  and 
that  the  proper  treatment  of  the  criminal  was,  accord- 
ingly, that  which  sought  his  cure  rather  than  his 
punishment.  He  claimed  that  this  was  an  advance  in 
iiumau  ibeling  as  well  as  in  intelligence.  It  might  be 
suggested  ti)at,  as  these  latter  day  views  of  criminology 
wei'e  the  result  of  special  studies,  or  studies  by  specialists, 
we  should  do  well  to  raise  the  question,  were  specialists,  as 
a  I'ule.  well  trained  in  philosophy.  Or,  to  put  it  more 
distinctl}-,  had  these  particular  specialists  any  fair  amount 
of  knowledge  of  the  ultimate  causes  of  the  various 
phenomena  of  the  universe  ?  Because  so  far  as  specialists 
were  deficient  in  general  philosophy,  so  far  must  we  guard 
agaiu.-it  being  led  to  avoid  generalising  on  the  I'e.sults  of 
their    undoubtedly     valuable    accumulations    of    evidence. 


Proceedinc/s  of  the  Royal  Society  of  Victoria.     24^ 

The  Kev.  Hastings  Raslulall  objected  to  the  retributive 
theory  of  punishment,  and  expected  that  with  the  necessary 
nioraiization  of  communities,  the  sphere  of  criminal  law 
ought  gradually  to  extend  ;  while  Dr.  Ferdinand  Tonnies, 
of  Kiel  University,  asserted  that  all  punishment  as  punish- 
ment should  cease,  though  he  had  no  better  substitute  to 
offer  than  a  system  of  fines  ;  while  he  looked  to  further 
research  foi-  other  means  of  preventing  crime  than  could  be 
found  in  the  threatened  or  real  consequences  of  what  the 
criminal  had  done.  Professor  Seth,  in  discussing  both  these 
opinions,  raised  the  question  whether  the  newer  and  older 
views  of  punishment  were  mutually  exclusive,  and  if  not, 
what  was  their  relation  to  one  another,  and  seemed  to 
favour  the  idea  that  punishment  must  be  reformatory  only. 
He  asserted  "  that  society  was  now  so  securely  oi'ganised, 
that  it  could  afford  to  be  not  only  just,  but  generous  as 
well."  Would  not  this  be  rather  dangerous  pleading  if 
adopted  in  our  Law  Courts?  Most  thinking  people 
would  agree  with  Professor  Seth  in  this  adverse  criticism 
of  the  diverse  views  advanced  by  Mr.  Rashdall  and 
Dr.  Tonnies,  and  also  when  he  much  qualified  the 
idea  that  crime  was  a  "  pathological  phenomenon,"  by 
urging  that  ''it  wns  only  an  analogy  or  metaphor 
after  all,  and  like  all  metaphors,  might  easil}'  prove 
misleading  if  taken  as  a  literal  description  of  the  facts;" 
that  "to  resolve  all  badness  into  insanit}^  did  not  conduce  to 
clear  thinking,"  and  that  "  normal  crime,  if  it  had  anything 
to  do  with  insanity,  was  rather  a  cause  than  a  result." 
He  said  that  "  To  reduce  crime  to  a  '  pathological  phenome- 
non '  was  to  sap  the  very  foundation  of  our  moral  judgment, 
merit  as  well  as  demerit,  reward  and  punishment,  were 
thereby  undermined.  Such  a  view  might  be  scientific,  it 
was  not  ethical,  for  it  refused  to  recognise  the  commonest 
moral  distinctions."  One  of  the  articles  of  the  Inter- 
national Criminological  Association,  quoted  by  Dr.  Tonnies, 
said,  "  Punishment  was  an  act  of  justice,  and  the  essence 
of  punishment  was  retribution.  From  this  standpoint, 
satisfaction  was  the  primary  object  of  punishment, 
and  the  other  objects  included  reformation  and  deter- 
rence." We  might  justly  take  this  as  a  fair  statement 
of  the  facts.  While  we  acknowledged  that  retribution 
could  only  be  made  in  trivial  crimes  against  the 
property  or  person,  when  crimes  became  more  serious, 
retribution  became   more   or  less  impossible.      Punishment 


250     Proceedings  of  the  Royal  SocieUj  of  Victoria. 

then  must  be  deterrent,  while  all  punishment  should  be 
inflicted  with  a  view  to  the  ultimate  reform  of  the 
criminal,  bearing  in  mind  that  in  order  to  secure  refor- 
mation tlie  conscience  of  the  criminal  must  be  awakened. 
He  must  be  brought  to  see  that  his  punishment  is 
just  before  we  could  hope  for  any  betterment.  In 
"  The  Criminal,"  by  Havelock  Ellis,  one  could  not  fail 
to  be  struck  with  the  mass  of  evidence  gathered  in  the 
biological  and  pathological  examination  of  the  criminal. 
But  it  must  be  observed  that  a  fair  examination  of  the 
evidence  led  to  the  opinion  that  the  bulk  of  what  were 
called  criminal  physiological  characteristics  were  also  to  be 
met  with  in  the  non-criminal  and  respectable  classes,  while 
many  of  the  so-called  criminal  characteristics  were  due  to 
the  professional  exercise  of  crime.  Lombrosa  had  been 
often  cited  as  the  greatest  living  exponent  of  criminal 
anthropology,  yet  he  was  called  rash  and  unscientific.  Ellis 
spoke  of  his  work  as  "  by  no  means  free  from  faults.  His 
style  was  abnipt ;  he  was  too  impetuous,  arriving  too 
quickly  at  conclusions,  lacking  in  critical  faculty  and  in 
balance.  Thus  at  an  early  date  he  was  led  to  over-estimate 
the  atavistic  element  in  the  criminal,  and  at  a  later  date  he 
has  pressed  too  strongly  the  epileptic  affinities  of  crime." 
Yet  this  was  the  authority  who  was  often  quoted,  especially 
by  medical  witnesses  who  advocated  irresponsibility  of  the 
criminal.  Of  all  his  vast  mass  of  in vestigiition,  extending  to 
about  3(),()()0  cases,  Lombrosa  himself  had  declared  that 
"  perhaps  not  one  stone  would  remain  upon  another,  but 
that,  if  this  was  to  be  the  fate  of  his  work,  a  better 
edifice  would  arise  in  its  place."  To  illustrate  the 
length  to  which  enthusiastic  specialists  w^ould  go  in 
advocating  their  own  views,  l)espine,  who  wrote  a 
good  work  in  1868,  "  Psychologic  Naturelle,"  had  con- 
sidered the  criminal  as  "  morally  mad,"  and  therefore 
irresponsible,  and  had  said,  "  No  physiologist  had  yet 
occupied  himself  with  the  insanity  of  the  sane."  Was 
not  this  evidence  of  the  "illusions  of  enthusiasm?"  We 
might  treat  the  question  of  responsibility  from  an  ethical, 
a  metaphysical,  a  clinical,  or  a  practical  point  of  view, 
and  we  must  arrive  at  the  conclusion  that  sane  or 
insane,  eveiy  criminal  must,  for  the  protection  of  society, 
be  treated  as  responsible.  He  had  been  asked  to  say 
something  on  the  treatment  of  the  criminal,  but  he 
thought    that     hardly    came    within     the    scope     of     the 


Fyuceedings  of  the  Royal  Society  of  Victoria.     251 

present  enquiry.  It  would  be  sufficient  to  say  that  tb(^ 
trend  of  modern  thought  was  in  the  direction  of  abolish- 
ing fixed  limits  to  sentences  ;  that  our  prisons  should 
be  made  into  Reformatories,  where  every  hour  would  be 
profitab]}'  occupied,  and  that  when  prisoners  were  reported 
fit,  they  should  be  allowed  out  on  parole.  Also,  that 
the  surest  way  to  check  tbe  increase  of  the  criminal 
class,  is  to  remove  criminal  children  to  a  healthy  environ- 
ment. 

Mr.  RuSDEN  remarked  that  Colonel  Goldstein  had  omitted 
to  mention  the  system  of  indeterminate  sentences,  which  he 
believed  to  be  one  which  would  soon  be  adopted.  Mr. 
Havelock  Ellis  mentioned  it  as  having  been  introduced  into 
America  some  time  ago.  To  Mr.  Frederick  Hill  belonged 
the  honour  of  first  suggesting  this  fruitful  reform.  Lunatics 
were  dealt  with  on  this  principle.  A  lunatic  was  not 
liberated  until  two  medical  gentlemen  certified  that  the 
treatment  to  which  lie  had  been  subjected  had  been 
successful,  and  that  he  was  fit  to  be  released.  At  present 
tlie  law  fixed  a  maximum,  and  in  some  cases,  a  minimum 
sentence,  and  the  criminal  was  released  very  much  the  worse 
for  his  imprisonment  after  a  shoi't  definite  period,  fixed 
without  regard  to  reason.  If  the  system  of  indeterminate 
sentences  were  given  a  fair  trial,  he  thought  it  would  be 
found  much  more  satisfactory  than  that  at  present  in  vogue. 
He  did  not  believe  it  was  possible  to  reform  a  man  who  had 
grown  accustomed  to  commit  crimes,  but  with  first  offenders 
this  system  might  be  veiy  successful. 

Mr.  Alexander  Sutherland  said  that  whilst  there  was 
little  to  cavil  at  in  Colonel  Goldstein's  i)aper,  there  was  one 
point  as  to  which  he  thought  he  detected  an  uncertain 
sound — viz.,  with  regard  to  the  nature  of  punishment. 
Colonel  Goldstein  had  said  that  punishment  might  either 
be  retributive,  deterrent,  or  reformatory.  If  punishment 
were  reformatory,  it  ceased  to  be  punishment  at  all.  One 
could  not  logically  speak  of  reformatory  punishment.  In 
that  case,  it  was  simply  a  mode  of  treating  criminals  whicli 
was  reformatory.  He  agreed  with  Mr.  Rusden  that,  if  a 
man  were  allowed  to  grow  up  a  criminal,  he  could  not  be 
reformed.  The  leading  authorities  were  agieed  on  that 
point.  Bej'ond  the  age  of  10  or  12,  the  chance  of  reform 
was  apparently  slight.  If  a  man  lived  up  to  the  age  of  20 
as  a  criminal,  nothing  practically  would  reform  him.     Not 


2'r2     Proceedings  of  the  Royal  Socleti/  of  Victoria. 

only  vvei-e  there  ^ood  authorities  for  that  statement  amongst 
the  leading  writers  in  England,  but  it  could  be  shown  that, 
even  in  this  colony,  such  was  the  fact.  Lord  John  Russell 
had  inaugurated  the  Penfold  scheme  —a  reformatory  schema 
in  whi(;h  criminals,  when  improved  to  a  certain  extent,  were 
to  be  sent  to  Australia  with  so  much  money  in  their  pockets. 
Two  shiploads  of  criminals,  certificated  as  having  been 
reformed,  had  been  sent  to  Australia  about  1849  or  1850, 
Two-thirds  of  the  men  who  had  entered  the  scheme  never 
reached  Australia,  and  those  that  were  sent  were  the  best. 
These  men  had  not  improved  the  population ;  but  our 
records  of  crime  showed  that,  instead  of  being  reformed, 
when  they  had  got  a  sum  of  money  in  their  pockets,  and 
were  landed  on  a  new  shoi-e  to  start  a  new  career,  they  had 
turned  out,  as  a  rule,  misei'able  failures.  But  it  was  fair  to 
remember,  that  the  ranks  of  crime  included  many  characters. 
There  was  the  criminal  who  was  of  an  energetic  character, 
and  whose  energies  had  been  directed  into  an  unfortunate 
channel.  There  was  the  man  who  had  mutinied  in  the 
Army,  or  the  man  who  had  merely  knocke.i  a  hare  over 
which  happened  to  run  in  front  of  him.  Then  again,  there 
was  the  man  whose  daughter  had  been  ruined  by  some 
wealthy  man,  and  who  had  avenged  himself  These  were 
not  criminals  in  the  proper  sense  of  the  term.  They  were 
on  a  different  footing,  and  might  have  a  fan^  chance  in  a 
new  countiy,  where  that  very  energy  and  impetuosity  that 
had  carried  them  into  a  wrong  grove  in  one  dii-ection,  might 
make  them  most  successful  in  another.  Many  of  the  world's 
gi-eatest  men  would  have  been  great  nuisances  if  they  had 
taken  a  wrong  ti-ack  ;  Lord  Clive  was  an  instance.  Marl- 
borough, too,  whose  energy  would  perhaps  have  been 
thrown  into  a  wrong  channel  if  his  country  had  not 
needed  his  services,  had  found  an  outlet  for  it  in  slaughter- 
ing Fienchmen,  and  so  had  become  a  hero.  Passing 
from  the  question  of  reformation,  there  remained  the 
theories  of  retribution  and  deterrence.  No  philo.sophical 
people  would  hold  that  punishment  should  be  retributive — ■ 
that  if  a  man  were  struck,  he  should  be  resolved  to  return 
the  blow,  merely  as  a  matter  of  retribution,  although 
he  would  be  perfectly  entitled  to  take  precaution 
to  prevent  a  repetition  of  the  act.  Retribution  was 
not  according  to  modern  views,  and  there  only  i-emained 
the  deterrent  view,  which  should  be  widened  out  into 
a  question  of  placing  a  sufficiently  strong  deterrent  motive 


Proceedings  of  the  Roijal  Society  of  Victoria.     25o 

in  the  balance  of  the  motives  tliat  actuated  a  nian  to 
ensure  his  actions  talking  a  right  direction.  There 
were  three  great  objections  to  the  view  that  criminals 
should  be  eliminated  by  simply  putting  them  to  death  : 
— First,  there  was  the  practical  one,  that  the  feelin<4 
of  the  race  was  against  it.  The  second  objection  was,  that 
the  process  would  have  to  be  repeated  time  after  time.  ]i' 
the  least  desirable  people  were  singled  out  at  the  present 
moment  and  got  rid  of,  although  the  remainder  of  the 
population  would  be  improved,  but  in  twenty  or  thirty  years 
thei'e  wcnild  be  just  as  much  difference  between  the  then 
respectable  classes  and  the  lower  classes  as  there  was  now. 
and  these  would  have  to  be  exterminated.  In  a  couple  of 
centuries,  people  of  the  character  of  the  judges,  who  sat 
upon  the  bench  at  the  present  moment,  would  be  chloro- 
formed as  being  objectionable  people.  Then  again,  was 
everybody  who  broke  the  laws  to  be  chloroibrmed  ?  At 
present  penalties  were  awarded  on  a  graded  scale,  and  there- 
were  felonies,  misdemeanors,  (fcc.  The  weeding  out  process 
would  require  an  arbitraiy  scale.  Would  they  let  off  first 
offenders,  or  would  they  draw  the  line  at  the  second  offence  ? 
However  it  was  arranged,  such  violent  and  arbitrary  lines 
must  appear  objectionable  to  the  public  conscience. 
The  third  objection  was  that  nature  had  arranged  the 
matter  in  her  own  way  in  a  far  more  efficacious  style. 
Although  the  genus  criminal  seemed  to  occur  in  a  sporadic- 
way,  it  really  obeyed  certain  laws.  To  make  his  meaning- 
plain,  he  would  draw  their  attention  to  the  extensive  area 
from  which  we  inherit  our  natures.  Everybody  had  two 
parents,  and  four  grandparents,  and  eight  great  grandparents, 
and  so  on.  In  the  fourth  generation,  there  were  thirty-two 
ancestors,  and  in  the  sixth,  sixty-four.  In  the  course  of  a 
century  and  a  half,  these  sixty-four  ancestors  had  each 
contributed  a  sixty-fourth  part  to  any  one  individual's 
characteristics.  Generally,  there  was  a  certain  accidental 
blending  of  all  these  sixty-four  characters,  so  as  to  produce 
a  particular  result.  Take  for  instance  the  case  of  a  nmsician. 
Out  of  the  sixty-four,  there  might  perhaps  have  been  six  or 
eight  wIjo  were  rather  above  the  average  in  music.  It 
generally  happened  there  were  as  many  below  the  average 
as  woidd  balance  this,  and  then  the  result  was  an  ordinary 
person  who  was  neither  much  above  nor  below  the  average  in 
musical  capacit3\  But  where  it  happened  that  a  certain 
number  of  the  sixty-four  were  rather  above  the  average,  and 


2o-l<     Proceedings  of  the  Royal  Society  of  Victoria. 

there  were  none  much  below  it,  and  where,  added  to  a 
musical  capacity,  there  was  sufficient  industry  and  inven- 
tiveness, the  result  was  a  musician.  The  result  of  all 
this  theory  was  that  ci'irahials,  if  not  compelled  to  herd 
together,  would  work  out  their  own  salvation  in  genera- 
tion after  generation.  If  they  were  compelled  to  herd 
together,  the}^  invariably  died  out.  A  criminal  woman 
very  rarely  left  posterity  that  would  survive  two  or 
three  generations.  With  regai'd  to  men,  too,  in  following 
up  the  history  of  Austi-alia  he  had  been  surprised  to  observe 
that  there  was  so  little  trace  of  convict  blood  that  liad  been 
poui-ed  out  on  these  shores  so  profusely.  Australia  was  not 
less  moral  than  any  similar  Anglo-Saxon  community.  The 
question  was  sometimes  asked,  why  we  were  not  deeply 
tinted  with  the  convict  element.  There  were  two  classes  of 
convicts.  One  class  who  had  not  inherited  the  cjiminal 
character,  but  who  had  been  sent  out  for  committing  crimes, 
chiefly  by  reason  of  ttieii'  super-abundant  and  mis-directed 
energy,  had  made  excellent  settlers,  whilst  the  real  criminals 
had  been  killed  by  drinking  the  plentiful  rum  of  the  early 
settlement  days,  or  being  knocked  on  the  head  in  brawls, 
and  had  left  no  posterit3^  If  nature  were  left  to  work  in 
her  own  way,  the  matter  would  come  right  in  the  end.  The 
criminal  nature  would  either  not  perpetuate  itself,  or  the 
average  would  be  rectified  in  succeeding  generations  as  the 
area  extended.  Therefore,  the  crude  notion  of  chloroforming 
the  criminal  should  be  disregarded  for  several  reasons.  It 
was  not  in  accordance  with  the  humanitarian  views  of  the 
present  day,  and  it  was  founded  on  a  wrong  impression  of 
the  law  of  heredit}'.  If  criminal  were  made  to  paii-  with 
criminal,  the  I'esult  would  no  doubt  be  a  very  bad  race 
indeed,  supposing  they  bred.  But  they  would  not ;  and, 
moreover,  they  did  not  and  could  not  be  compelled  to  pair 
wi^h  one  another.  In  conclusion  he  believed  that,  at  the 
present  time,  things  should  be  allowed  to  remain  as  they 
were,  and  that  the  pressure  of  public  opinion,  and  where 
necessary,  of  sharp  public  punishment,  would  cast  sufficient 
weight  on  the  right  side  of  the  motive  to  induce  people  to 
act  as  best  suited  the  community.  Beyond  this,  there  was 
no  need  to  interefere.  We  should  follow  the  old  lines  as 
much  as  possible,  making  our  laws  as  humanitarian  as  the 
interests  of  society  would  allow. 

The  Peesideis'T  suggested  that  a  definition  of  criminality 
and  insanity  miglit  be  desirable. 


Froceeding.s  of  the  Royal  Society  of  Victoria,.      255 

Mr.  White  asked  Mr.  Sutherland  upon  what  statistics  he 
had  based  his  statemeiit  that  the  early  convicts  were  dying 
out.  Not  long  ago  the  early  criminal  records  of  Tasmania 
had  been  sent  to  xMel bourne  and  destroyed,  and  shortly 
afterwards  the  same  luid  been  done  with  those  of  New 
South  Wales.  He  would  be  much  relieved  if  it  could  be 
proved  that  the  ciiminal  class  v.^as  dying  out,  but  he  was 
unable  to  take  such  an  optimistic  view,  and  would  like 
to  ask  Mr.  Suthei'land  the  basis  of  his  statement. 

Dr.  Jamieson  said  he  hardly  thought  a  definition  of 
either  criminality  or  insanity  was  practicable.  He  doubted 
whether  there  was  any  definition  of  insanity  except  the  bald 
and  unsatisfactory^  one  that  it  was  some  de])artuie  from  the 
normal  mental  condition,  about  which  there  could  be  no 
doubt.  As  to  criminalit}',  he  thought  thei'e  was  more 
difficulty  still.  The  idea  of  criminality  varied  indefinitely 
almost  from  one  generation  to  another,  and  in  different 
races.  In  fact  within  a  very  short  time  it  had  been  made 
an  offence  punishable  by  law  to  buy  or  sell  a  pound  of 
sugar.  It  was  not  an  offence  for  a  man  to  sell  a  cigar  or 
glass  of  whisky  at  8  o'clock  at  night,  but  it  was  an  ofience 
to  sell  a  pound  of  sugar  at  that  hour.  The  idea  seemed  to 
be  that  criminalitj-  was  the  doing  of  certain  things  which  the 
majority  had  agreed  ought  not  to  be  done,  and  the  doing  of 
which  should  be  punished.  Such  an  offence  was  called  a 
crime.  It  was  well  known  that  there  were  habitual 
criminals,  people  who  made  it  their  business  to  commit 
offences  against  the  law,  but  it  was  difficult  to  get  at  the 
root  and  origin  of  this  tendency.  He  believed  a  great  many 
offences  were  due  to  intellectual  defects.  There  was  a  clear 
enough  distinction  between  a  man  being  weak  and  being 
wicked,  but  it  was  quite  certain  that  weakness  very  soon 
led  to  wickedness,  and  a  person  who  was  weak  in  body 
or  mind  ran  far  greater  risk  of  falling  into  criminality  than 
a  person  fairly  endowed  with  mental  ability  and  physical 
power.  The  man  who  was  mentally  weak  was  liable  to  be 
led  into  crime  by  stronger  minded  and  less  scrupulous 
persons,  and  the  man  who  was  weak  in  body  was  at  a 
disadvantage  iti  earning  his  living,  and  fell  into  such  straits 
that  he  was  tempted  to  break  the  law  and  ap])ropriate  that 
to  which  he  had  no  right.  This  habit  would  grow  in  both 
cases,  and  he  did  not  see  why  a  person  without  any  special 
criminal    tendency    might    not,    if    constantly    exposed    to 


256      Proceedings  of  the  Royal  Soeiety  of  Victoria. 

temptation,  become  by  force  of  liabit  a  habitual  criminal. 
This  much  was  clear  enough,  and  capable  of  proof,  but  wlien 
it  came  to  a  (juestion  of  moi-al  delect,  it  was  difficult  to  s;  y 
how  lar  it  was  natural  and  how  far  acquired.  It  was  just 
as  ])robable  that  people  failed  in  moral  capacit}'  just  as  they 
did  in  intellectual  capacity,  but  it  did  xiot  iiecessaiily  follow 
that  both  defects  should  be  co-existent  in  the  same  pei'son. 
A  man  might  be  strongly  endowed  mentally,  and  yet  Ix' 
weak  in  moral  qualities,  and  it  had  been  observed  that  some 
people  who  were  well  endowed  morally  were  rather  poor  in 
intellectual  constitution.  But  it  was  difficult  to  say  how  lai 
what  Avas  called  moral  detect  was  natuiul,  and  how  far  it 
was  acquired.  It  might  be  admitted  as  a  likely  enough 
thing  that  there  were  peojjle  insufficiently  endowed  with 
moral  qualities,  who  readily  enough  became  criminals, 
independently  of  their  intellectual  capacit}^,  and  as  a  meic 
matter  of  theory  it  might  be  admitted  that  the  moral 
endowment  could  be  so  poor  that  the  person  of  necessity 
became  a  criminal.  He  would  not  like  to  say  on 
theoretical  grounds  that  this  was  not  so,  but  the  difficulty 
was  m  proving  it  to  be  so,  and  to  recognise  a  theoieticai 
deficiency  of  moi-al  endowments,  apart  from  pure  mental 
capacity,  as  a  ground  of  iri-esponsibilities,  was  excessively 
dangerous  doctrine.  But  this  seemed  to  be  the  doctrine 
held  by  the  modern  school  of  criminal  anthropologists,  who 
went  so  far  as  to  sa}^  that  they  could  tell  pretty  accurately 
what  would  be  the  physical  characteristics  of  the  habitual 
criminal.  He  did  not  think  any  of  them  would  profess  to 
be  able  to  tell  from  the  physical  characteristics  of  a  man, 
without  knowing  anything  about  his  conduct,  whether  or 
not  he  was  a  criminal,  and  this  was  the  difficulty  of  safely 
apyjlying  the  doctrine,  however  rational  it  might  be  as  a 
general  principle,  to  individual  cases.  For  that  reason  he 
thought  that  any  attempts  to  save  criminals  from  the 
consequences  of  their  actions  on  such  grounds  should 
certainly  not  be  encouraged,  but  should  on  the  contiaiy 
be  discouraged  very  strongly.  With  regard  to  the  ques- 
tion of  punishment,  he  agreed  with  Mr.  Sutherland 
that  ret'oT-mation  could  not  properly  be  spoken  of  as 
a  form  of  punishment,  although  the  criminal  might 
regard  in  that  light  any  etlbrts  made  by  the  authorities 
in  that  direction.  He  feared  that  retiilmtion  could  not  be 
got  rid  of  If  one  man  inflicted  injuries  u])on  another  that 
could  be  measured  pecuniarily,  he  was  fined  to  a  proportionate 


ProceeiUii(/-i  of  the  Roijal  Socistij  of  Victoria.     'Ihl 

extent.  Witliout  a  doubt  there  was  retribution  in  that. 
The  man  who  had  not  the  money  to  pay  was  entitled  to 
be  punished  by  having  something  taken  out  of  him  as  a 
quid  pro  quo.  He  was  not  prepared  to  drop  the  old-fashioned 
idea  of  retribution.  Of  course  jvunishment  was  also  deterrent. 
It  liad  a  deterrent  effect  upon  the  offender  himself,  inasmuch 
as  things  were  made  so  unpleasant  ftjr  him  that  he  would 
not  be  likely  to  repeat  the  offence.  The  extreme  deterrent 
was  the  taking  away  of  life.  After  all,  people  value  their 
life  more  highly  than  anything  else,  and  death  was  the 
sti-ongest  possible  deterrent  the  law  could  inflict.  It 
practically  amounted  to  this,  that  the  community  recognised 
certain  crimes  as  being  of  such  an  atrocious  character,  that  it 
sim})ly  decreed  that  those  who  committed  them  should  not 
only  be  banished  from  human  society,  but  should  have  their 
existence  terminated,  not  only  as  a  strong  deterrent,  but  to 
prevent  the  possibility  of  the  act  ever  being  repeated  by 
them.  It  was  not  done  with  any  view  of  exterminating 
the  breed,  that  was  a  hopeless  task  he  was  afraid.  The 
meeting  was  greatly  indebted  to  Colonel  Goldstein  for 
the  manner  in  which  he  had  brought  the  matter  befoi-e 
it.  He  was  mistaken  in  thinking  that  because  a  man  had 
malformation  of  the  brain,  or  some  disease,  that  he  was 
therefore  insane.  He  did  not  think  many  members  of  the 
medical  profession  would  hold  such  ci'ude  doctrines  as  that. 
It  was  quite  certain  that  there  might  be  diseases  of  the 
brain  without  any  indication  of  insanity,  and,  although  it 
was  very  unlikely,  there  might  be  mental  derangement 
without  any  demonstrable  disease  of  the  brain.  As  to  other 
matters,  he  was  sure  there  were  not  many  who  w^ould  be 
inclined  to  differ  greatly  from  what  Colonel  Goldstein  had 
said. 

The  Rev.  E.  H.  Sugden  said  that,  in  a  definition  of 
crisiiinality,  something  was  wanted  which  would  connect 
the  selling  of  an  article  after  hours  and  the  taking  of  life  in 
cold  blood.  The  anti-social  spirit  that  both  acts  displayed 
constituted  them  crimes  against  society.  It  seemed  to  him 
that,  in  order  to  lessen  the  criminal  po))ulation,  the  artificially 
made  crimes  against  society  ought  to  be  lessened  as  much  as 
possible.  A  glance  at  the  list  of  indictable  offences  would 
show  that  ninety-nine  out  of  every  hundred  were  not  wrong 
in  themselves,  but  were  wrong  because  society  had  made 
them  wrono-.      Takina  the  view  that  the  criminal  was  an 


258      ProceciliiKjs  of  the  Royal  Scclcfy  of  Vidofia. 

offeiuler  against  society,  it  seemed  to  hi  in  tliat  the  proper 
piinishnjent  would  l)e  to  send  liini  to  Coventry,  if  some 
reasonable  method  of  doing  so  coul(i  1  e  indicated.  If  a 
reasonalily  fertile  and  productive  part  of  the  earth  could  l;e 
fenced  off  and  made  a  dumping  ground  for  ci'iminals,  where 
they  could  be  left  to  develope  a  State  for  themselves  on 
their  own  anti-social  lines,  it  would  be  a  veiy  comfortable 
thing  for  Society,  and  perhaps  a  very  salutary  discipline  for 
themselves.  Tliis  he  was  afraid  was  somewhat  Utopian, 
but  if  the  social  instinct  could  be  aroused  in  tiie  so-called 
criminal  classes,  a  great  deal  of  crime  would  be  prevented. 
If  a  man  could  be  taught  to  entertain  a  real  and  tender 
regard  for  any  living  thing,  a  great  deal  had  been  done  to 
pievent  him  from  committing  crime.  In  spite  of  Mr. 
Sutherland  and  others,  the  Christian  Clnnch  still  believed 
that  the  criminal  could  be  reformed,  if  he  could  onl}'  be  got 
to  love  someone,  and  that  the  one  Person  whom  it  was 
easiest  and  most  effectual  to  bring  him  to  love  A^as  our 
common  Master.  It  seemed  to  him  these  were  the  lines  on 
which  the  best  results  would  be  obtained.  Tlie  natural 
punishment  for  one  who  manifested  tlie  anti-social  spirit 
would  be  to  shut  him  out  of  society,  and  the  only  i-emedy 
would  be  the  })romotion  of  the  social  spirit  in  him. 


Thursday,  July  ]4fA. 

The  President  (Professor  Kernot)  occupied  tlie  chair. 

On  the  motion  of  Mr.  Griffiths,  seconded  by  Mr. 
RbSDEN,  the  minutes  of  the  preceding  meeting  were  taken 
as  read  and  duly  confirmed. 

Mr.  G.  C.  W.  Officer,  Member,  and  Mr.  Strettle,  Associate, 
present  for  the  h'rst  time,  signed  the  Roll,  and  were  introduced 
to  the  meeting. 

Mr.  Donald  Clarke,  of  the  School  of  Mines,  Bairnsdale, 
was  elected  a  Country  Member. 

The  President  announced  that  the  following  gentlemen 
had  been  nominated,  and  would  be  balloted  for  at  the  next 
meeting  : — Rev.  Walter  Fielder,  Associate  ;  L.  J.  Balfour, 
Member  ;  Douglas  Howard,  Associate. 

Professor  Spencer,  in  the  absence  of  the  Hon.  Librarian, 
reported    that    since    the    last    meeting,    92    volumes    and 


ProceediiKjii  of  the   Roijal  Society  of  Victoria.      2')9 

periodicals  had  been  received,  and  the  Council  had  deter- 
mined to  proceed  with  the  binding  of  the  books  belonging 
to  the  Libraiy,  and  53  volumes  had  been  sent  to  the 
binders  for  that  purpose. 

The  Rev.  A.  Cresswell  read  a  paper  entitled  "  Notes 
on  the  Lilydale  Limestone,"  illustrating  his  remarks  hy 
specimens  and  blackboard  drawings. 

Mr.  Grip'FITHS  considered  that  the  geologists  of  Victoria 
were  under  a  debt  of  obligation  to  the  Rev.  Mr.  Cresswell, 
for  having -undertaken  the  description  of  this  interesting 
bed  of  limestone.  So  far  as  his  recollection  went,  this  was 
the  only  bed  of  crystalline  limestone  found  in  Victoria  in 
the  Silurian,  eithei-  upper  or  lower.  The  only  other  deposits 
of  crystalline  limestone  known  to  him  were  some  beds  in 
Gippsland,  in  one  of  which  Mr.  Sweet  had  discovered  some 
very  interesting  fossil  tish.  It  was  to  him  a  source  of 
wonder  that  the  bed  described  by  Mr.  Cresswell,  lying  so 
near  Melbourne,  had  remained  for  so  many  years  without 
.  any  description  that  was  accessible  to  the  public.  Mr. 
Cresswell  had  been  so  successful  as  to  obtain  a  large  number 
of  interesting  fossils,  and  no  doubt  his  paper  would  stimulate 
geologists  generally  to  pay  more  attention  to  the  bed  of 
limestone  at  Lilydale. 

Mr.  Dennant  regretted  very  much  he  never  had  an 
opportunity  of  visiting  Lilydale,  and  was  much  gratified 
to  see  that  this  matter  had  been  taken  up  by  Mr.  Cresswell. 
He  was  particularly  glad  that  some  palfeontological  work 
would  embellish  the  pages  of  the  "Transactions."  He  did 
not  know  whether  this  bed  was  definitely  known  to  extend 
furtiier  in  the  east,  but  he  had  heard  it  said  by  those  who 
knew  the  country,  that  there  were  outcrops  of  limestone 
t)n  the  Upper  Yarra,  about  fifteen  miles  to  the  north-west 
of  the  Emerald  Township  and  Gembrook,  but  he  had  never 
had  an  opportunity  of  personal!}^  verifying  the  fact.  It 
would  be  interesting  to  know  whether  this  was  really  an 
extension  of  the  limestone  which  outcropped  at  Lilydale. 

Ml'.  Sweet  said  he  had  visited  the  limestone  beds  at 
Lilydale  on  several  occasions,  and  had  taken  considerable 
interest  in  tliern.  He  was  therefore  very  pleased  that 
Mr.  Cresswell  had  taken  the  matter  up,  and  hoped  he  would 
continue  his  labours  until  he  had  given  them  a  complete 
list  of  all  he  had  found. 

S   -2 


200      Proceedings  of  the  Rotjal  Society  of  Victoria. 

Mr.  Pritchard  said  there  were  some  other  specimens 
which  he  had  obtained  on  a  recent  visit,  and  which  were 
very  interesting.  The  first  was  a  coral  which  Mr.  Cresswell 
had  mentioned,  namely,  Heliolites.  A  large  quantity  of 
this  material  had  been  obtained  on  the  last  visit,  and  was 
in  a  very  perfect  state  of  preservation,  so  that  it  would  be 
a  good  specimen  for  description.  There  was  another  fossil 
which  occurred  there  rather  commonly,  but  which  u\)  to 
the  present  had  not  received  any  mention  at  all,  although 
it  had  been  discovered  four  ov  five  years  ago.  It  was  a 
kind  of  operculum,  which  had  been  handed  to  Professor 
McCoy,  but  had  never  been  descr-ibed.  It  seemed  to  corres- 
pond with  the  Gyclonema  a/ustralis,  and  he  thought  it 
would  be  well  if  something  were  done  in  connection  with 
the  specimens  he  had  mentioned.  There  were  several  other 
specimens  of  the  coral  type,  some  of  which  were  very 
nteresting,  and  which  he  had  not  been  able  to  identify  at 
all.  He  would  be  glad  to  hand  them  over  to  any  Member 
who  would  undertake  a  description  of  them. 

Mr.  Hall  said  that  Mr.  Pritchard  had  forgotten  to 
mention  that  one  of  the  specimens  of  which  he  had  spoken, 
the  operculum  of  some  shell,  had  been  discovered  wedged 
into  the  mouth  of  an  Euomphalus.  Whether  it  belonged  to 
the  Euomphalus  or  not  was  a  matter  which  would  admit  of 
discussion.  They  were  all  obliged  to  Mr.  Cresswell  foi-  his 
interesting  paper. 

The  President  said  that  he  had  been  in  the  vicinity  of 
the  Upper  Yarra  a  good  many  times,  but  did  not  remember 
noticing  any  limestone.  However,  he  had  not  been  on  a 
geological  ex])edition,  and  it  was  perfectly  possible  there  was 
plent}^  of  limestone  which  he  had  not  noticed. 

Rev.  Mr.  Cresswell,  in  reply,  .said  that  if  any  limestone 
did  exist  on  the  Upper  Yai'ra,  it  was  just  in  position  to  be  a 
repetition  by  flexure  of  the  limestone  at  Lil3^dale.  It  was  a 
mistake  to  suppose  there  were  no  other  beds  of  Upper 
Silurian  limestone  in  Victoria.  There  were  two  very 
extensive  thick  beds  in  Gippsland,  one  at  Walhalla,  near 
Cooper's  Creek,  and  another  at  Deep  Creek,  about  seven  oj- 
ten  miles  from  Walhalla.  They  were  nearly  parallel  seams 
of  limestone,  and  might  be  a  repetition  by  flexure.  The 
fossils  Avere  very  similar  in  both.  These  beds  had  been 
known  for  a  considerable  number  of  years,  and  were  alluded 
to  in  the  Geological    Survey  Reports.      In  conclusion,   he 


Proceedings  of  the  Royal  Society  of  Victoria.     2G1 

expressed  his  indebtedness  to  Mr.  Pritchard,  who  had  lent 
him  the  fossil  which  formed  the  chief  matter  he  had  spoken 
about  that  evening,  and  recorded  his  thanks  to  Mr. 
David  Mitchell  and  his  foreman,  Mr.  Fuller,  for  statistical 
information. 

A  "  Preliminary  Note  on  the  Glacial  Deposits  of  Bacchus 
Marsh,"  by  Messrs.  C.  G.  W.  Officer  and  L.  J.  Balfour,  was 
read  by  Mr.  Officer. 

Mr.  Dennant  said  that  there  were  a  number  of  questions 
raised  by  the  paper  which  he  would  like  an  opportunity  of 
speaking  upon  at  considerable  length,  and  as  time  would  not 
permit  of  a  protracted  discussion  that  evening,  he  moved 
"That  the  discussion  on  the  paper  be  postponed  till  the 
next  meeting." 

Mr.  Cres.SWELL  seconded  the  motion  for  the  reasons  stated 
l)y  Mr.  Dennant. 

Tile  motion  was  agreed  to. 

Report  of  the  Ceemation  Committee.* 

The  Report  of  the  Cremation  Committee  was  read  by  the 
Hon.  Secretary,  Mr.  RusDEN. 

On  the  motion  of  Professor  Spencer,  seconded  by  Di-. 
Brett,  the  Report  was  received. 

The  President  said  that  a  model  had  been  procured 
which  would  ilkistrate  the  working  of  the  Gorini  incinerator, 
which  was  the  process  adopted  at  Milan,  where  ciemation 
appeared  to  be  practised  to  a  very  considerable  extent,  and 
in  a  way  not  calculated  to  offend  or  disgust  in  any  respect. 
The  Crematory  and  its  surroundings  were  pleasant  and 
attractive.  Milan  was  one  of  the  largest  and  busiest  cities 
in  the  northern  parts  of  Italy,  and  the  Crematory  was  as 
near  to  the  centre  of  Milan  as  the  present  Melbourne 
Cemetery  was  to  the  centre  of  Melbourne. 

Mr.  F.  Chamberlain  produced  the  model  and  explained 
its  construction. 

Dr.  Gresswell  was  heartily  in  accord  with  the  strong- 
recommendations  of  the  Cremation  Committee.  He  felt 
that  it  was  a  reform  that  was  bound  to  come,  although 
he  was  not  ver}^  sanguine  as  to  its  coming  quickly.  How- 
ever, he   felt   satisfied    with    the    progress    being    made    as 

*  VUU  Stipri',  p.   2'22. 


202     Froceedinij^  of  ike  Royal  ISociety  of  Victoria. 

indicated  in  the  Report,  and  hoped  that  the  Report  would 
eiiconrfige  those  who  were  interested  to  make  further  etibrts. 
With  reference  to  the  observations  in  the  Report  in  regard 
to  Japan,  it  was  an  interesting  fact  tliat  for  a  very  long 
time  past.  Cremation  had  been  the  rule  in  Japan,  but  upon 
the  introduction  of  European  civilisation  into  Japan  fifteen 
or  twenty  years  ago,  it  was  thought  right  to  follow  the 
European  customs  in  the  matter,  and  substitute  burial  tor 
Cremation.  They  soon  discovered  their  error,  and  reverted 
to  their  former  practice. 

Dr.  Brett  moved,  "  That  the  Report  be  adopted  by  the 
Society,  and  printed  in  its  Proceedings."  He  took  consider- 
able interest  in  the  subject  of  Cremation,  and  had  seen' it 
practised  in  nearly  every  country  in  the  world. 

Mr.  G.  A.  Syme  seconded  the  motion,  which  was  carried 
Unanimously. 


Tlturciilai],  AuytiM  Mth. 

The  President  (Professor  Keknot)  occupied  the  chair. 

The  minutes  of  the  [)receding  meeting  were  read  by  the 
Secretary,  and  duly  confirmed. 

Dr.  Barrett,  Member,  and  Mr.  Barnard,  Associate,  present 
for  the  fii'st  time,  signed  the  Roll,  and  were  introduced  to  the 
meeting. 

The  President  announced  that  the  following  gentlemen 
had  been  nominated,  and  would  be  balloted  for  at  the  next 
meeting : — W.  H.  Steele,  M.A.,  Associate  ;  Frederick 
Chamberlain,  Member;  Alfred  Stillwell,  Member;  and 
A.  Purdy,  M.A.,  Associate. 

The  following  gentlemen  were  balloted  for,  and  duly 
elected  : — The  Rev.  Walter  Fielder,  Associate ;  Douglas 
Howard,  Esq.,  Associate  ;  and  L.  J.  Balfour,  Esq.,  Member. 

The  Librarian's  Report  stated  that  69  publications  liad 
been  received  from  various  parts  of  the  world,  and  -i-i  bound 
volumes  from  the  binders. 

Adjourned  discussion  on  "  Pj-eliminary  Note  on  the 
Glacial  Deposits  of  Bacchus  Marsh,"  by  C.  G.  W.  Officer, 
B.Sc,  and  L.  J.  Balfour. 

-    Mr.  Officer  said  that  since  the  paper  was  read  he  had, 
together    with     Mr.    Balfour,    traversed    the    whole    of   the 


Proceedings  of  the  Royal  Society  of  Victoria.     203 

district  covered  by  the  paper,  and  he  wished  to  make  one  or 
two  corrections.  The  first  was  with  regard  to  the  sections 
described  on  the  Myrniong  Creek.  It  had  been  stated  that 
the  ghicial  deposit  was  overlaid  by  older  basalt.  This  part 
of  the  map  was  outside  of  that  published  by  the  Geological 
Survey,  and  he  would  like  to  say  that  this  basalt  was 
probably  to  be  referred  to  the  upper  and  newer  basalt,  and 
not  the  older.  In  the  not-e,  it  had  also  been  stated  that  a 
certain  section  was  probably  an  example  of  contorted  till. 
On  further  examination,  however,  this  had  proved  to  be  not 
till,  but  what  was  called  Mesozoic  sandstone,  and  the  apparent 
contortions  were  due  to  concretionary  action.  Also  at  another 
section  where  the  glacial  deposit  was  overlaid  with  sandstone, 
it  had  been  difficult  to  decide  whether  that  sandstone  was 
simply  associated  with  till,  or  belonged  to  the  Mesozoic 
sandstone  in  the  surrounding  district.  They  were  now  of 
opinion  that  the  overlying  sandstone  .was  probably  of 
Mesozoic  age.  On  the  Korkuperrimul  Creek  the  glacial  till 
was  overlaid  by  basalt,  which  they  thought  was  to  be  assigned 
to  the  upper  basalt.  But  the  evidence  on  which  the  distinc- 
tion between  upper  and  lower  basalt  was  often  drawn, 
seemed  to  be  somewhat  feeble. 

The  Rev.  Mr.  Cresswell  said  he  had  a  few  criticisms  to 
offer  upon  the  paper  which  Mr.  Officer  had  kindl}^  lent  him, 
and  he  would  begin  by  recording  his  appreciation  of  the 
value  of  the  paper.  It  was  a  most  interesting  and  complete 
paper,  although  he  was  not  able  to  agree  with  all  the 
conclusions  arrived  at  by  the  authors.  From  the  evi- 
dence adduced  by  other  observers,  as  for  instance  Mr. 
Selwyn,  Sir  Richard  Daintree,  Dr.  Lendenfeld,  and  Mr. 
Dunn,  there  could  be  no  doubt  that  in  former  ages 
there  had  existed  distinct  glaciers  in  the  Alpine  districts 
of  Victoria  and  New  South  Wales,  but  it  seemed  to  liini 
to  be  very  questionable  whether  those  glaciers  extended 
any  distance  from  those  particular  regions.  He  would 
begin  by  making  a  general  criticism  upon  the  terms 
used.  He  thought  it  somewhat  misleading  to  appl}'-  the 
term  "till"  to  two  formations  which,  according  to  the 
authors'  showing,  were  so  very  widely  separated  in  the 
geological  series — one  being  apparently  a  Pleistocene  glacial 
deposit  of  some  kind,  and  the  other  being  a  glacial  depo.sit 
belonging  to  the  Miocene  age.  He  thought  it  far  better  to 
keep    the    term  "till"  for    well-known    and    acknowledged 


264^     P roceed'mrjs  of  the  Royal  Society  of  Victoria. 

de[)osit.s  of  glacial  nature  beloDging  to  the  Pleistocene  age. 
To  a]3ply  the  term  to  both  formations  tended  to  confusion. 
He  was  one  of  those  who  believed  that  till  was  not  a 
Moraine  jn-ofonde,  but  owed  its  origin  to  Moraine  matter, 
redistributed  par-tly  by  aqueous  action,  and  the  boulders 
contained  in  it  were  mostly  the  result  of  icebergs  which  had 
broken  away  from  glaciers  and  deposited  the  debris  in  the 
clay.  Still  less  was  he  inclined  to  believe  that  these 
particular  formations  were  instances  of  a  ground  Moraine, 
and  he  doubted  that  they  necessarily  indicated  that  there 
had  l)een  glaciers  on  the  very  spot  where  they  had  been 
found.  With  regard  to  the  ,  striated  pebbles,  no  one 
could  have  the  smallest  doubt  as  to  their  being  striated, 
and  having  been  striated  by  glacial  action.  They  were 
evidently  striated-glaciated  pebbles.  He  very  much 
doubted,  however,  that  they  had  been  scratched  by 
any  rocks  where  they  were  now  found.  So  far  as 
his  memory  served  him,  the  Upper  Mesozoic  sandstones 
wei-e  particularly  soft,  and  incapable  of  scratching  these 
pebbles,  and  he  therefore  imagined  that  they  must  neces- 
sarily have  come  from  a  very  considerable  distance — 
probably  from  the  Alpine  regions  of  either  New  South 
Wales  or  Victoria— and  5iot  from  the  neighbourhood  where 
they  v/ere  now  found.  These  remarks  ap)plied  to  the 
upper  till,  or,  as  he  would  call  it,  the  upper  glacial  formation. 
With  i-egai-d  to  the  lower  glacial  formation,  he  was  not 
very  well  acquainted  with  the  nature  of  the  Silurian  rocks 
in  the  neighbourhood,  but  unless  they  contained  quartzites 
very  abundantly,  he  did  not  know  of  any  rock  likely  to  be 
capable  of  imparting  the  stride  to  these  pebbles  in  the 
Silurian  area.  He  believed  that  these  pebbles,  although  no 
doubt  glaciated,  had  been  brought-  from  a  considerable 
distance  by  alluvial  and  other  action.  As  to  the  stride  on 
the  Silurian  rocks,  he  would  be  very  sorrj^  indeed  to  insult 
their  friends'  powers  of  observation,  by  implying  that  thej'' 
could  possibly  have  made  a  mistake  if  the}'  had  had  the 
opportunity  of  observing  them  upon  an  extended  scale  ;  but 
considering  that  they  liad  not  had  such  an  opportunity,  but 
had  only  found  the  strise  here  and  there  in  small  patches,  he 
would  venture  to  ask  them  whether  the}'  might  not  have 
mistaken  the  unequal  wearing  of  the  edges  of  the  rock. 
He  understood  from  the  paper  that  these  grooves  were  not 
found  in  the  Mesozoic  sandstone,  but  only  in  connection 
with    the   lower  drift  on   the    Silurian  rocks,   and  he  was 


Proceedings  of  the  Royal  Society  of  Victoria.      265 

particularly  struck  by  the  remark  made  in  the  paper,  that 
these  grooves  invariably  lan  north  and  south,  that  was  to 
say,  exacth'  coincident  with  the  strike  of  the  strata,  and 
these  Silurian  strata  were  tilted  up  at  an  angle  of  70  degrees. 
He  did  not  know  the  extent  of  the  patches  uncovered,  but 
considering  that  the  observers  were  most  anxious  no  doubt 
to  see  what  they  saw,  was  it  not  possible  that  they  might 
have  mistaken  the  unequal  wearing  of  the  edges  of  the 
Silurian  strata.  He  had  frequently  seen  on  the  smooth 
upturned  edges  of  the  Silurian  strata  appearances  of  very 
deep  grooves,  owing  to  the  unequal  wearing.  But  even 
supposing  them  to  be  veritable  grooves,  it  was  not  beyond 
the  range  of  possibility  that  they  might  have  been  formed 
by  an  iceberg  charged  with  hard  pebbles  underneath 
grounding  in  the  neighbourhood  on  the  spot.  With  re- 
gard to  the  roche  moulonnde,  he  laboured  under  the  dis- 
advantage of  never  having  been  on  the  particular  spots 
which  had  been  mentioned,  so  that  he  was  unable  to 
judge  from  his  own  observation  whether  they  were  really 
instances  of  roche  moutonnee.  He  would  just  conceive  it 
possible,  therefore,  that  the  hummocky  appearance  might 
have  been  caused  by  diluvial  action.  For  instance,  at 
Lilydale,  there  was  an  appearance  of  hummocky  rock, 
which  was  simply  caused  by  the  action  of  the  waves 
on  the  seashore.  Messrs.  Otficer  and  Balfour  objected 
to  the  idea  of  marine  action  having  anything  to  do 
with  the  phenomena  they  instanced,  but  he  would 
venture  to  point  out  that  tliere  was  indisputable  evi- 
dence that  the  whole  continent  had  been  submerged 
thousands  of  feet  under  the  i^ea.  He  had  seen  gravel 
on  the  top  of  Mount  Useful  covered  with  basalt,  and 
this  was  generally  put  down  on  the  geological  maps  as 
marine  gravel.  There  was  some  doubt  as  to  the  age 
of  that  particular  gravel,  but  there  could  be  no  doubt  as 
to  the  age  of  the  drifts  at  Gastlemaine  and  Bendigo,  and 
other  places,  which  would  represent  a  submergence  of  at 
least  2000  feet.  It  was  a  very  moot  point  as  to  whether 
these  drifts  had  been  caused  by  y)luvial  action,  extending 
over  a  considerable  pluvial  period  ;  but  he  believed,  with 
Selwyn  and  others,  that  they  were  caused  b}^  marine  action. 
Although  he  had  appeared  to  criticise  the  paper  somewhat 
advei-sely,  he  quite  admitted  it  was  a  very  debateable 
question.  Not  being  an  extreme  glacialist,  he  was,  perhaps, 
inclined   to  minimise   the  evidence  adduced,   but   he   could 


20 G     Proceedings  of  the  Royal  Society  of  Victoria. 

quite  conceive  tliafc  others  who  were  extreme  glacialists 
would  concede  that  the  authors  had  fully  proved  tlieir 
point. 

Mr.  Griffiths  said  that  for  purposes  of  discussion  the 
paper  might  be  divided  into  two  parts — the  part  which  was 
purely  descriptive,  and  the  part  which  offered  explanations 
of  the  appearances  described.  The  latter  part  might  be 
divided  into  three  principal  propositions  which  were  put 
forward  although  not  formally  stated.  The  first  was  that 
there  were  evidences  locally  of  two  glacial  e})Ochs — one  earl}^ 
in  the  Permian,  the  other  early  in  the  Tertiary.  The 
second  was  that  the  boulder  clay  of  each  of  these  was  due 
to  land  ice,  and  not  to  marine  transport  by  icebergs.  The 
third  was  that  the  submergence  of  the  continent  sufficient 
to  ffoat  an  iceberg  at  Bacchus  Marsh,  would  reduce  the 
land  surface  to  such  a  small  area  that  it  would  be  too 
limited  in  area  to  breed  icebergs,  and  too  warm  to 
accumulate  ice  on  account  of  its  insularity.  With  respect 
to  the  descriptive  part  of  the  paper,  he  had  found 
many  discrepancies  between  the  descriptions  given  b}' 
the  authors  of  the  paper  and  the  statements  made  by  the 
officers  of  the  Geological  Survej^  The  Government  officers 
had  given  a  section  showing  a  thick  bed  of  what  had  since 
been  termed  Trias.-ic  conglomerate,  which  the  authors  of  the 
paper  had  attributed  to  a  different  period,  but  did  not  give 
a  tripartite  division,  which  Messrs.  Officer  and  Balfour 
stated  to  exist  in  the  section  desciibed  on  the  Werribee 
River.  The  Government  officers,  who  were  men  of  ex- 
perience, had  failed  to  recognise  any  glaciated  rocks  in 
this  sect-ion,  although  they  had  stated  that  a  glacial  con- 
glomerate existed  in  the  distiict.  Of  course,  it  sometimes 
happened  that  through  want  of  sufficient  data,  errors  had 
crept  into  the  Geological  maps,  and  he  simply  pointed 
this  out  as  showing  that  it  wa.s  advisable  to  carefully 
weigh  the  evidence  adduced  before  accepting  it.  Although 
the  Government  geologists  had  not  seen  their  way  to 
describe  the  bottom  member  of  this  section  as  a  glacial 
deposit,  they  had  pointed  out  that  the  Mesozoic  sand- 
stone was  composed  of  two  members,  the  upper  being  a 
sandstone  and  the  lower  being  a  conglomerate,  and  stated 
that  this  conglomerate  was  due  to  marine  action.  They 
recognised  a  difference,  but  attributed  it  to  a  different 
cause.       With   regard    to   the    ffrst  proposition,   that   there 


Pr<H'eedii)(js  of  the  Royal  Society  of  Victoria.     267 

was  evidence  of  two  glacial  epochs,  it  certainly  did 
se<'in  upon  examination  of  the  specimens  and  pho- 
tographs as  if  there  was  evidence  to  show  that  the 
bottom  member  ]iad  had  a  o-lucial  origin.  There  were 
un<loubted  grooves  and  stria?,  and  the  Silurian  rocks 
were  certainly  marked  as  if  a  plane,  with  grooves,  such 
as  a  carpenter  would  use  in  his  ornamental  work,  had 
passed  over  them.  Such  was  the  appearance  of  rocks 
which  had  undei'gone  grinding  by  the  passage  over 
them  of  a  glacier.  There  was  a  good  deal  of  weight 
in  Mr.  Ciesswell's  criticism  with  regard  to  the  rocks 
having  a  north  and  south  direction  so  far  as  their  strike 
■was  concerned,  and  the  edges  of  the  Silurian  rock  being 
uptilted  to  a  high  angle,  and  varying  degrees  of  hard- 
ness and  wearing  unequally,  and  all  this  would  have  to 
be  taken  into  consideration  before  coming  to  a  liual  conclu- 
sion ;  but,  nevertheless,  looking  at  the  photographs  of  the 
portion  of  the  rock  that  had  been  uncovered,  it  certainly 
suggested  a  glacial  origin  for  the  I'ock  to  his  mind.  In  the 
second  pla.ce,  the  boulders  in  the  till  were  certainly  to  a 
large  extent  striated,  and  as  suggested,  appeared  to  have 
been  de})Osited  there  by  ice.  There  was  also  another  feature 
which  was  favourable  to  the  proposition  suggested  by  the 
authors,  namely,  the  great  variety  of  rock  which  was  found 
in  this  clay  bed.  Such  a  heterogeneous  collection  of  rocks 
collected  together  in  one  place  without  any  stratification 
at  all,  and  most  of  them  more  or  less  striated,  was  cer- 
tainly to  his  mind  suggestive  of  glacial  action,  and 
pointed  to  a  glacial  origin  for  the  deposit.  Then  again 
the  rocks  were  not  only  varied,  but  they  did  not 
c;)rresj»ond  with  the  rocks  in  the  neighbourhood.  If  the 
conglomerate  was  due  to  coastal  action,  it  would  agree  with 
the  rocks  found  in  the  locality,  but  that  fact,  that  this  was 
not  the  case,  indicated  that  the  boulders  had  been  brought 
some  considerable  distance  ;  ;ind  it  was  well  known  that  a 
glacier  does  collect  rocks  in  this  manner,  and  that  in  a 
coastal  conglomerate  no  such  heterogeneous  collection  of  I'ocks 
is  found.  'J'his  was  another  ]x>int  in  favour  of  the  hypo- 
thesis of  the  authors.  It  seemed  to  him  that  the  specimens 
and  descriptions  all  favoured  a  glacial  origin  for  this  bed, 
but  whether  one  could  go  further  and  agree  with  the  authors 
of  the  paper,  that  the  bed  was  of  Permian  age,  was  another 
matter.  In  arriving  at  that  conclusion,  it  was  necessar}''  to 
take  a  number  of  matters  into  consideration.     In  the  first 


'lUH     Frvceedimjs  of  the  Royal  Society  of   Victoria. 

place,  tlie  conglomerate  was  associated  with  a  l)ed  of  rock 
which  was  known  to  be  of  Mesozoic  age.  In  the  second 
place,  there  were  no  Permian  rocks  described  in  Victoria,  and 
it  was  well  known  that  there  was  a  great  break  in  the 
sequence,  the  sandstor.es  which  were  the  Upper  Devonian 
rocks  being  the  last  rocks  met  with  before  reaching  the 
Mesozoic  sandstones.  There  was  no  such  break  in  the 
neighbourhood  of  Sydney,  where  there  was  a  complete 
sequence  from  the  Upper  Devonian  to  the  Oolite.  On  the 
Sydney  side  the  country  was  sinking  and  the  deposits 
accumulating,  but  in  Victoria,  the  other  end  of  the  sea-saw, 
the  country'  was  elevated  and  erosion  was  going  on.  There- 
fore, in  Victoria,  one  did  not  look  for  Permian  rocks,  and  if 
this  were  a  bed  of  Permian  age,  it  was  an  interesting  fact 
which  required  more  evidence  than  was  at  present  avail- 
able. Its  association  with  the  Mesozoic  rocks  had  led 
the  Government  geologists  to  associate  it  with  the  beds 
above  it  with  regai'd  to  age,  and  he  did  not  see  any 
reason  for  disturbing  that  conclusion.  With  regard  to 
the  upper  glacial  deposit,  the  deposit  on  top  of  the 
Mesozoic  sandstone,  the  rocks  in  the  clay  were  of  the 
same  lieterogeneous  character,  and  included  granites  and 
porphyries  which  were  stiiated,  and  had  all  the  general 
characteristics  of  a  glacial  deposit ;  but  there  was  one  very 
great  ditterence  between  the  tvfo  beds,  as  had  been  pointed 
(jut  by  Mr.  Cresswell.  The  lower  bed  lay  upon  a  surface 
that  had  liecn  smoothed  and  planed  apparently  by  the 
action  of  ice,  but  the  uppe}'  bed  lay  upon  a  sandstone  surface 
as  rugged  and  rough  as  a  mountain  peak;  and  in  the 
fractu]-es  wliich  were  found  in  this  rugged  suriace  hard  clay, 
very  much  like  a  boulder  bed,  and  rocks,  including  granites, 
had  been  jammed  down  hard,  and  presented  a  very  different 
appearau'ce  indeed  to  the  bed  which  lay  below  it  upon  the 
Silurian.  If  the  upper  bed  were  due  to  glacial  origiu, 
certainly  the  circumstances  were  very  different  to  those  of 
the  lower  bed.  No  heavy  mass  of  ice  had  ever  passed  over 
this  till,  because  if  it  had,  it  would  have  planed  all  the  soft 
sandstone  as  smooth  as  the  Silurian  had  been  planed  below 
it.  Therefore,  if  it  were  due  to  glacial  action,  the  till  and 
rocks  must  have  been  deposited  where  they  were  now  found 
by  the  thin  edge  of  the  glacier,  an  edge  which  had  no 
weight,  but  which  at  the  same  time  was  sufficieutl}'  strong 
to  cai-ry  a  burden  of  rocks  and  tip  then;  out  Iaterall3^  This 
was  a  possible  explanation.     It  might  also  have  been  caused 


Proceedings  of  the  Royal  Society  of  Victoria.     209 

by  a  veiy  small  glacier  indeed  from  some  steep  mountain 
close  by,  removed  by  erosion,  the  glacier  itself"  having  no 
v/oight  or  body,  but  able  to  bring  down  stones  and  tilt 
them  out.  This  was  a  very  strong  point  of  difference 
between  the  two  beds  of  so-called  glacial  till.  Then, 
with  regard  to  the  age  as.signed  to  the  upper  body  as 
being  Miocene,  tie  failed  to  see  that  there  was  any  evi- 
dence at  present  that  would  enable  them  to  assign  such  an 
early  age  to  the  bed.  Messrs.  Officer  and  Balfour  stated 
that  it  was  overlaid  by  a  Miocene  lava,  but  he  understood 
them  now  to  retract  that  statement,  and  to  sa}'  they  believed 
that  the  lava  was  the  recent  lava  which  was  found  all  about 
the  little  cavities.  Miocene  lava  was  found  there,  but  the 
Government  geologists  had  not  rej)resented  it  as  overlying 
the  Mesozoic  sandstones,  but  stated  that  it  was  intrusive 
and  pushed  its  way  through.  They  represented  it  as  pro- 
jecting from  below,  and  the  Mesozoic  sandstone  lying  upon 
it.  The  later  lava  was  distributed  all  over  the  country, 
and  lay  sometimes  on  the  Silurian,  and  sometimes  on  the 
Mesozoic  sandstones,  and  sometimes  upon  the  conglomerates, 
which  had  been  described  by  the  authors.  Therefore,  there 
was  no  evidence  that  he  could  see,  which  would  justify 
these  gentlemen  in  attributing  to  this  upper  boulder  clay 
the  age  which  they  had  given  to  it.  Their  second  pro- 
position was  to  the  effect  that  the  boulders  of  both 
these  epochs  were  due  to  land  ice  and  not  to  icebergs. 
With  regard  to  that,  the  giooves  and  strite  of  the  Silurian 
ran  from  north  to  south.  As  Mr.  Cresswell  had  pointed 
out,  that  might  be  produced  by  the  character  of  the  rock 
itself ;  but  if  there  were  grooves  and  strise  there,  they 
were  just  as  one  would  expect  to  find  them,  because  the 
highland  there  was  always  found  to  the  north  of  this  point, 
and  ice  travelling  from  the  mountain  cap  must  ha\*e  taken  a 
n(jrth  and  south  dii-ection.  Had  these  marks  been  produced 
by  the  grounding  of  icebergs,  one  would  expect  to  find  some- 
thing quite  different.  Icebergs  would  travel  first  of  all  upon 
the  general  trend  of  the  coast,  which  was  from  west  to  east. 
They  would  travel  with  the  currents  of  those  seas,  and  as 
the  prevailing  winds  in  this  locality  were  from  west  to  east, 
so  the  currents  were  from  west  to  east,  and  one  would 
expect  to  find  the  icebergs  travelling  with  the  currents  and 
with  the  winds  and  along  the  coast  line,  all  three  of  which 
ran  from  west  to  east.  Therefore,  if  they  produced  any 
strife  at  all,  theSe  stria?  would  run  from  west  to  east.     But 


270      P rucecdiiKjti  of  Ihe    lioi/ai  Soclely  of  Yiclorhi. 

it  was  known  that  icebergs  when  they  grounded  did  not 
produce  strict,  in  fact  could  not  do  so.  When  an  iceberg 
grounded  it  swung  round  on  its  heel,  and  if  it  produced  an_y 
marks  at  all  on  the  rock,  in  the  first  place  it  bruised  the 
rocks,  and  in  the  second  ]:)hice  the  rocks  which  had  been 
embodied  in  the  iceberg  and  touched  the  rocky  bottom 
in  the  bed  of  the  sea,  produced  marks  wliich  were  arcs  of 
circles.  Icebergs  never  made  straight  lines.  Then  there 
was  another  line  of  argument,  which  to  his  mind  entirely 
disposed  of  the  iceberg  theory.  The  Mesozoic  sandstone 
was  essentially  of  fresh  or  brackish  water  formation. 
The  only  fossils  found  in  the  Mesozoic,  were  two  fresh 
water  mollusca  and  the  vegetable  remains  of  ferns.  A])art 
from  this,  the  form.ation  liad  all  the  characteristics  of  sedi- 
mentary beds  which  had  been  formed  in  a  lake.  It  was 
known  to  most  geologists,  that  the  Mesozoic  sandstone  was  a 
fresh  water  lake  deposit.  Our  mountains  at  the  time  it  was 
formed  were  very  much  higher  than  they  are  now,  and  a 
series  of  lakes  were  formed  between  their  shelving  sides,  and 
as  the  lakes  got  tilled  up  with  the  sand  which  now  formed  the 
Mesozoic  sandstone,  the  water  rose  higher.'  There  was  very 
little  doubt  that  it  was  never  anything  else  than  a  shallow 
fresh  water  lake,  perhaps  of  considerable  dimensions.  He 
would  like  to  knovv  how  icebergs  were  going  to  float  in  fresh 
shallow  water.  An  icebeig  had  one  part  above  water  and 
eight  parts  below.  How  were  icebergs  to  float  ?  This  was 
not  a  marine  deposit,  and  although  2500  feet  in  thickness, 
we  might  depend  upon  it  this  Mesozoic  sandstone  had 
accumulated  gradually,  and  as  it  accumulated  at  the  bottom 
of  the  lake,  the  water  had  risen.  No  iceberg  ever  could 
have  floated  in  these  waters,  and  therefore,  in  his  opinion, 
no  icebergs  could  ever  have  caused  these  marks.  There 
was  also  another  important  piece  of  evidence  that  should 
not  be  disregarded.  Around  all  the  remains  of  all  these 
ancient  lakes  in  Victoria,  below  the  sandstone  bed  was 
found  a  bed  of  conglomerate.  This  was  apparently  the 
case  at  Bacchus  Marsh,  with  regard  to  the  conglomerate 
under  discussion.  The  officers  of  the  Government  Geological 
Survey  attributed  the  bottom  member  to  the  action  of 
the  water  on  the  coast.  Wherever  the  margin  of  this 
sandstone  was  found,  the  conglomerate  was  found  under 
it.  On  the  whole,  the  evidence  was  rather  favourable 
to  a  glacial  origin  for  the  bottom  deposits  at  Bacchus 
Marsh,  and  he  attributed  them  to  land  ice,*  not  to  icebergs. 


Proceedings  of  flie   Royal  Society  of  Vicfoiid.      271 

With  regard  to  the  third  proposition,  that  the  immersion 
of  the  Continent  sufficient  to  tioat  icebergs  would  reduce  the 
land  sui'tkce  to  such  a  small  area  that  it  would  iuive  a  mild 
insular  climate.  If  tliese  beds  were  due  to  a  deposit  in  the 
lakes,  they  were  not  due  to  immersion  in  the  ocean,  and  the 
argument  of  the  authors  fell  through,  because  the^v  assumed 
that  the  high-water  level  was  the  high-water  level  of  the 
ocean,  whereas  it  appeared  to  him  to  be  th(-  high-water 
level  of  various  fresh-water  lakes. 

Mr.  Dexis'ANT  said  he  would  allude  princi[)ail3'  to  the 
claim  made  by  these  gentlemen  for  a  post-Miocene  glacial 
epoch,  or  at  any  rate,  even  if  not  j)ost-Miocene,  some  portion 
of  the  Tei"tinrv  time,  which  would  include  the  Eocene, 
Miocene,  and  Pliocene.  Consequently,  if  there  were  glaciers 
in  Victoria  at  that  time,  the  climate  must  have  been  a  cold 
one,  for  it  would  be  impossible  to  have  a  glacier  with  the 
temperature  the  same  as  at  present.  If  we  started  with 
the  Eocene,  it  was  well  known  that  there  was  a  very  rich 
fauna  in  the  Eocene,  perhaps  the  richest  of  any  found  in 
any  part  of  tlie  world,  but  it  was  essentially  a  tropical 
fauna,  and  no  one  who  had  examined  the  launa  of  the 
Eocene  period  would  doubt  ibr  a  moment  but  that  he  was 
in  the  same  latitudes  as  the  West  Indies  and  the  Tropics 
generally.  Passing  from  the  Eocene  to  the  Miocene,  the 
climate  was  certainly  getting  cooler,  but  was  still  very 
much  warmer  even  than  our  present  climate.  The  shells 
indicated  a  climate  becoming  more  and  more  like  the 
present,  but  ver}'  far  indeed  from  being  a  glacial  one. 
Passing  to  the  Pliocene,  during  the  last  twelve  years,  two 
very  rich  marine  deposits  had  been  found  in  the  Pliocene, 
one  in  the  older  Plioceiie  near  Adelaide,  and  another  in  the 
west  of  our  own  Colony,  at  Limestone  Creek  ;  in  each  of 
these  there  was  a  rich  fauna.  The  climate  indicated  was 
slightly  colder  than  that  of  the  Miocene,  and  in  both 
deposits  was  found  a  large  number  of  living  shells.  In  the 
late  Pliocene  or  almost  Pleistocene  of  Victoria,  the  living 
shells  amounted  to  81  per  cent.,  but  they  were  not  those 
generally  found  on  the  present  shores  of  Victoria,  or  of 
Southern  Australia,  but  those  more  frequently  found  living 
in  the  northern  pai'ts  of  the  Continent.  Consequently,  at 
the  time  they  were  deposited  the  climate  was  warmer  than 
now,  and  approximated  to  the  climate  of  the  northern  parts 
of  the  Continent.     It  might  be  concluded  that  in  the  older 


272      Froceedin'js  of  fl/c  Ihiijal  Sociefy  of  Victoria. 

Tertiary  ])erio<l,  there  must  liave  been,  on  the  wl)(jle,  a 
warm  climate,  and  during  that  })eriod  it  would  be  impossible 
to  expect  that  any  glacial  phenomena  could  have  been 
produced.  Besides  this,  no  shells  had  been  found  that 
would  indicate  Arctic  conditions.  It  was  well  known  that 
in  the  glacial  till  of  Europe,  Arctic  shells  were  frequently 
found,  and  it  was  possible  to  trace  these  deposits  by  the  shells. 
Where  then  could  the  Tertiary  glacial  epoch  of  Victoria 
be  placed  ?  PalfPontolog•icali3^  there  was  no  room  for  it.  It 
might  perhajjs  be  mentioned  that  the  sea  was  certainly  close 
to  Bacchus  Marsh  during  Tertiary  times.  Undoubted 
evidence  of  this  had  been  given  by  Mr.  Reginald  Murray  in 
one  of  his  reports.  A  statement  had  also  been  made  that 
the  pebbles,  &c.,  which  had  been  found,  were  not  known  to 
exist  in  Victoria.  He  would  like  to  know  what  these  were, 
for  no  list  of  rocks  was  given  as  those  not  found  in  any 
other  part  of  the  colony.  Then  again,  amongst  these  rocks, 
granite,  schist,  felspar  and  sandstone  were  mentioned,  but  no 
mention  was  made  of  the  Tertiary  limestone  oi'  any  rock  of 
undoubted  Tertiary  age.  If  this  were  a  post-Miocene  or  late 
Tertiary  deposit,  he  thought  we  should  have  some  of  these 
rocks  amongst  those  which  had  been  transported. 

Mr.  James  Dunn  said  that  the  conglomerate  w*ljich  he 
regarded  as  of  glacial  origin  lay  at  the  base  of  the  coal  measures. 
If  the  conglomerates  that  he  described  were  the  same  as 
those  described  by  the  authors  of  the  paper,  it  was  out  of  the 
question  to  speak  of  roches  moutonnees.  In  fact  those  who 
had  passed  through  Bacchus  Marsh  would  have  observed  the 
rounded  appearance  of  the  liiils.  This  was  characteristic  tf 
the  Mesozoic  deposits  of  every  part  of  the  colon}- ,  and  was 
certainly  due  to  diluvial  action,  and  he  did  not  think  any 
weight  could  be  laid  upon  that  chaiucteristic  feature  of  the 
landscape  as  indicating  any  glacial  action  whatever.  He 
was  glad  the  matter  had  been  brought  forward,  and  the 
authors  of  the  paper  had  done  veiy  good  service  in  making- 
such  earful  observations,  which  would  enable  those  who 
wished  to  do  so  to  examine  the  spots  referi'ed  to  for 
themselves. 

Professor  Spencer  said  that,  twelve  days  since,  he  had 
gone  with  Mr.  Dunn  to  the  de))osit  he  had  described,  for  tlie 
purpose  of  being  shown  what  were  undoubtedly  roches 
moutonnees.  There  could  not  be  the  slightest  doubt  about 
the  presence  of  these  at  Derinal. 


Froeeedinijs  of  the  Roijal  iSoc'iefij  of  Vlcturla.     273 

Mr.  Pritchard  said  he  would  like  to  make  a  few  remarks 
on  the  diversity  of  opinion  as  to  the  age  of  these  beds.  Tlie 
Bacchus  Marsh  beds  had  been  originally  set  down  as  Tiiassic, 
and  the  coal  measures  in  Newcastle  and  in  the  neighbour- 
hood of  Sj'dtiey  were  originally  set  down  as  belonging  to  the 
Mesozoic  period,  so  that  originally  the  Bacchus  Marsh  sand- 
stones had  been  placed  on  a  lower  level  than  the  Newcastle 
coal  series.  xVt  the  present  time,  the  Newcastle  coal  series 
were  known  to  belong  to  the  carboniferous  age,  and  the  oidy 
fossil  remains  which  had  been  hitherto  found  in  the  Bacchus 
Marsh  sanstones  were  three  species  of  the  genus  Gangamoj)- 
teris.  The  genus  itself  had  been  found  in  the  coal  measures  of 
New  South  Wales,  in  connection  with  Glossopteris  and  othei- 
geneiu,  which  were  now  looked  upon  as  Mesozoic,  and  which 
had  always,  up  to  the  present  time,  l)een  looked  upon  as 
characteristically  Mesozoic,  but  having  been  found  together 
with  characteristic  Pakieozoic  plants  and  marine  fossils,  they 
were  looked  upon  now  as  an  extension  of  the  range  of  the 
genus  Glossopteri.s.  This  would  seem  to  point  to  the 
conclusion  that  the  Bacchus  Mar.sh  sandstones  might  belong 
to  the  carboniferous  age.  Some  authorities  looked  upon  the 
Bacchus  Marsh  sandstone  as  belonging  to  the  cai'boniferous 
period. 

Mr.  Dennaxt  said  he  only  referred  to  the  roches  mouton- 
nees  in  connection  with  the  claim  made  for  Tei'tiary  age.  He 
understood  that  Mr.  James  Dunn  placed  his  deposit  in  the 
cai'boniferous  era,  and  evidently  I'eferrecl  to  a  diftei'ent  epoch 
to  that  which  these  gentlemen  referred  to  when  they  spoke 
of  a  post-Miocene  glacial  epoch.  If  the  fauna  did  not 
indicate  necessarily  the  climate,  at  all  events  any  glacial 
epoch  that  might  have  occurred  during  Tertiary  times  nuist 
have  been  of  a  veiy  spasmodic  nature. 

Mr.  Cresswell  asked  whether  the  proposition  that  the 
upper  glacial  bed  was  a  post-Miocene  deposit  had  not  been 
withch-awn. 

Mr.  Officer  said  that  in  the  paper  it  had  been  stated 
that  this  bed  probably  belonged  to  the  Tertiary,  but  they 
had  not  attempted  to  assign  it  to  any  particular  era  in 
Tertiary  times.  In  fact  they  had  expressh^  stated  that  they 
were  unable  to  find  out  its  relation  to  the  Miocene  beds. 

Mr.  Dexxant  said,  that  being  the  case,  most  of  his 
remarks  need  not  have  been  made,  for  he  was  only  claiming 


274-      Proceediv(js  of  the  Royal  Socie/ij  of  Victoria. 

that  tliere  was  no  evidence  of  a  Tertiary  glacial  epoch  in 
Victoria  in  that  neighbourhood. 

Dr.  Dendy  said  there  seemed  to  he  one  as])ect  ot"  the 
question  that  had  not  been  touched  upon,  viz.,  its  bearing 
v)n  the  latest  glacial  theory  in  Europe  and  America.  lie 
believed,  according  to  this  theory,  the  glacial  epoch  was 
attributed  to  astronomical  causes,  and  it  was  a  remarkable 
fact  tliat  according  to  this  theory,  if  they  had  had  a  glacial 
epoch  in  the  northern  hemisphere,  it  followed  as  a  natural 
consequence  that  there  must  have  been  one  in  the  southern 
heniisj)here  alternating  with  it.  The  European  geologists, 
according  to  this  theory,  had  confidently  predicted  that  we 
should  find  in  Australia  evidences  of  a  Tertiary  glacial 
epocli.  Therefore,  he  trhouglit  it  probable  that  the  glacial 
evidences  discovered  by  Messrs.  Balfour  and  Officer  might 
be  Tertiaiy.  With  regard  to  the  question  of  climate  in 
connection  with  the  fauna,  tlie  ftict  that  trojiical  fauna  was 
found  in  some  of  these  Miocene  rocks  was  rather  a  strong 
proof  in  favour  of  the  glacial  theoij^  because  it  had  been 
shown  that  in  the  glacial  epoch  in  the  northern  hemisphere 
there  had  been  a  series  of  unusually  warm  periods  altern- 
ating with  a  series  of  unusually  cold  ones.  We  should 
therefore  expect  to  find  fauna  of  tropical  character  in 
connection  with  any  glacial  epoch  which  might  have 
iiappened  heie. 

Mr.  Officer,  in  reply,  said  that  with  regard  to  the  term 
"  Till,"  he  thought  it  a  very  good  term  indeed  to  api)ly  to 
any  deposit  which  could  be  shown  to  be  moraine  profonde. 
As  to  Mr.  Cresswell's  contention  that  till  was  not  ground 
moraine,  but  was  due  to  water  action,  the  boulders  having 
been  transported  by  icebergs,  that  was  a  theory  which 
would  not  bear  inspection.  As  to  the  roches  moutonndes, 
gentlemen  did  not  seem  to  be  quite  satisfied  as  to  the  genuine- 
ness of  tlie  article.  He  had  seen  many  examples  of  roches 
moutonne'es,  but  he  had  seen  very  few  better  specimens  than 
those  he  had  described.  With  regard  to  the  age  which  they 
had  assigned  the  lower  deposit,  they  had  stated  in  the  paper 
that  it  was  .simply  a  matter  of  probability.  Their  remarks 
had  been  based  on  the  fact  that  in  Europe  and  South 
Africa,  the  glacial  conglomerates  were  of  Permian  age. 
Mr.  Griffiths  had  stated  that  in  New  South  Wales  there 
was  no  break  from  the  Devonian  period  to  the  Mesozoic. 
Professor   David    had    stated    that    at   the    close    of    lower 


Froceedlmj'i  <>f  the   Roijal  Society  of    Victoria.      275 

(Carboniferous  times  there  was  a  distinct  break  in  tlie  flora, 
and  at  the  close  of  Perrao-carboniferous  times  there  was  also 
a  distinct  break  in  the  flora.  He  was  inclined  to  agree  with 
Mr.  Pritchard  with  regard  to  the  Mesozoic  sandstones.  They 
had  been  assigned  to  Mesozoic  age,  simply  on  the  evidence 
(if  tin-ee  species  of  Gangamopteris.  Seeing  that  these  occurred 
associated  with  Glossopteris  in  Permo-carboniferous  beds  in 
New  South  Wales,  and,  as  it  had  been  stated  by  Professor 
David  that  Gangamopteris  was  a  more  primitive  form  than 
Glossopteris,  it  would  almost  seem  as  if  these  beds  were  of 
an  earlier  age  than  Mesozoic.  Mr.  Griffiths  had  also  said 
he  did  not  think  that  an}^  ice  ever  passed  over  the  till  at 
the  quarry  where  the  Iractui-e  in  the  sandstone  occurred. 
On  liis  last  visit  to  the  same  quarry,  he  had  found  a  similar 
fracture  filled  with  till-bearing  striated  stones  at  a  much 
higher  level,  and  about  half-a-mile  further  up  the  creek 
there  was  a  great  thickness  of  this  till,  exposed  at  a  height 
between  sixty  or  seventy  feet.  If  that  had  been  accumulated 
under  a  glacier,  the  glacier  which  could  have  accumulated  it 
must  have  extended  much  further  down  the  valley,  and  it 
was  probable  it  did  over-ride  these  stones.  The  sandstone 
rock  was  very  soft,  and  woukl  not  show  strise.  It  had  been 
subjected  to  much  denudation.  The  rocks  also  dipped  at  a 
considerable  angle  up  to  35  degrees,  and  a  glacier  coming 
down  the  v.Jley  of  soft  sandstone  would  be  i-ather  likely  to 
fracture  them  and  give  them  a  rugged  appearance.  On  the 
whole,  he  did  not  think  anything  had  been  said  which 
would  lead  them  to  suppose  these  deposits  were  due  to 
anything  else  than  glacier  ice. 

Mr.  Steele  read  a  paper  on  "The  Conductivity  of  Cop])er 
Sulphate  Solutions." 

The  President  said  that  as  it  was  now  past  ten  o'clock, 
the  other  papers  would  be  held  over  till  the  next  meeting. 


Thur-sdai/,  Septntiher  Sth. 

The  President  (Professor  Kehnot)  in  the  chair. 

The  minutes  of  the  last  meeting  were  read  and  confirmed. 

Mr.  Hogg  signed  the  Roll  Book  and  was  introduced  to 
the  members. 


276      Pi-oceedingn  of  tlie   lioijal  Socleti/  of  Victoria. 

Mr.  Fi-edrick  Cliamljerlain  and  Mi-.  Alfred  Stilhvell  were 
elected  Members,  and  Mr.  A.  Purdie,  M.A.,  and  Mr.  W.  H. 
Steele,  M.A.,  Associates. 

The  PiiKSlDENT  welcomed  to  the  meeting  Proi'essoi' 
Haswell,  of  the  University  of  Sydney,  and  President  of  tlip 
Linnasan  Society  of  New  South  Wales. 

The  Librarian's  re|)ort  showed  that  1)8  new  volumes  had 
been  added  to  the  Library. 

Dr.  Barrett  read  a  papei-  on  "Snake-bite." 

In  reply  to  Mr.  Ellery,  Dr.  Barrett  paid  that  snake-bite 
was  usually  not  a  dangerous  affection  in  Victoria.  The 
natural  remedy  for  a  severe  bite  was  the  expulsion  of  the 
poison  by  downward  bandaging.  He  thought  a  great  deal 
was  to  be  said  in  favour  of  strychnine. 

Mr.  Elleuy  instanced  two  cases  of  recovery  from  snake- 
bite, but  in  one  case  the  man  was  subject  to  epilepsy  ever 
afterwards.  In  the  other  case,  injections  of  ammonia  had 
the  effect  of  causing  a  cure. 

In  reply  to  a  question  by  Dr.  Brett,  as  to  the  length  of 
time  taken  in  the  absorption  of  the  poison,  Dr.  Barrett 
said  it  was  impossible  to  state  how  long  it  would  take  for 
the  poison  to  take  effect  if  it  were  injected  into  the 
sub-cutaneous  tissue.  If  the  poison  were  shot  into  the 
vein,  no  bandaging  would  save  the  patient. 

Mr.  Hogg  considered  that  ammonia  and  strychnine  were 
not  antidotes,  strictly  speaking,  but  merely  had  the  effect 
of  making  a  patient  recover  from  a  comatose  state. 

Mr.  Fenton  said  that  in  Victoria  in  ten  years  there  were 
thirty-eight  deaths  from  snake-bite,  but  a  great  many  of 
those  were  insignificant  bites.  Not  more  than  about  six  of 
those  cases  were  over  twenty  years  of  age.  The  I'emainder 
were  all  young  children.  In  India  there  were  22,000  deaths 
from  snake-bite,  and  that  would  give  about  ninety  ])er  mean 
of  population. 

Dr.  Jamieson  said  that  no  statistics  wei'e  kept  of  the 
number  of  cases  of  snake-bite.  His  impression  was,  that 
real  cases  of  snake-bite  were  nmch  less  irequent  than  the 
supposed  cases,  and  the  symptoms  usually  presented  by 
those  su])j)osed  to  liax'e  been  bitten  were  not  due  to  snak(^- 


I'roceediiuis  of  the  Royal  Socieiy  of  Victoria.      Til 

l)ite  at  all.  The  stiyelniine  treatment  for  real  snake-bite  was 
not  at  all  irrational.  The  poison  could  not  be  extracted 
once  it  was  absorbed,  but  the  patient  could  be  kept  alive  by 
stimulants — alcohol,  ammonia,  or  strychnine.  The  last  was 
a  rational  remedy  if  cautiousN^  used,  and  if  the  patient  could 
lie  kept  alive  for  a  sufficient  length  of  time  the  poison  would 
be  thrown  off  b}^  the  kidneys,  or  might  be  rendered  inert  by 
the  action  of  the  liver.  The  injection  of  permanganate  of 
))Otash  was  to  his  mind  an  irrational  mode  of  treatment.  It 
interfered  with  the  circulation,  and  it  could  only  act  on 
the  poison  by  meeting  it  on  the  spot  were  it  existed  and 
destroying  it  in  a  chemical  way,  as  any  similai-  substance 
would"  be  destroyed  in  the  test  tube.  It  was,  therefore, 
haphazardous  ti-eatment,  as  it  was  uncertain  if  the 
permanganate  would  meet  the  poison.  The  time  occupied 
in  injecting  the  ]»ermanganate  might  be  utilised  to  better 
advaiitage  by  excision  of  the  bitten  part,  or  by  suction  or 
pressure. 

Mr.  Lucas  was  of  opinion  that  the  best  thing  to  do  was 
to  keep  the  [tatient  alive,  if  i)ossible,  by  sCimulants,  until 
the  ])roteid  was  ox\'dised. 

Professor  Haswkll  agreed  with  Dr.  Barrett  as  to  the 
i'allacy  of  statistics  on  this  subject.  He  was  of  opinion  that 
the  only  light  on  the  matter  was  to  be  obtained  by  means 
of  experiments  on  animals  carefully  conducted,  with  very 
careful  and  accurate  weighing  and  measuring  of  the  poison 
and  the  antidotes,  and  the  effects  of  both.  He  was  glad 
to  announce  that  there  was  a  prospect  of  some  results  being 
oVitained  from  experiments  of  this  nature.  Dr.  Martin, 
Demonstrator  (jf  Physiology  at  tlie  Sydney  University, 
was  engaged  in  researches  as  to  the  etfects  of  the  poison  of 
the  Australian  snakes. 

Mr.  Frost  had  some  experience  in  estimating  the  time 
occui)ied  by  the  poison  in  circulating  through  the  system. 
He  had  caused  a  tiger  snake  to  bite  a  rat,  and  the  rat 
was  dead  in  a  minute  and  a  half.  It  was  probable  that 
the  poison  was  injected  into  the  vein.  The  tiger  snake 
))ossessed  fully  three  times  more  poison  than  any  other 
snake.  He  had  seen  a  tiger  snake  emit  poison  at  the  third 
successive  bite  which  would  be  sufficient  to  kill  a  small 
animal.  It  was  difficult  in  experiments  with  small  animals 
to    estimate    the    amount    of  strychnine    necessary    to    kill 


278      Proceedhi(/s  of  the  Roycd  Societij  of  Victoria. 

the  animal.  After  injecting  sti-ychnine  into  a  rat,  the  rat 
recovered  from  snake-bite,  hut  afterwards  it  died  from  the 
effects  of  the  strychnine. 

Dr.  Barrett  said  that  in  the  Medical  Journal  for  187(5 
would  be  found  a  collection  of  replies  to  a  circular  issued  by 
Dr.  McCrae  to  medical  men.  He  got  a  return  of  253  cases 
of  .snake-bite,  and  10  per  cent,  had  died  without  any 
treatment.  It  was  interesting  to  note  that  Australian 
snakes  ejected  only  a  small  quantity  of  poison,  while  the 
Indian  snake  ejected  a  very  large  quantit}^  It  was  a 
question  whether  the  doses  were  in  proportion  to  the  size 
of  the  animals  met  with  by  the  snakes.  In  India  of  course 
the  animals  would  be  much  larger  than  those  met  with 
in  Australia.  He  agreed  with  the  opinion  expressed  by 
Piofessor  Hasweli,  that  careful  experimenting  in  the  labora- 
tory is  the  only  means  of  settling  the  question  as  to  the  size 
of  the  doses. 

Professor  Spencer  read  some  notes  on  "The  Structure  of 
the  Poison  Fang  in  certain  Australian  Snakes." 

Professor  Haswell  said  that  he  had  an  opportunity  of 
inspecting  Professor  Spencer's  sections,  and  there  could  be 
no  doubt  that  they  proved  his  deductions. 

A  paper  b}^  Mr.  A.  J.  Campbell,  F.L.S.,  on  "Three  Rare 
Species  of  Eggs,"  was  then  taken  as  read. 

Dr.  Dendy  read  a  paper  on  a  "  Synopsis  of  the  Australian 
Calcarea  Heterocoela,  with  a  pi-opused  Classification  of  the 
Group  and  Descriptions  of  some  New  Genera  and  Species." 

An  exhibition  of  specimens  followed,  and  the  meeting 
terminated. 


Thursdaij,   October  VMh. 

Mr.  White  (Vice-President)  m  the  Chair. 

The  minutes  of  the  last  meeting  were  read  and  confirmed. 

Dr.  Dendy  read  the  Librarian's  Report,  which  showed 
that  110  new  publications  had  been  added  to  the  Library. 

A  paper  by  Mr.  T.  S.  Hall,  M.A.,  on  "  Two  New  Tertiary 
Stylasterids,"  was  read  by  Mr.  Pritchard. 

Mr.  Pritchard  considered  that  the  paper  was  very 
interesting,  on  account  of  its  being  the  first  description  of 


Proceedliu/s  of  the  Roijal  Socletu  of  Victoria.     27i) 

Stj-lasterids  from  Australian  Tertiaries.  Since  looking  over 
Mr.  Hall's  paper,  he  had  found  numerous  Stylasterids  in  his 
own  collection. 

Dr.  Dendy  read  notes  on  "  The  Method  of  Reproduction 
of  Geonemertes  australiensis." 

Mr.  E.  F.  J.  Love,  M.A.,  exhibited  and  exjilained  Professor 
Rowlands'  Photogra})hs  of  the  Solar  Spectrum. 


Thursday,  Novemher  \Oth. 

The  President  (Professor  Kernot)  in  the  Chair. 

The  minutes  of  the  preceding  meeting  were  read  and 
coniirmed. 

Professor  A.  Liversidge,  F.R.S.,  was  elected  an  Honor;uy 
Member. 

Mr.  Steele  signed  the  book,  and  was  introduced  to  the 
Members  ;  Mr.  Isaac  Tip])ing,  C.E.,  was  nominated  as  an 
Associate. 

The  following  Members,  composing  the  Antarctic  Com- 
mittee, were  re-elected : — The  President,  and  Messi's.  Ellery, 
Rusden,  and  Griffiths. 

The  following  Members,  composing  the  Port  Phillip 
Biological  Committee,  were  re  elected : — Professor  Spencer, 
Dr.  Dendy,  Rev.  A.  W.  Cresswell,  and  Messrs.  Bale,  Lucas, 
McGiilivray,  and  Bracebridge  Wilson. 

The  Members  composing  the  House  Committee,  with  Mr. 
Blackett  as  Convener,  were  re-elected. 

Mr.  Love  presented  and  read  the  Report  of  the  Gravity 
Survey  Committee.*  The  President  and  Professors  Lyle 
and  Masson,  and  Messrs.  Elleiy,  White,  and  Love,  were 
re-elected  as  Members  of  Conmiittee. 

The  Preside^it  explained  that  the  apparatus  which  had 
been  used  by  the  Committee,  and  which  was  set  uj)  in  the 
Observator}',  was  in  perfect  order  and  fit  for  use. 

Mr.  W.  H.  Steele,  M.A.,  read  a  paper  on  "  Physical 
Constants  of  Tliallium." 

Mr.  Love  said  the  paper  was  a  valuable  contribution  to 
electrical  science.     It  liad  bi'ought  out  an  important  point, 

*  ]'ide  Supra,  p.  218. 


'280      Proceeding.s  of  the  Royal  Society  of  Victoria. 

iiaineh',  that  silver  wire  as  now  ()l)tained  is  very  mucli 
more  pure,  and  altogether  very  much  better  than  what  was 
supposed  to  be  pure  silver  twenty -five  years  ago.  The 
metal  J\latheson  worked  with  iu  making  the  experiments 
for  the  British  Association,  was  supposed  to  he  the  purest 
that  could  be  got,  and  up  to  the  present  that  had  never 
been  dis|)uted,  l>ut  after  Mr.  Steele's  work  there  need  be  no 
hesitation  in  asserting  that  silver  Aviro  had  been  produced 
in  this  Colony  of  a,  considerably  higher  degree  of  purity 
than  what  was  supposed  to  be  chemically  {)ure  some  years 
ago. 

Profi-snr  Masson  desired  to  know  what  steps  Mr.  >Steele 
took  tii  ascertain  the  degree  of  purity  of  his  Thallium.  He 
also  wished  to  know  how  far  Mr.  Steele's  coefficient  of  specific 
resistance  agreed  with  Mathesoii's  determiii;Uiou  of  some 
yeai's  ago. 

Ml-.  Steele  said  he  was  not  then  in  a  |:)osition  to  reply 
to  the  ([uestioti  relating  to  the  degree  of  purity  of  his 
Thallium.  As  to  the  conductivity-  of  Thallium,  the 
'•Dictionary  of  Uhenustry"  referred  to  it,  hut  the  results 
were  not  given  with  absolute  certainty.  According  to  the 
fig-nves  he  had  ([Uoted  in  his  pai)er,  the  conductivity  of 
Thalliuui  is  slightly  better  than  lead,  but  a  great  deal  worse 
than  tin. 

Mr.  Bl.yckett  considered  that  the  lead  used  by  Mr. 
Steele,  being  ordinary  commercial  lead,  would  have  gi\-en 
better  results  if  it  had  been  purified  by  re-crystallisation. 

Mr.  Steele  said  that  the  purity  of  the  lead  made  no 
difference.  He  merely  measured  the  specih'c  resistance  of 
lead,  and  he  might  have  used  an  alh\y. 

The  PiiESiDENT  said  that  the  elasticity  of  the  Constants 
was  of  interest. 

'vlr.  Steele  believed  that  the  jjarticular  specimen  of 
Thallnmi  he  had  used,  was  imported  into  the  Colony  for 
the  pur[)i).se  of  having  its  elasticity  tested,  and  that  had 
been  ilone  l>y  Mr.  William  Sutherhuid,  prioi'  to  his  use  of 
the  s{K:'ciu;cn. 

A  paper  (~»n  "'flu;  Lichenology  of  Victoria.  Part  I,"  by 
the  Rev.  F.  R.  M.  Wilson,  was  presented  by  Mr.  \V.  H. 
Arclier.  F. L.S.,  and  taken  as  read. 


Proceedliujs  of  thr   lii>i/nl  ^Societij  of   Vicforia. 


•281 


ill-.  D.  McAlpink  road  notos  i)re[)are(I  by  biuiself  and 
Mr.  P.  W.  Farmer,  M.B.,  Ch.  B.,  "On  a  Poisonous  Species 
(if  Honieria  found  at  Pascoe  Vale,  causing  the  death  of 
<'attle  and  otlier  animals  feedinti,'  ujiou  it." 

Mr.  Blackett,  ill  lej)!}'  to  the  President,  explained  that 
Mr.  Wilkinson,  with  his  assistance,  had  been  making  an 
iiivestigati()n  of  the  plant,  and  lie  was  contident  that  there 
was  no  trace  of  alkaloid  in  it.  They  did  not  liope  to 
isolate  any  particular  poison  from  the  plant.  The  corrosion 
of  the  mucous  membrane  of  the  cow's  stomach,  wdiich  was 
rt'ferred  to  in  the  paper,  ought,  if  it  existed,  to  be  detected 
from  the  extract.  He  di'ank  a  s[)Oonful  of  the  extract, 
but  it  did  not  produce  any  after  effect,  alth(»ugh  he  found 
it  disagreeable  and  nauseous 

Air.  McAlpine  said  that  the  symptoms  couhl  be  better 
explainetl  when  the  chemical  analysis  was  completed,  and 
a  second  paper  would  be  presented  when  that  was  done. 
Altiiough  two  extracts  had  lieen  experimented  on  with 
negative  results,  yet  there  was  the  fact  that  the  rabbits 
had  died.  The  corrosion  of  the  stomachs  of  the  animals 
that  had  eaten  the  [jlant  was  a  surprising  thing,  and  he 
hoped  that  chemical  anal3^sis  would  throw  light  on  the 
symptoms  recorded  in  the  notes. 

Mr.  White  asked  if  the  plants  were  more  dangerous  at 
certain  seasons  than  at  others. 

Mr.  McAlpine  said  that  in  West  Australia,  at  certain 
^'asons,  it  was  known  not  to  cause  death.  Cows  seemed  to 
I  void  the  plant. 

Mr.  White  said  that  might  prove  that  the  flowers  and 
not  the  leaves  were  poisonous. 

Mr.  McAlpine  thought  that  the  bulbous  part  contained 
the  Doison. 


Thitrsdaij,  Becember  blh. 
The  Pi'esident  (Professor  Keiixot)  occupied  the  chair. 

The  minutes  of  the  ])receding   meeting    were    read    and 
■ontirmeil. 

Mr.   Isaac  'J'lpping  was    ballotted   for  as  a  Member,  and 
Icclar.'d  duly  elected. 


282     Proceedings  of  the  Royal  Society  of  Victoria. 

Rev.  F.  R.  M.  Wilson  and  Mr.  P.  W.  Fanner  were  nomin- 
ated as  Members. 

The  President  said  that  Mr.  Lucas,  who  had  been  well 
known  to  those  present  for  many  years  past  as  a  member  ot 
the  Society,  and  a  writer  of  valuable  papers,  and  also  as  a 
Member  of  the  Council,  was  about  to  leave  the  Colony, 
having  accepted  the  headmastership  of  the  xs'ewingtou 
College  in  Sydney.  He  thought  that  ever}'  member  would 
agree  with  him  when  he  said  they  were  veiy  sorry  indeed 
to  lose  Ml-.  Lucas.  He  trusted  that  his  removal  to  Sydney 
would  be  a  step  in  the  direction  of  prosperity  and  emolument, 
and  in  the  name  of  the  Society  he  wished  Mr.  Lucas  every 
happiness,  prosperity,  and  success  in  the  new  sjjhere  which 
he  was  about  to  fill.  He  did  not  know  whether  he  would 
continue  his  connection  with  this  Society,  or  whether  his 
contributions  would  fall  into  the  hands  of  the  Ro3'al  Society 
of  New  South  Wales.  He  might,  perhaps,  be  ])ermitted  to 
express  a  liope  that  he  would  not  be  in  a  ver}'  great  hurry 
to  abandon  the  Royal  Society  of  Victoria  for  that  of  the 
sister  Colony. 

Mr.  Lucas  said,  I  thank  you  very  much,  Mr.  President 
and  Gentlemen,  for  the  very  kind  way  in  which  you  have 
taken  leave  of  me.  I  have  not  the  remotest  intention  ot 
severing  ni}'  connection  with  this  Society.  I  should  be  very 
sorry  indeed  to  lose  touch  with  the  many  friends  and  fellow 
workers,  the  advantage  of  whose  society  I  have  enjoyed 
during  the  last  ten  years.  I  have  every  confidence  in  the 
future  of  this  Society,  and  no  one  will  watch  its  progi-ess 
and  success  with  more  interest  and  pleasure  than  myself 

Dr.  Dendy,  the  Hon.  Librarian,  reported  that  77  publica- 
tions hail  been  received  since  the  last  meeting.  The 
liberality  of  the  Council  had  enabled  them  to  send  !)o 
complete  volumes  to  the  binder. 

Professor  Spencer  said  that,  according  to  the  rules  of  the 
Society,  the  Officers,  and  some  of  the  Members  of  the 
Council  i-etired,  but  were  eligible  for  re-election.  He  miglit 
say  it  was  no  use  nominating  Mr.  Sutherland  to  the  position 
of  Secretary  for  the  following  year,  as  he  was  about  to  leave 
the  Colony  for  an  extended  visit  to  Europe,  and  would  not  be 
able  to  serve  the  Society  for  the  next  three  years.  So  far 
as  he  (Professor  Spencei')  was  concerned,  he  did  not  want  to 
sever  his  connection  with  the  Council,  if  he  could  help  it. 


FroeeediiKjs  of  the  Roj/al  Soc/eti/  of  Victor ia.      2So 

He  woiiIJ  be  at  the  fii'st  meeting  next  year,  but  not  at  any 
of  the  subsequent  ones.  He  thought  it  miglit,  perhaps,  be 
lett  with  the  Council  to  appoint  someone  to  do  his  work 
while  he  was  away. 

The  President  thought  it  would  be  wise  to  fall  in  with 
Professor  Spencer's  request.  If  the  Society  gave  him  leave 
of  absence  to  visit  Europe,  he  would  still  remain  Secretary 
to  the  Society,  and  as  the  Society's  Secretary  might  be  able 
to  represent  the  Society  to  the  Societies  of  Europe  and 
America,  and  thus  do  useful  work. 

Dr.  Barrett  said  the  Medical  Society  had  a  custom  of 
empowering  all  distinguished  members  visiting  Europe  to 
represent  the  Society  at  all  scientific  gatherings.  A  written 
document  to  that  effect  was  given,  and  was  veiy  often  found 
iiseful  in  travelling.  He  tliought  it  the  very  least  this 
Society  could  do. 

Mr.  Hogg  moved,  "That  Professor  Spsncer  be  granted 
leave  of  absence  to  visit  Europe  and  America,  and  that  he 
be  empowered  by  a  suitable  letter  to  represent  this  Society 
at  the  meetings  of  other  Societies  and  scientific  bodies." 

Dr.  Barrett  seconded  the  motion,  which  was  carried. 

Mr.  White  .said  it  often  happened  that,  by  the  1st  Mai'ch, 
scarcely  one  nomination  had  been  received,  and  to  save  the 
Council  from  the  invidious  position  of  always  nominating 
themselves,  he  would  suggest  that  some  gentleman  present, 
who  was  not  a  Member  of  tlie  Council,  would  move  that  the 
old  Officers  and  retiring  Members  of  Council,  should  be 
re-elected  as  far  as  possible.  This  would  not  prevent  other 
nominations,  if  any,  being  received  before  the  1st  March. 

Mr.  Lucas  nominated  all  the  i-etiring  Members  of  Council 
who  were  eligible  for  re-election. 

Mr.  F.  A.  Campbell  seconded  the  nomination. 
Professor  SPE^X'ER  nominated   Mr.  Hogg  as  a  Member  of 
the  Council. 

Dr.  Dekdy  seconded  the  nomination. 

Mr.  White  nominated  Mr.  F.  A.  Campbell  as  a  Member 
of  the  Council. 

Mr.  Griffiths  seconded  the  nomination, 

Mr.  Griffiths  said  it  had  been  suggested  to  him  that  it 
would  be  only  a  giaceful  act  on  the  part  of  tlie  Society  to 


2N I      ProceeJii)>js  of  the   R(>>fal  Uncldy  of  Victoria. 

ackiiowk-dge  tlie  services  oi  Mr.  Sutberland  as  Secretary, 
and  he  begoed  to  move,  " 'J^hat  we  ]jlace  upon  the  minutes 
the  Society's  appreciation  of  the  services  of  Mr.  Sutherland, 
and  regret  that  lie  is  about  to  leave  us  for  some  time." 

Mr.  RusDKN  seconded  the  motion,  which  was  carried. 

A  pa[>er  entitled  "Some  Fallacious  Observations  on 
Sneezing''  was  read  by  Dr.  J.  W.  Bauiiett. 

The  PhksI!)i:.\t  said  that  having  done  a  good  deal  of 
sneezing  himself  during  the  past  fortnight,  since  the  grass 
had  become  dry  and  the  smell  of  hay  had  been  about,  his 
im|)ression  was  tliat  a  large  portion  if  not  the  v/hole  of  the 
blast  (-fair  came  through  the  mouth. 

Dr.  JamikS()N  said  that  like  most  people  he  had  been  led  by 
his  reading  to  think  that  sneezing  wasapurjiosive  act,  carried 
out  with  a  view  to  expelling  some  foreign  body  from  the  nose. 
Since  talking  to  Dr.  Barrett  about  it,  however,  he  had  made 
liUservations  for  himself,  and  had  conie  to  the  conclusion 
that,  as  a  matter  of  fact,  in  the  act  of  sneezing  the  air  was 
exploded  thr<mgh  the  mouth.  Still,  he  would  not  like  to 
say  it  vvas  always  so,  as  a  little  observation  amongst  children. 
led  one  to  lieiieve  that  a  good  blast'of  air  nmst  sometimes 
])ass  throiii'li  the  nose,  because  it  was  not  uncommon  to 
observe  a;i  ejection  of  mucus  following  too  soon  after  to  be 
the  result  of  exudation  in  consequence  of  the  act.  However, 
he  agreed  veiy  fully  with  what  Dr.  Barrett  had  said.  The 
explanation  usually  given  in  l)ooks  rested  on  the  a.ssuni[)tion 
that  such  striking  phenomena  could  not  be  without  purpose, 
and  as  coughing  had  such  a  clear  purpose,  it  was  easy 
and  natural  foi-  people  to  think  that  .sneezing  had  a  similar 
purpose.  Still,  Dr.  Barrett  seemed  to  be  pushing  the 
matter  to  the  other  extreme  when  he  said  that,  as  a  matter 
of  fact,  the  upper  part  of  the  nasal  passages  was  actually 
closed  off,  with  the  deliberate  intention  of  preventing  air 
going  through  the  nose  in  sneezing.  It  was  very  true  that 
many  reflexes  had  a  definite  object  in  view.  Winking 
protected  the  eye  ;  the  pupil  contracted  so  as  to  guard 
the  back  of  the  eye  from  the  sudden  entrance  of  strong 
light  ;  but  certainly  moj-e  striking  phenomena  than  even 
coughing  or  sneezing  did  occur  which  were  en.tirely  purpose- 
less, oi  ill  some  cases  worse  than  ])urposeless.  In  one's 
praeti:-e-,  one  soMKjtimes  came  across  the  ca;-e  of  a  child 
\rhicli   ha<i  sustained  a  severe  burn  on  the  skin,  and  some 


Pi'oceedinfjs  of  lite  Rof/al  Socletf/  of  Victoria.      285 

time  afterwards  it  mioht  happen  that  the  child  was  seized 
with  violent  general  >  convulsions.  The  child's  violent 
struofglings,  and  the  comatose  condition  which  followed,  ha<l 
no  effect  in  healing  the  burn  or  allaying  the  irritation.  We 
were  not,  therefore,  of  necessity  to  suppose  that  any  violent 
manifestation,  such  as  sneezing,  served  a  good  purpose. 
The  irritation  of  the  nose  which  usually  preceded  sneezing 
w«s  very  regularly  accompanied  with  watering  of  the  eyes, 
so  that  the  flow  of  the  water  from  the  eyes  would  largely 
contribute  to  the  washing  out  process.  The  question  was 
one  of  considerable  interest,  and  it  was  very  striking  to  iind 
that  nearly  all  the  best  authorities  on  physiology  seemed,  as 
lie  really  believed  after  thinking  the  matter  over,  to  bo 
manifestly  wrorg  in  the  interpretation  the}'  put  upon  it. 

Mr.  Hogg  said  that  several  instances  occurred  to  him 
which  seemed  to  show  that  a  blast  came  through  the  nose 
in  tlie  act  of  sneezing.  If  one  irritated  the  nose  with  snuff, 
immediately  one  sneezed  a  portion  of  the  snuff'  was  blown 
on  the  handkerchief,  and  it  was  very  obvious  that  the  snuff 
had  come  down  the  nasal  passages.  It  could  hardly  be 
supposed  that  it  had  been  washed  down,  it  certainly  seemed 
to  have  been  blown  down  by  a  blast  of  air.  If  one  were  so 
unfortunate  as  to  sneeze  when  in  the  act  of  drinking,  the 
liquid  would  be  forced  down  the  nasal  passage,  which 
seemed  to  indicate  that  there  was  some  blast  of  air  down 
the  nose.  Whether  it  was  entirely  through  the  nose,  or 
entirely  through  the  throat,  was  another  question.  Different 
]jeo])le  sneezed  in  different  waj's,  and  the  noise  produced 
was  different.  A  friend  of  his  in  sneezing  made  a  noise 
exactly  like  others  did  in  coughing ;  it  was  a  noise  not 
coming  from  the  mouth  or  nose,  but  from  the  pharynx,  as  in 
coughing.  Of  course  the  question  could  be  set  at  rest 
by  experimenting,  but  it  seemed  to  him  that  there  must 
unquestionably  be  a  blast  down  the  nostrils. 

Dr.  Dkxdy  said  he  was  very  susceptible  to  hay  fever,  as  it 
was  called,  but  he  thought  it  was  a  great  mistake  to  call  it 
hay  fever.  When  he  lived  in  London,  if  he  emerged  from 
the  underground  I'ailway,  the  sudden  strong  light  an<l 
the  dust  would  cause  him  to  sneeze  violently.  The  smell  of 
grass  would  bring  it  on,  and  so  would  the  dust  of  Melbourne, 
and  it  was  always  accompanied  with  violent  watering  of  the 
eyes  and  running  of  the  nose.  He  was  inclined  to  agree 
with  Dr.  Barrett. 


•28G      Proceed  in  < I  )<  of  the   Royal  Socief//  of  Victorio. 

Mr.  RUSDEN  said  lie  was  often  troubled  with  ])ersisterit 
Hiieezing.  Usually  the  Vilust  came  through  the  mouth,  but 
lie  had  tried  the  experiment  of  closing  the  mouth  tightly, 
when  an  explosion  through  the  nose  took  ])lace  and  expelled 
the  foreign  body  that  was  the  cause  of  the  trouble. 

Mr.  Griffiths  said  the  matter  had  been  brought  before 
liim  by  Dr.  Barrett  a  few  days  ago,  and  observations  that 
he  had  made  since  then  seemed  to  confirm  what  bad  been 
said  by  several  gentlemen,  for  it  appeared  that  the  air  bla.st 
did  proceed  through  the  mouth  and  not  through  the  nose. 
On  one  occasion,  finding  himself  engaged  in  a  series  of 
sneezes,  he  bad  determined  to  make  a  strong  efFoi't  to  close 
liis  mouth,  and  had  found  very  great  difficulty  indeed  in 
keeping  his  mouth  closed  ;  in  fact,  had  failed  to  make  a 
complete  closure.  So  far  as  he  had  succeeded  in  making  a 
])erfect  closure,  he  found  that  the  air  forced  its  way,  at  the 
cost  of  very  considerable  inconvenience,  through  the  nasal 
passage,  and  seemed  to  blow  some  mucus  from  the  glottis 
into  the  posterior  part  of  the  nose,  and  in  that  way  set  up 
considei-able  irritation  and  annoyance,  which  would  not 
have  been  experienced  liad  the  sneeze  been  allowed  to 
])ursue  the  normal  course.  It  had  occurred  to  him  that  the 
curtain  of  the  soft  palate  might  be  forcibly  closed  for  the 
ver}^  purpose  of  preventing  the  foreign  matter,  such  as 
mucus  lying  in  the  glottis,  from  being  shot  straight  out  of 
the  air  passage  of  the  throat  right  into  the  back  })ai-t  of  the 
nose,  on  account  of  the  irritation  that  would  be  set  up  by 
transferring  this  substance  from  the  glottis  to  the  nose. 
Such  a  transler  would  be  followed  by  considerable  incon- 
venience, giving  rise  to  furtlier  efforts  to  expel  from  the  back 
of  the  nose  that  which  had  no  right  to  be  there,  and  which 
would  not  be  there  but  for  attempting  the  \musual  course  of 
trying  to  stop  the  mouth. 

Mr.  Haig  said  that  his  experience  in  sneezing  was 
certainly  that  the  blast  came  entirely  through  the  mouth. 
In  fact  he  felt  that  the  nasal  passage  was  closed  both  before 
and  after  the  sneeze.  The  inspiration  before  the  sneeze  was 
always  through  the  moutli,  never  through  the  nose.  Some 
of  the  discharges  that  were  spoken  of  as  coming  through  the 
nose  might  come  afterwards  ;  as,  for  instance,  the  snufl". 
The  sneeze  might  come  first  with  a  blast  through  the  mouth, 
then  the  watery  discharge  would  take  place  through  the 
nose,  and  with  it  the  snuff. 


Proceedinfjs  of  the  Royal  Socletj/  of  Victoria.     2S7 

Mr.  Field  said  that  if  the  nose  were  held  wlien  about  to 
sneeze  the  noise  in  one's  ears  was  ahnost  deafening.  He 
had  found  that  by  placing  his  finger  upon  a  certain  nerve 
upon  the  top  of  the  head  when  about  to  sneeze,  the  sneeze 
would  be  stopped  altogether. 

Dr.  Barrett,  in  reply,  said  a  considerable  amount  of 
information  had  been  furnished  by  the  discussion.  Members 
on  the  whole  were  inclined  to  agree  with  him,  that  in 
uncomplicated  sneezing  the  bulk  of  the  air  passed  through 
the  mouth.  With  regard  to  the  remarks  of  Mr.  Hogg,  if 
any  voluntar}'  effort  wei'e  made,  the  statement  that  the  air 
passed  through  the  mouth  no  longer  held  good.  Unless  it 
was  a  pure  uncomplicated  sneeze,  it  was  impos.sible  to  reason 
with  any  degree  of  accuracy.  If  the  mouth  were  closed 
firmly  enough,  the  air  was  bound  to  go  though  the  nose,  or 
something  had  to  give  wa}'.  If  you  have  the  whole 
force  of  the  expiratory  muscles  at  work,  it  is  questionable 
which  is  strongest,  tlie  tissues  of  the  pharynx  or  the 
expiratory  muscles.  If  one  sneezed  with  a  closed  mouth, 
one  stood  a  good  chance  of  breaking  the  membrane,  for  a 
tremendous  pressure  would  be  put  upon  a  membrane,  which 
might  not  be  very  sound.  In  pure,  uncomplicated  sneezing, 
the  bulk  of  the  air  passed  through  the  mouth.  The  case  of 
children,  as  instanced  by  Dr.  Jamieson,  was  in  point.  Most 
children  had  enlargement  of  the  pharyngeal  tonsil,  which 
gave  rise  to  the  loud  breathing  and  the  bulk  of  the  ear 
troubles  in  children.  When  this  was  the  case,  one  could  no 
longer  reckon  upon  the  air  going  the  proper  way.  He  did 
not  think  the  inconvenience  spoken  of  by  Mr.  Griffiths  was 
due  to  expulsion  of  mucus  from  the  glottis.  Under  the 
circumstances  mentioned,  Mi-.  Griffiths  had  got  his  palate 
inverted  in  part,  and  then,  his  palate  acting  feebly,  a  certain 
amount  of  mucus  dro[)ped  down  into  the  pharynx.  With 
regard  to  hay  fever,  wliat  was  called  hay  fever  really  meant 
abnormal  sensibility  of  the  aperture  of  the  no.se,  and, 
consequently,  unusual  liability  to  stimulant.  Thus,  a 
stimulant  which  would  produce  practically  no  result  in  one 
person  with  normal  sensibility,  would  produce  an  abnormal 
result  in  a  person  suffering  from  hay  fever,  and  [)roduce,  not 
only  a  flow  of  water  from  the  nose,  but  also  from  the  eyes. 
T''he  last  speaker  had  referred  to  the  stoppage  of  the  sneeze 
by  a  pressure  on  a  nerve.  A  sneeze  could  be  sto[)ped  in  a 
score   of    ways.       How    it    was   done    would    be    rather   a 


288      Proceedivj/s  of  the  Ixoijid  Society  of  Vidoria. 

formidable  niattei'  to  discus.s  (yH-liand,  but  the  additional 
stimulus  set  up  V)y  another  nerve  would  destroy  the  ett'ect 
of  the  stimulus  imparted  liy  the  tirst.  Thus,  a  |>istol  shot, 
fired  alongside  a  person  about  to  sneeze,  would  pi-obably  |))it 
an  end  to  the  sneeze. 

Dr.  Dendy  then  read  a  note  on  a  "  New  S))eeies  of  Leu- 
cosolenia  from  the  Neighbourhood  of  Port  Phillip  Heads." 

The  meeting  then  terminate^]. 


LAWS. 

Amended  to  December,  1892. 


I.  Tlie  Society  shall  be  called  "The  Royal  Society  Name. 
of  Victoria." 

II.  The    Royal    Society  of   Victoria  is  founded  for  oiyeots. 
the  advancement  of   science,   literature  and    art,  with 
especial  reference  to  the  development  of  the  resources 

of  the  countr}^ 

III.  The  SocietN'  shall  consist  of  Ordinary  Members  Mombeisand 
residing  vi^ithin  ten  miles  of  Melbourne  ;  Country  ^^"•"^'■'*'^'^- 
Members  residing  be3^on(l  that  distance  :  Life  Members 

(Law  XXV),  Honorary  Members  (Law  XXIV), 
Corresponding  Members  (Law  LII),  and  Associates 
(Laws  XXV,  XXVI,  and  LIII),  ail  of  whom  shall  be 
elected  by  ballot. 

IV.  His    Excellency  the    Governor  of   Victoria,  for  Patron, 
the  time  being,  shall  be  invited  to  accept  the  office  of 
Patron  of  the  Society. 

V.  There  shall  be  a  President,  and  two  Vice-Presi-  officers. 
dents,    who,    with    twelve    other    Members,    and    the 
following  Honorary  Officers,  viz..  Treasurer,  Librarian, 

and  two  Secretaries  of  the  Society,  shall  constitute  the 
(Council. 

V[.  The  Council  shall  have  the  management  of  the  Management. 
atfai)-s  of  the  Society. 

VII.  The  Ordinary  Meetings  of  the  Society  shall  be  o.dhiary 
held    once    in    every  month  during  the  Session,  from       ^«*>"gs. 
March    to     December    inclusive,    on    days    fixed    and 
subject  to  alteration  by  the  Council  with  due  notice. 

VIII.  In  the  second  week  in  March,  there  shall  be  Annual  General 
an  Annual  General  Meeting,  to  receive  the  report  of    "  ''^  "'^^• 
the  Council,  and  elect  the  Officers  of  the  Society  for 

the  ensuing  year. 

IX.  All    Office-bearers    and     Members    of    Council  iiethementof 
except    the  .six  junior  or   last  elected  Members,  shall 
retire  from  office  at  the  Annual  General   Meeting    in 
March.     Should  a  senior  Member's  seat  become  vacant 

U 


Ortioer 


290    Proceed in<i.^  oj  Royal  Society  of   Victoria. 


Votes  required. 


Address  by  the 
President. 


Duties  of 
Presidei 


in  the  couise  of  the  year,  it  shall  be  held  by  his 
successor  (under  Law  XI 11)  as  a  senior  Member,  who 
shall  retire  at  the  next  Annual  General  Meeting.  The 
names  of  such  Retiring  OHicers  are  to  be  announced 
at  the  Ordinary  Meeting  in  December.  The  Officers 
and  Members  of  Council  so  retiring  shall  be  eligible  foi' 
the  same  or  any  othei-  office  then  vacant. 

X.  The  President,  Vice-Presidents,  Treasurer,  Secre- 
taries, and  Librarian  shall  be  sepai'ately  elected  by 
ballot  (should  such  l)e  demanded),  in  the  above-named 
order,  and  the  six  vacancies  in  the  Council  shall  then 
be  filled  up  togethe)'  by  ballot  at  the  General  Meeting 
in  March.  Those  member-s  only  shall  be  eligible  for 
any  office  who  have  been  pro]:)osed  and  seconded  at  the 
Ordinary  Meeting  in  December,  or  b>  letter  addressed 
to  one  of  the  Secretaries,  and  received  by  him  before 
the  1st  March,  to  be  laid  before  the  Council  Meeting- 
next  before  the  Annual  Meeting  in  March.  The 
nomination  to  any  one  office  shall  be  held  a  nomina- 
tion to  any  office,  the  election  to  which  is  to  be 
subsequently  held.  No  ballot  shall  take  place  at  any 
meeting  unless  ten  members  be  present. 

XL  No  Member,  whose  subscription  is  in  arrear. 
shall  take  part  in  the  election  of  Officers  or  other 
business  of  the  meeting. 

XIL  An  address  shall  be  delivered  b}'  the  President 
of  the  Society  at  eithei-  a  Dinner,  Conversazione,  or 
extra  meeting  of  the  Society,  as  the  Council  may 
determine  in  ea.ch  year. 

XIII.  If  any  vacancy  occur  among  the  Officers, 
notice  thereof  shall  be  inserted  in  the  summons  for  the 
next  meeting  of  the  Society,  and  the  vacancy  shall  be 
then  filled  up  by  ballot. 

XIV.  The  President  shall  take  the  chair  at  all 
meetings  of  the  Society  and  of  the  Council,  and  shall 
regulate  and  keep  order  in  all  their  proceedings  ;  he 
shall  state  questions  and  pi-opositions  to  the  meeting, 
and  re])ort  the  result  of  })allots,  and  carr^'  into  efi'ect 
the  regulations  of  the  Society.  In  the  absence  of  the 
President,  the  chair  shall  be  taken  l)y  one  of  the  Vice- 
Presidents,  Treasurer,  or  Ordinary  Member  of  Council, 
in  order  of  seniority. 


Laivs.  291 

XV.  The    Treasurer    may,    imuiediatel}'    after    lii.s  Duties  of 
election,  appoint  a  Collector  (to  act  dnrini^'  ]»leasare), 
subject   to  the   approval   of  the    Council  at   its    next 
meeting.     The  iluty  of  tiie  Collector  shall  be  to  issue 

the  Treasurer's  notices,  and  collect  subscriptions.  The 
Treasurer  shall  receive  all  moneys  ])aid  to  the  Societ}', 
and  shall  deposit  the  same  before  the  end  of  each 
month  in  the  bank  approved  by  the  Council,  to  the 
credit  of  an  account  opened  in  the  name  of  the  Royal 
Society  of  Victoria.  The  Treasurer  shall  make  all  pay- 
ments ordered  by  the  Council  on  receiving  a  written 
authoiity  fiom  the  chaiiinan  of  the  meeting.  All 
cheques  shall  be  signed  by  himself,  and  countersigned 
by  one  of  the  Secretaries.  No  payments  shall  be  made 
except  by  cheque,  and  on  the  authority  of  the  Council. 
He  shall  keep  a  detailed  account  of  all  receipts  and 
expenditure,  present  a  report  of  the  same  at  each 
Council  meeting,  and  prepare  a  balance-sheet  to  be  laid 
before  the  Council,  and  included  in  its  Annual  Report. 
He  shall  also  produce  his  books  whenever  called  upon 
to  do  so  by  the  Council. 

XVI.  The  Secretaries  shall  share  their  duties  as  they  Dutiesof 
may  find  most  convenient.  One  or  other  of  them  shall 
conduct  the  correspondence  of  the  Society  and  of  the 
Council,  attend  all  meetings  of  the  Society  and  of  the 
Council,  take  minutes  of  theii'  proceedings,  and  enter 
them  in  the  proper  books.  He  shall  inscribe  the  names 
and  addresses  of  all  Members  and  Associates  in  a  book 
to  be  kept  for  that  purpose,  from  which  no  name  shall 
be  erased  except  by  order  of  the  Council.  He  shall 
issue  notices  of  all  meetings  of  the  Society  and  of  the 
Council,  and  shall  have  the  custody  of  all  papers  of 
the  Society,  and,  under  the  direction  of  the  Council, 
superintend  the  jirinting  of  the  Transactions  of  the 
Society. 

XVII  The  Council  shall  meet  on  any  day  within  one  .Meetings  of 
week  before  every  Ordinary  Meeting  of  the  Society. 
Notice  of  such  meeting  shall  be  sent  to  every  Member 
at  least  two  days  previou.sly.  No  business  shall  be 
transacted  at  any  meeting  of  the  Council  unless  five 
Members  be  present.  Any  Member  of  Council  absent- 
ing himself  from  three  consecutive  meetings  of  Council, 
with<jut  satisfactory  explanation  in   writino-,  shall  be 

U^2 


Secretaries. 


Council. 


292     Proceediuf/s  of  Roijid  Sociefu  of  Victoria. 


Special  Meet! 
of  Ccmnoil. 


dliecial  Gem 
Meetings 


Annual  Repi 


Expulsion  I 
Members 


Election  of 
Members  iiuil 
Associates. 


considered  to  have  vacated  his  ojiice,  and  the  election 
of  a  Member  to  till  his  place  shall  ite  proceeded  with  at 
the  next  Ordinaiv  Meeting  of  Members,  in  accordance 
with  Law  XIII.  ^ 

XVIII.  One  of  the  Secretaries  shall  call  a  Special 
Meeting  of  Conncil  on  the  authority  of  the  President  or 
of  three  members  of  the  Conncil.  The  notice  of  such 
meeting  sliall  specify  the  object  for  which  it  is  called, 
and  no  other  business  shall  be  entertained. 

XIX.  The  Council  shall  call  a  Special  Meeting  of  the 
Societ}^,  on  receiving  a  requisition  in  writing  signed 
by  twenty-foui-  members  of  the  Society,  s]^ecifying  the 
purj)ose  for  which  the  meeting  is  retpiired,  or  upon  a 
resolution  of  its  own.  No  other  business  shall  be  enter- 
tained at  such  Meeting.  Notice  of  such  meeting,  and 
tiic  purpose  for  which  it  is  summoned,  shall  be  sent  to 
every  Member  at  least  ten  days  before  the  meeting. 

XX.  The  Council  shall  annually  prepare  a  Report 
of  the  Proceedings  of  the  Society  during  the  past  year, 
embodying  the  Balance-sheet,  duly  audited  by  two 
Auditors,  to  l)e  ajjpointed  for  the  year  at  the  Ordinary 
Meeting  in  December,  exhibiting  a  statement  of  the 
present  position  of  the  Society.  This  Report  sliall  be 
laid  before  the  Society  at  the  Annual  Meeting  in  March, 
No  paper  shall  be  read  at  that  meeting. 

XXI.  If  it  shall  come  to  the  knowledge  of  the 
Council  that  the  conduct  of  an  Officer,  a  2vlember,  or 
an  Associate  is  injurious  to  the  interest  of  the  Society, 
and  if  two-thirds  of  the  Council  present  shall  be 
satisfied,  after  opportunity  of  defence  has  been  afforded 
to  him,  that  such  is  the  case,  it  may  call  upon  him  to 
resign,  and  shall  have  the  power  to  expel  him  from  the 
Society,  or  remove  him  from  any  office  therein  at  its  dis- 
cretion. In  every  case,  all  proceedings  shall  be  entered 
upon  the  minutes. 

XXII.  Every  candidate  for  election  as  Member  or  as 
Associate  shall  be  proposed  and  seconded  by  Members 
of  the  Society.  The  name,  the  address,  and  the 
occupation  of  every  candidate,  with  the  names  of  his 
proi)oser  and  of  his  seconder,  shall  be  comnumicated  in 
writing  to  one  of  the  Secretaries,  and  shall  be  read  at  a 
meeting  of  Council,  and  also  at  the  following  meeting 


Laivs,  293 

of  the  Society,  and  the  ballot  shall  take  place  at  the 
next  following  Ordinary  Meeting  of  the  Society.     The  votes  required  to 
assent  of  at  least  five-sixths  of  the  number  voting  shall    '""'"  *' 
be  requisite  for  the  admission  of  a  candidate. 

XXIII.  Every    new    Member    or    Associate    shall  Members  shaii 
receive  due  notice  of  his  election,  and  be  supplied  with    ^'s^^  !'»"'«• 

a  copy  of  the  obligation*  together  with  a  copy  of  the 

Laws  of  the  Society.    He  shall  not  be  entitled  to  enjoy 

any  privilege   of  the   Society,  nor  shall  his  name  be 

printed  in  the  List  of  Members  until  he  shall  have  paid 

his  admission   fee   and   first  annual   subscription,  and 

have  returned  to  the  Secretaries  the  obligation  signed 

by   himself     He    shall,    at   the    first    meeting    of   the 

Society  at  which  he  is  present,  sign  a  duplicate  of  the 

obligation  in  the  Book  of  the   Laws  of  the  Society, 

after  which  he  shall  be  introduced  to  the  Society  by 

the  Chairman.     No  Member  or  Associate  shall  lie  at  conditious  of 

liberty  to  withdraw  from  the  Society  without  previ-     ^^signaion 

ously  giving  notice  in  writing  to  one  of  tlie  Secretaries 

of  his  intention  to  withdraw,  and  returning  all   books 

or  other  property  of  the    Society  in    his    possession. 

Members  and  Associates  will  be  considered  liable  for 

the  payment  of  all  subscriptions  due  from  them  up  to 

the  date  at  which  they  give   written    notice  of  their 

intention  to  withdraw  from  the  Society. 

XXIV.  Gentlemen  not  resident  in  Victoria,  who  are  Honorary 
distinguished  for  their  attainments  in  science,  literature,       *""  ^"" 
or  art,  may  be  |)ru})Osed  for  election  as  Honoi-ary  Mem- 
bers, on  the  rectjmmendation  of  an  absolute  majority  of 

the  Council.  The  election  shall  be  conducted  in  the 
same  manner  as  that  of  Oidinary  Members,  but  nine- 
tenths  of  the  votes  must  be  in  favour  of  the  candidate. 

XXV.  Ordiriar}^  Members  of  the  Society  shall  pay  suUscriptions 
two  guineas  annually.  Country  Members  and  Associates 

shall  ])ay  one  guinea  annually.     'J'hose  elected  after  the 

*  The  obligat^cu  referred  to  is  as  follows  : — 

EoYAL  Society  or  Victoeia. 
I,  the  nndersigucJ.  do  hereby  engage  that  I  will  endeavour  to  promote 
the  interests  and  welfare  of  the  Itoyal  Society  of  Victoria,  and  to 
observe  its  laws,  as  long  as  I  shall  remain  a  Member  or  Associate 
thereof. 

(Signed) 
Address 
Date 


294?    r roceed higs  of  Royal  Sochdy  of  Victoria. 


Life  Member- 
ship. 


Entrance  fees, 
Arc. 


Duration  of 
Meetings. 


Order  aud  mo 
of  conductii 
tlie  busiiies;^ 


first  of  July  shall  pay  only  half  of  the  subsci'iptiori  for 
the  current  year.  Ordinary  Menilters  nia}-  compound 
for  all  annual  subscriptions  of  the  ciurent  and  future 
years  by  paying  ^21 ;  and  Country  Members  may  com- 
pound in  like  manner  by  p.i^ying  £10  10s.  Any  Country 
Member  having  compounded  lor  his  subscri[)tion,  and 
eon-jing  to  reside  within  ten  miles  of  Melbourne,  must 
pay  either  the  balance  £10  10s.  of  the  Ordinary'  Member's 
com)n)sition,  or  one  guinea  annually  while  he  resides 
within  ten  miles  of  Melbourne.  The  subscri])tions 
shall  be  due  on  the  1st  of  January  in  every  year.  At 
the  commencement  of  each  year  there  shall  be  hung- 
up in  the  Hall  of  the  Society  a  list  of  all  Members  and 
Assi;c!ates,  ujion  which  the  ])ayment  of  their  subscrip- 
tion as  made  shall  be  entered.  During  Jul}',  notice 
shall  be  sent  to  all  Meuibers  and  Associates  still  in 
ari-eai's.  At  the  end  of  eiich  year,  a  list  of  those  who 
liave  not  paid  their  subscriptions  shall  be  prepared,  to 
bo  considered  and  dealt  with  by  the  Council. 

XXVI.  Newly-elected  Ordinary  find  Country  Mem- 
bers shaM  pay  an  entrance  fee  of  two  guineas,  in  addition 
to  the  subscription  ibr  the  curi'ent  year.  Honorary 
Memliers,  Corresponding  Members  and  Associates 
shall  not  be  I'eijuired  to  ]:»a,y  any  eii trance  fee  H'  the 
entrance  fee  and  subscription  be  not  paid  within  one 
month  of  the  notitication  of  election,  a  sect^nd  notice 
shall  be  sent,  and  if  payment  be  not  made  within  one 
month  from  the  second  notice,  the  election  sliall  lie  void. 
Associates,  on  seeking  election  as  Oixlinary  or  Coiuitry 
MemV»ei-s,  shall  comply  with  all  the  forms  prescribed 
for  the  election  of  Members,  and  shall  pay  the  enti'ance 
fee  prescribed  <ibove  of  Ordinary  or  Countr}'  Members 
respectively. 

XX VII.  At  tlie  Ordinary  Meetings  of  the  Society 
the  chair  shall  be  taken  ])unctiuilly  at  eight  o'clock, 
and  no  new  business  shall  be  taken  after  ten  o'clock. 

XXVIII.  At  the  Ordinary  Meetings  business  shall 
be  transacted  in  the  follo\ying  order,  imless  it  be 
specially  decided  otherwise  by  the  Chairman  : — 

Minutes   of  th(^    preceding    meeting    to    be    read, 
jimendiMl  if  incoi'rect.  and  ci>ntirmed. 


Laws.  295 

New    Members   and   Associates    to    enroll    their 

names,  and  be  introduced. 
Ballot    for    the    election    of    new    Members    or 

Associates. 
Vacancies  among   officers,  if  any,  to  be  filled  up. 
Business  arising  out  of  the  minutes. 
Communications  from  tlie  Council. 
Presents  to  be  laid  on  the  table,  and  acknowledged. 

Motions,  of  which  notice  has  been  given,  to  be 
considered. 

Notice  of  motion  for  the  next  meeting  to  be  given 
in  and  read  by  one  of  the  Secretaries. 

Papers  to  be  read. 

XXIX.  No  stranger  shall  speak  at  a  meeting  of  the  stranger. 
Society   unless    specially   invited    to    do    so    by   the 
Chairman. 

XXX.  Every    paper     before     being    read    at    any  Papera  to  be  first 
meeting  mast  be  submitted  to  the  Council.  comlciL'* 

XXXI.  The    Council   may  call  additional  meetings  Additional 
whenever  it  may  deem  it  necessiiry  to  do  so.  ^^  "'^^' 

XXXII.  Every  Member  may  introduce  two  visitors  visitors, 
to   the  meetings  of  the  Society  by  orders  signed  by 
himself. 

XXXIII.  Members  and  Associates    shall   have   the  Members  may 
privilege   of  reading   before    the   Society   accounts  of    '*'^*  papeis. 
experiments,  observations,  and  researches  conducted  by 
themselves,  or  original  papers,  on  subjects  within  the 

scope  of  tlie  Society,  or  descriptions  of  recent  dis- 
coveries, or  in\entions  of  general  scientific  interest. 
No  vote  of  thanks  to  any  Member  or  Associate  foi'  his 
paper  shall  be  proposed. 

XXXIV.  If  a    Member  or  Associate    be  unable  to  or  depute  oiher 
attend  for  the  purpose  of  reading  his  paper,  he  may 
delegate  to  any  Member  of  the  Society  the    reading 

thereof,  and  his  right  of  reply. 

XXXV.  Any     Member    or    Associate    desirous    of  Members  must 
reading  a  paper,  shall  give  in  writing  to  one  of  the    thlir"pape'is" 


2i)(;     ['rocecdiiKjs  of  lioijal  Soclefij  of  Victoria. 

Seci'etai'ies,  ten  clay.s  before  the  meeting  at  which  he 
desires  it  to  be  I'ead,  its  title  and  the  time  its  reading 
will  occupy. 

Papers  by  XXX VJ.    The    Council  may  tor  an 3'  special  reason 

laiigeis.  permit  a  paper  such  as  is  described  in  Law  XXXIII, 
not  written  by  a  member  of  the  Society,  to  be  read  by 
one  of  the  Secretaries  or  other  Members. 

Papers  beioug  to      XXXVII.    Evcry  papei'  read  befn-e  the  Society  shall 
the  Society.     |^^  ^^^^  y)roperty  thereof,  and  immediately  after  it  has 
been  read  shall  be  delivered  to  otie  of  the  Secretaries, 
and  shall  remain  in  his  custody. 

Papers  must  be        XXXVIII.    No    paper    shall    be    read    before    tlie 

original.  Socict^'    or    published     in     the     Ti-ansactions     unless 

approved  by  the  Council,  and  unless  it  consist  mainly 

of  origmal  matter  as  regards  the  facts  or  the  theories 

enunciated. 

Couuciiraay  XXXIX.    The  Couticil  may  refer  an\' j:)aper  to  any 

Mfmberr'^*"  Member  oi-  Members  of  the  Society,  to  re]X)rt  upon  the 
desirability  of  printing  it. 

Rejeote.i  XL.    Should    tlic    Couucll    dccidc    not    to  publish  a 

reTunied  ^^     paper,  it  shall  be  at  <jnce  returned  to  the  author. 

Members  may  XLI.    The  author  of   any  paj)er  which  the  Council 

tS pipe's"^  has  decided  to  publish  in  the  Ti'ansactions  may  have 

tifty  copies  of  his  paper  on  giving  notice  of  his  wish 

in  wilting  to  one  of  the  St;cretaries,  and  any  further 

number  on  paying  the  extia  cost  thereof 

Members  and  XLIl.    Every    Member    and    Associate    whose    sub- 

have'^Trans*"    scription  is   uot  ill   aiTear,  and    eveiy  Honorary   and 
actiouB.  Corresponding  Member  is  entitled  to  receive  one  copy 

of  the  Transactions  of  the  Society  as  published.  Newly- 
elected  Memljers  shall,  on  payment  of  their  entrance-fee 
and  subseri[)t,ion,  receive  a  copy  of  the  volume  of  the 
Transactions  last  published. 

Property.  XLIIl.    Evcry  book,  pamphlet,  model,  plan,  drawing, 

specimen,  preparation,  or  collection  presented  to  or  pur- 
chased by  the  Society,  shall  be  kept  in  the  house  of  the 
Societ}^ 

Library.  XLIV.    The  Library  shall  be  open  to  Members  and 

Associates  of  the  Societj-,  and  the  public,  at  such  times 
and  under  such  regulations  as  the  Council  may  deem  fit. 


Laws.  297 

XLV.    The  legal  ownei'ship  of  the  ))i'operty  of  the  Le^ai  owneMiiii. 
Society  is  vested  in  the  President,  the  Vice-Presidents,     ^'<' i"'"i'e''^>' 
and  tlie  Treasurer  for  the  time  being,  in  trust  for  the  use 
of  the  Society  ;  but  the  Council  shall  hav^e  full  control 
over  the  expenditure  of  the  funds  and  management  of 
the  property  of  the  Society. 

XLVI.    Every  Committee  appointed  by  the  Society  committbe.s 
shall  at  its  first  meeting  elect  a  Chairman,  who  shall    chaUmau 
subsequently  convene  the  Committee  and  briiig  up  its 
report.     He  shall  also  ol)tain  from  the  Treasurer  such 
grants  as  wv^y  have  been  voted  for  the  purposes  of  the 
Committee. 

XLVII.  All  Committees  and  individuals  to  whom  Heiwrt  before 
any  work  has  been  assigned  by  the  Society  shall  pre- 
sent to  tlie  Council,  not  later  than  the  1st  of  November 
in  each  year,  a  report  of  the  progress  which  has  been 
made ;  and,  in  cases  ^here  grants  of  mone}^  for  scientific 
purposes  have  been  entrusted  to  them,  a  statement  of 
the  sums  which  have  been  expended,  and  the  balance 
of  each  grant  which  remains  unexpended.  Every  Com- 
mittee shall  cease  to  exist  at  the  November  meeting, 
imless  then  re-appointed. 

XLVIII.  Grants  of  pecuniary  aid  for  scientific  pur-  Grants  e.xpi re. 
poses  from  the  funds  of  the  Society  shall  expire  on  the 
1st  of  March  next  following,  unless  it  shall  appear  by 
a  report  that  the  recommendations  on  which  they  were 
granted  iiave  been  acted  on,  or  a  contiiuiation  of  them 
be  ordered  by  the  Council. 

XLIX.    In  grants  of  money  to  Committees  and  indi-  Peis.mai 
viduals,  the  Society  shall  not  pay  any  personal  expenses    uVne  pali"*^ 
which  may  be  incurred  by  the  Members. 

L.  No  new  law,  or  alteration  or  repeal  of  an  exist- Alterations  of 
ing  law,  shall  be  made  except  at  the  Annual  General 
Meeting  in  March,  or  at  a  Special  General  Meeting 
summoned  for  the  purpose,  as  provided  in  Law  XIX, 
and  in  pursuance  of  notice  given  at  the  preceding 
Ordinary  Meeting  of  the  Society. 

LI.    Should  any  circumstance  arise  not  provided  for  Cases  not 
in  these  Laws,  the  Council  is  empowered  to  act  as  may    p''°^i'^«^*  ^o' 
seem  to  be  best  for  the  interests  of  the  Society. 


298    Proccedln(js  of  l{o>/al  Societ//  of  Vicloria. 

LIT.  The  Council  shall  have  power  to  propose 
gentlemen  not  resident  in  Victoria,  for  election  in  the 
same  manner  as  Ordinary  Members,  as  Corresponding 
Members  of  the  Society.  Tlie  Corresponding  Members 
shall  contribute  to  the  Society  papers  which  may  he 
received  as  those  of  Ordinary  Members,  and  shall  in 
return  be  entitled  to  receive  copies  of  the  Society's 
publications. 

LIII.  Associates  shall  have  the  privileges  of  Members 
in  respect  to  the  Society's  publications,  in  joining  the 
Sections,  and  at  the  Ordinary  Meetings,  with  the  excej)- 
tion,  that  tliey  shall  not  have  the  power  of  voting;  they 
shall  also  not  be  eligible  as  Officers  of  the  Society. 


MEMBERS 

OF 

£l)c  ^icrnal  Botictn  of  yictoria 


Pat  I!  ox. 

Ho])etoun,    His  Excellency  The  Kiglit  Hon.  John   Adrian  Louis 
Hope,  G.C.M.a.,  Seventh  Earfof. 

HONOKAHY    MemBEKS. 

Agnew,  Hon.  J.  \V.,  M.E.O.,  M.!).,  Hobart,  Tasmania. 
Bancroft,  J.,  E^sq  ,  M.D.,  Brisbane,  Queensland. 
Clarke,  Colonel  Sir  Andrew,  K.C.M.G.,  C.B.,  C.I.E.,  London. 
Forrest,  Hon.  J.,  C.M.G.,  Surveyor-General,  West  Australia. 
Hector,  Sir  James,  K.C.M.G.,  M.D.,  E.R.S.,  Wellington,  K.Z. 
Liveisidge,  Professor  A.,  F.R.S.,  Univer.sity,  Sydney. 
Neumeyer,  Professor  George,  P!i.  1).,  Hainburg,  Germany. 
Russell,  H.  C,  Esq.,  F.R.S.,  F.R.A.S.,  Observatory,  Sydney,  KS.  W. 
Scott,  Rev.  W.,  M.A.,  Kurrajong  Heights,  N.S.W. 
Todd,  Charles,  Esq.,  C.M.G.,  F.R.A.S.,  Adelaide,  S.A. 
Verbeek,  Dr.  R    D.  M.,  Buitenzorg,  Batavia,  Java. 

Life  Membeus. 

Bage,  Edward,  jun.,  Esq.,  Crawford,  Fulton-street,  St.  Kilda. 
Barkly,   His    Excellency  Sir    Henry,   G.C.M.G.,   K.C.B.,   Carlton 

Club,  London. 
Bosisto,  Joseph,  Esq.,  C.M.G.,  Richmond. 
Butters,  J.  S.,  Esq.,  323  Collins-street. 

Eaton,  H.  F.,  Esq.,  Treasury,  Melbourne. 

Elliott,  T.  S.,  E.sq.,  Railway  Department,  Spencer-street. 

Elliott,  Sizar,  Esq.,  J.P.,  Were-strcet,  Biighton  Beach. 

Fowler,  Thomas  W.,  Esq.,  Carlyle-street,  Upper  Hawthorn.- 

(iibbons,   Sidney   W.,    Esq.,   F.C.S.,  care  of  Mr.    Lewis,   Chemist, 

341  Bourke  street. 
Gilbert,  J.  E.,  Esq.,  Money  Order  OtRce,  G.P.O.  Melbourne. 


300     Proceed ln(js  of  the  Royal  Society  of  Victoria. 

Hovvitt,  Edward,  Esq.,  Piathinine.s-iortd,  Auburn. 

Love,  E.  F.  J.,  Esij.,  M.A.,  Queen's  (Jollege,  University. 

Mueller,  ikron  F.  Von,  K.C.M.G.,  M.D.,  Ph.D.,  F.R.S.,  Arnold- 
street,  South  Yarra. 

Nicholas,  William,  Esq.,  E.G.S.,  Melbourne  University. 

Rusden,  11.  K.,  Esq.,  F.R.G.S.,  Ockley,  Marlton  Crescent, 
St.  Kilda. 

Selby,  (Jr.  W.,  Esq.,  99  Que^n-street. 

White,  E.  J.,  Esq.,  F.R.A.S.,  Melbourne  Observatory. 
Wilson,  Sir  Samuel,  Knt.,  Oakleigh  Hall,  East  St.  Kilda. 

Ordinary  Members. 

Allan,  Alexander  C,  Esq.,  Fitzroy-street,  St.  Kilda. 
Allan,  M.  J.,  Esq.,  17  Delbridge-street,  North  Fitzroy. 
Archer,  W.  H.,  Esq.,  J.P.,  F.L.S.,  F.I.A.,  Alverno,  Grace  Park, 
Hawthorn. 

Bage,  William,  Esq.,  M.I.C.E.,  319  Collins-street. 

Baltbui-,  Lewis  J.,  Esq.,  Tyalla,  Toorak. 

Barnard,  F.,  Esq.,  49  High-street,  Kew. 

Barnes,  Benjamin,  Esq.,  Queen's  Terrace,  South  Melbourne. 

Barracchi,  Pietro,  Esq.,  R.E.  and  C.E.  Italy,  F.R.A.S.  Eng., 
Observatory,  Melbourne. 

Barrett,  J.  W.,  Esq.,  M.D.,  34  Collins-street. 

Bevan,  Rev.  L.  D.,  LL.D.,  D.D.,  Congregational  Hall,  Russell- 
street. 

Beckx,  Gustave,  Esq.,  Queen's  Place,  St.  Kilda  Ptoad. 

Bennetts,  W.  R.,  jun.,  Esq.,  129  Rathdovvn-street,  Carlton. 

Blackett,  C  R.,  Esq.,  J. P.,  F.C.S.,  Charlesfort,  Tennyson-street, 
South  St.  Kilda. 

Brett,  J.  Talbot,  Esq.,  M.R.C.S.,  S5  Collins-street. 

Campbell,   F.   A.,    Esq.,   C.E.,   Working    Men's  College,   Latrobe- 

street. 
Candlei-,  Samuel  Curtis,  Esq.,  Melbourne  Club. 
Cohen,  Joseph  B.,  Esq.,  A.R.I.B.A.,  Public  Works  Department, 

Melbourne. 
Coane,  J.  M.,  Esq.,  C.E.,  care  of  Coane  and  Grant,  Fourth  Floor, 

Prell's  Building.s,  Queen-street. 

Danks,  John,  Esq.,  391  Bourke-street. 

David.son,  William,  Esq.,  C.  E.,  Inspector-General  of  Public 
Works,  Melbourne. 


Lid  of  Members.  801 

Dendy.  Arthur,  Esq.,  D.Sc,  F.L8.,  Uuivei-.sity,  Melbourne. 
Deiinaiit,  T.,  Esq.,  JFG.S.,  F.C.S.,  Lyiidlmr.st  Crescent,  Hawthorn, 
Dunn,  Frederick,  Esq.,  306  Littk^  Fliiiders-street 

Ellery,   R.   L.   J.,   Esq.,   O.M.(i.,    F.R.S.,    F.R.A.S.,   Observatory, 
Melbourne. 

Foord,  George,  Esq.,  F.C.S.,  Royal  Mint,  Melbourne. 
F'ox,  W.,  Esq.,  U  Robe-street,  St.  Kilda. 

Goldstein,  J.  R.  Y.,  Esq.,  Office  of  Titles,  Melbourne. 
Gotch,  J.  S.,  Esq.,  109  All>ert-street,  East  Melbourne. 
Griffiths,    G.    S.,    Esq.,    F.R.G.S.,    Waratah,     Washington-street, 

Toorak. 
Grut,   Percy  de  Jersey,   Esq.,  E.  S.   &  A.  C,   Bank,   Collins-street 

West. 

Hake,  C.  N.,  Esq.,  F.C.S.,  Melbourne  Club,  Melbourne. 
Halley,  Rev.  J.  J.,  Congregational  Hall,  Russell-street. 
Hart,  Ludovico,  Esq.,  i)  Tivoli-road,  South  Yarra. 
HefFernan,  E.  B.,  Esq  ,  M.D.,  10  Brunswick-street,  Fitzroy. 
Hogg,  H.  R.,  Esq.,  ]  6  Market  Buildings,  Flinders  Lane  West. 
Hewitt,  A.  W.,  Esq.,  P.M.,  F.G.S.,  Secretary  Mines  Department, 
Melbourne. 

Inskip,  Geo.  C,  Esq.,  F.  R.I.B.A.,  St.  James'  Buildings,  William- 
street. 

Jager,  Ernest,  Esq.,  North-street,  Ascot  Vale. 
James,  E.  M.,  Esq.,  M.R.C.S.,  2  Collins-street. 
Jamieson,  James,  Esq.,  M.D.,  56  Collins-street. 
Joseph,   R.   E.,    Esq.,    Electric    Light    Company,   Sandridge-road, 
Melbourne. 

Kernot,  Professor  W.  C,  M.A.,  C.E.,  University,  Melbourne. 

Lucas,  A.  H.  S.,  Esq.,  M.A.,  B. Sc,  Newington  College,  Sydney, 

New  South  Wales. 
Lucas,  William,  Esq.,  113  Leopold-street,  South  Yiirra. 
Lyle,  Professor  T.  R.,  M.A.,  University,  Melbourne. 
Lynch,  William,  Esq.,  St.  James'  Buildings,  William-street. 

M'Coy,   Professor    Sir   F.,   K.C.M.G.,  D.Sc,   F.R.S.,   University, 

Melbourne. 
M'Alpine,  D.,  Esq.,  F.C.S.,  5  Wallace-street,  Toorak. 
Main,  Thomas,  Esq.,  City  Surveyor's  Office,  Melbourne. 
Maloney,  William,  Esq.,  57  Roden-street,  West  Melbourne. 
Masson,  Professor  Orme,  M.A.,  D.Sc,  University,  Melbourne. 


302      Piocee(.(in(/s  of  the  iioi/al  Society  of  Victoria. 

Matliew,  Rev.  John,  Oobuig. 

Moerliii,  C,  Esq.,  Observatory,  Melbourne. 

Moors,  H.,  Esq.,  Chief  Secretary's  Office,  Melbourne. 

Morris,  Professor,  M.A.,  University,  Melbourne. 

Morrison,     Alexander,     Esq.,     M.D.,     472     Albert-street,     Ea.st 

Melbourne. 
Muntz,  T.  B.,  Esq.,  C.E.,  358  Collins-street. 

Nanson,  Professor  E.  J.,  M.A.,  University,  Melbourne. 
Neild,  J.  E.,  Esq.,  M.D.,  Biltou  Hou.se,  21  Spring-street. 
Newbery,  J.  Cosmo,  Esq.,  JJ.  Sc,  C.M.G.,  Technological  Museum 
Nimmo,  W.  H.,  Ksq.,  Melbourne  Club,  Melbourne. 

Officer,  C.  G.  W.,  Esq.,  B.Sc,  Toorak. 

Oldfield,  Lenthal,  Esq.,  36  Nicholson-street,  Fitzroy. 

Prince,  J  ,  Esq.,  Henry-street,  Windsor. 

Ridge,  Samuel  H.,  E-q.,  B.A.,  257  Victoria  Parade,  E.  Melbourne. 
Resales,     Henry,     Esq.,     F.G.S.,    Alta    Mira,    Grandview-grove, 

Aniiadale. 
Rudall,  -J.  T.,  Esq.,  F.R.C.S.,  coraer  of  Spring  and  Collins-streets, 

Melbourne. 
Rule,  O.  R.,  Esq.,  Technological  Museum,  Melbourne. 

Sargood,  Hon.  Sir  Frederick  T.,  K.C.M.G.,  M.L.C.,  Elsternwick. 

Shaw,  Thomas,  Esq.,  Woorywyrite,  Camperdown. 

Spencer,  Professor  W.  Baldwin,  M.A.,  University,  Melbourne. 

Stillwell,  A.,  Esq.,  195a  Collins  street. 

Sugden,  Rev.  E.  H.,  B.A.,  B.Sc,  Queen's  College,  Carlton. 

Sutherland,  Alex,  Esq.,  M.A  ,  Carlton  College,  Royal  Park. 

Sweet,  George,  Esq.,  Wilson-street,  Brunswick. 

Syme,  G.  A.,  Esq.,  M.B.,  F.R.C.S.,  74  Collins  street. 

Tipping,  Isaac,  Esq.,  C.E  ,  90  Henry-street,  Prahran. 
Ti.sdall,  H.  T.,  Esq.,  F.L.S.,  Washington  street,  Toorak. 
Topp,    C.    A.,    Esq.,    xM.A.,    LL.B.,    F.L.S.,    Grandview    Grove, 
Armailale. 

Whitley,  David,  Esq.,  26  Qii(!en-street 

Wight,  Gerard,  Esq.,  Phceiiix  Chambers,  Market-street. 

Country  Members. 

Bland,  R.  H.,  Esq.,  Clunes. 

Cameron,  John.  Esq.,  Orbost,  Victoria. 

Clarke,  Donald,  Esq.,  School  of  Mines,  Bairnsdale. 

Conroy,  James  Macdowall,  Esq.,  Wingham,  Manning  River,  N.S.W. 


List  of  Memhei-ii.  303 

Dawson,  J.,  Esq.,  Rennyliill,  Oamperdow!). 
Desmond,  John,  Esq.,  Warrnanibool. 
Dobson,  A.  Dudley,  Esq.,  Warrnauiltool. 

Edwards,  Henry,  Esq.,  Princess  Tlieatre,  Melbourne. 

Field,   William   Graham,   Esq.,   C.E.,  Railway  Engineer-in-Chiefs 
Department,  Melbourne. 

Hall,  T.  S.,  Esq.,  M.A.,  School  of  Mines,  (Jastlemaine. 
Hunt,  Robert,  Esq.,  Royal  Mint,  Sydney,  N.S.W. 

Ivey,  James,  Esq.,  Ballarat. 

Keogh,  Laurence  F.,  Esq.,  Brucknell  Banks,  Cobden, 

Loaghrey,  B.,  Esq.,  M.A.,  C.E.,  City  Surveyor,  Wellington,  N.Z. 

MacGillivray,  P.  H.,  Esq.,  M.A.,  M.R.C.S.,  Bendigo. 
Mackay,  J.  B.  Lillie,  Esq.,  F.C.S.,  School  of  Mines,  Bendigo. 
Manson,  Donald,  Esq.,  Elgin  Buildings,  Sydney,  N.S.W. 
M'lnday,  J.,  Esq.,  Huberton,  North  Queensland. 
Murray,  Stewart,  Esq.,  C.E.,  Kyneton. 

Naylor,  John,  Esq.,  Stawell. 

Oddie,  James,  Esq.,  Dana-sti'eet,  Ballarat. 

Oliver,  C.  E.,  Esq.,  C.E.,  Victoria  Water  Supply,  Rialto,  Collins- 
street,  Melbourne. 

Powell,    Waiter    D.    T.,    Esq.,    Hai-bour    Department,    Brisbane, 

Queensland. 
Purdie,  A.,  Esq.,  M.A.,  School  of  Mines,  Kyneton. 

Shaw,  W.  H.,  Esq..  Phrenix  Foundry,  Ballarat. 

Vickery,  S.  K.,  Esq  ,  Ararat. 

Wakelin,  T.,  Esq.,  B.A.,  Greytown,  Wellington,  New  Zealand. 
Wall,  John,  Esq.,  Town  Hall,  Sebastopol,  Ballarat. 
Williams,  Rev.  W.,  F  L.S.,  Toorak. 

Wilson,  J.   Bracebridge,  Esq.,  M.A.,  C.  of  E.  Grammar  School, 
Geelong. 

Corresponding  Members. 
Bailey,  F.  M.,  Esq.,  The  Museum,  Brisbane. 
Clarke,  Hyde,  Esq.,  32  St.  George's  Sipiare,  London,  S.W. 


304      Froceedinf/s  oj  the   Royal  Societ;/  of  Victoriu. 

Etlieridge,    Robert,    Esq.,    jiin.,    F.CI.S.,    Department   of     Mines, 

Sydnev,  N.S.W. 
Sfcirton,  James,  Esq.,  M.  U.,  F.L.S.,  15  Nowton-street,  Glasgow. 
Ulrich,  Professor  G.  H.  F.,  F.G.S  ,  Dunedin,  Otago,  N.Z. 
Wagner,  William,  Esq.,  LL.D.,  Philadelphia,  U.8.A. 

Associates. 
Askew,  David  C,  Esq.,  410  P)Oiirke-street. 

Page,  C,  Esq.,  M.D.,  81  Toorak-road,  South  Yarni. 

Paker,  Thomas,  Esq.,  Pond-street,  Abbotsford. 

Pale,  W.  M.,  Esq.,  Walpole-street,  Hyde  Park,  Kew. 

Parnaid,  P.  J.  A.,  Esq.,  M.A.,  Queen's  College,  Carlton. 

Pest,  Heinrich,  Esq.,  care  of  Mr.  H.   Falkner,  Nightingale-street, 

Palaclava. 
Pooth,  John,  Esq.,  C.E.,  Rennie-street,  Coburg. 
Powen,  W.  W.,  E.sq.,  3o8  Collins-street. 
Prockenshire,  W.  H.,  Esq.,  C.E.,  Railway  Department,  Kerang. 

Champion,  H.  V.,  Esq.,  Council  Chambers,  Williamstown. 
Chase,  L.  H.,  Esq.,  Queensberry- street,  Carlton,  or  Railway  De])art- 

ment,  Selborne  Chambers. 
Craig,  Robert,  Esq.,  Training  College,  Grattan-street. 
Cresswell,  Rev.  A.  W.,  M.A.,  St.  John's  Parsonage,  Camberwell. 

Danks,  A.  T.,  Esq.,  42  Pourke-street. 
Dawson,  W.  S.,  Esq.,  Runnymede,  Essendon. 

Edwards,  J.   E.,  Esq.,  Colonial   Telegraph  Exchange,   133  Little 
Collins-street. 

Falk,  David,   Esq.,  care  of  Messrs.  Falk  and  Co.,  Little  Collins- 
street  East. 
Fielder,  Rev.  W.,  Noiwood,  Mitchell-street,  St.  Kilda. 
Finney,  W.  H.,  Esq.,  Pridport-street,  South  Melbourne. 
Fison,  Rev.  Lorimer,  M.A.,  Essendon. 
Fraser,  J.  H.,  Esq.,  Railway  Department,  Melbourne. 

Gabriel,  J.,  Esq.,  Simi>son"s-road,  Collingwood. 

Harper,  Andrew,  Esq.,  M.A,.  Toorak. 

Hart,  J.  S.,  E.sq.,  M.A.,  P.Sc,  University,  Melbourne. 

Holmes,  W.  A.,  Esq.,  Telegraph  Engineer's  Office,  Railway  Dejai  t- 

nient.  Spencer-street,  Melbourne. 
Howard,    Douglas,    Esq.,     M.A.,     Hawthorn     Grammar     School, 

Power-street,  Hawthorn. 


List  of  Member.^.  305 

Hubbard,  J.  R.,  Esq.,  99  Queen-stieet,  Mell)Ourne. 

Ingainells,  F.  N.,  Esq.,  Observatory,  Melbourne. 

Kernot.  Frederick  A.,  Esq.,  Royal  Park,  Hotimiu. 
Kiiklaud,  J.  B.,  Esq.,  F.C.S.,  University,  Melbourne. 

Lambert,  Thomas,  p]sq.,  Bank  of  New  South  Wales,  Chiltern. 
Lindsay,  James,  JEsq.,  172  Bouverie-street,  Carlton. 

Maclean,  C.  W.,  Esq.,  Walsh-street,  South  Yarra. 

Melville,  A,  G.,  Esq.,  Mullen's  Library,  Collins-street. 

Mills,  H.  W.,  Esq.,  Glan-y-mor,  Brighton. 

Moors,  Miss  Florence  Donald,  Punt-road,  South  Yarra. 

Moors,  E.  M.,  Esq.,  University,  Sydney,  N.S.W. 

Murray,  T.,  Esq.,  C.E.,  Yictoiia  Water  Supply  Department,  Melb, 

Paul,  A.  W.  L.,  Esq  ,  Male-street,  North  Brighton. 

Phillips,  A.  E.,  Esq.,  99  Queen-street. 

Pritchard,  G.  B.,  Esq.,  Mantell-strei-t,  Moonee  Ponds. 

Quarry,  Herbert,  Esq.,  Alma  Cottage,  Macaulay-road,  Kensington. 

Remfry,  Josiah  Richard,  Esq.,  Branxholme,  Yictoria. 
Riddel!,  Mrs.,  F.S.Sc.  Lond.,  21  May-road,  Toorak. 
Robertson,  John  Steele,  Esq.,  B.A.,  Universit}',  Melbourne. 
Ro.ss,  Joseph,  Esq.,  M.D.,  Warrnambool. 

Schafei-,  R.,  Esq.,  17  Union-street,  Windsor. 

Slater,  H.  A.,  Esq.,  121  Collin.s-street. 

Smibert,  G.,  Esq.,  G.P.O.,  Melbourne. 

Smith,  Elderson,  Esq.,  London  Chartered  Bank,  410  Brunswick- 
stieet,  Fitzroy. 

Smith,  Mrs.  Elderson,  London  Chartered  Bank,  410  Brunswick- 
street  Fitzroy. 

Steele,  W.  H.,  Esq.,  jNI.A.,  Sherwood-street,  Richmond. 

Stewart,  C,  Esq.,  9  Murphy-street,  South  Yarra. 

Strettle,  W.  S.,  Esq.,  Ormond  College,  Carlton. 

Tate,  Frank,  Esq.,  B.A.,  Traitiing  College,  Grattan-street. 
Wil&more,  C.  N.,  Esq.,  B.Sc,  University,  Melbourne. 


LIST  OF  THE  INSTITUTIONS  AND  LEARNED 
SOCIETIES  THAT  RECEIVE  COPIES  OF  THE 
"TRANSACTIONS  AND  PROCEEDINGS  OF  THE 
ROYAL  SOCIETY  OF  VICTORIA." 


England. 

Agent-General  of  Victoria 

Anthropological  Institute 

Biological  Society  of  Liverpool 

Bodleian  Library 

British  Museum 

Colonial  OflBce  Library     ... 

"Electrician"    .. 

Foreign  OiEce  Library 

Geological  Society 

Institute  of  Mining  and  Mechanical  Engineers 

Institution  of  Civil  Engineers 

Linnsean  Society 

Literary  and  Philosophical  Society 

Marine  Biological  Laboratory 

Natural  History  Museum 

Naturalists'  Society 

"Nature" 

Owens  College  Library     ... 

Patent  Office,  :^ 5  Southampton  Buildings 

Philosophical  Society 

Royal  A.siatic  Society 

Royal  Astronomical  Society 

Royal  Colonial  Institute 

Royal  Gardens 

Royal  Geographical  Society 

Royal  Microscopical  Society 

Royal  Society    ... 

Statistical  Society 

University  Library 

Scotland. 
Botanical  Society 
Geological  Society 
Royal  College  of  Pljysicians'  Laboratory 


London 

London 

Liverpool 

Oxford 

London 

London 

London 

London 

London 

Newcastle 

London 

London 

Liverpool 

Plymouth 

London 

•     Bristol 

London 

Manchester 

London 

Cambridge 

London 

London 

London 

Kew 

London 

London 

London 

.     London 

Cambridge 


Edinburgh 
Edinburgh 
Edinburgh 


List  of  Institutions,  &c. 


307 


Royal  Observatory 

Royal  Physical  Society     ... 

Royal  Society   ... 

Royal  Scottish  Society  of  Arts 

Scottish  Geographical  Society 

University  Library 

University  Library 


Edinburgh 
Edinburgh 
Edinburgh 
Edinburgh 
Edinburgh 
Ediiiburgh 
.     Glasgow 


Ireland, 

Natural  History  and  Philosophical  Society 
Royal  Dublin  Society 
Royal  Geological  Society 
Royal  Irish  Academy 
Trinity  College  Library  ... 


Belfast 
Dublin 
Dublin 
Dublin 
Dublin 


Germany. 

Gessellschaft  fiir  Erdkunde              ...               ...  ...         Berlin 

Grossh.  Hessische  Geologische  Anstalt           ...  Darmstadt 

Konigl.  Botanische  Gesellschaft      ...               ...  Regensburg 

Konigl.  Offentl.  Bibliothek              ...              ...  ...     Dresden 

Konigl.  Preussische  Akademie  der  Wissenschaften       ...         Berlin 

Konigl.  Sachs  Gesellschaft  der  Wissenschaften  . . .       Leipzig 

Konigl.  Societat  der  Wissenschaften               . . .  Gottingen 

Naturfoi-schende  Gesellschaft           ...               ...  ...       Emden 

Naturforschende  Gesellschaft           ...               ...  ...        Halle 

Naturforschende  Gesellschaft           ...               ...  ...      Leipzig 

Naturliistorisch  Medizinischer  Verein             ...  Heidelberg 

Naturhistorische  Gesellschaft           ...               ...  ...   Hanover 

Naturhistorisches  Museum                ...               ...  Hamburg 

Naturhistorisches  Museum                ...               ...  ...   Hanover 

Naturwissenschaftlicher  Verein        ...               ...  ...     Bremen 

Naturwissenschaftlicher  Verein       ...               ...  Frankfurt 

Oberhessische  Gesellschaft  fiir  Natur  <k  Heilkunde        ...     Giessen 

Schlesische  Gesellschaft  fiir  Vaterland.  Cultur.  ...      Breslau 

Verein  fiir  Erdkunde        ...              ...               ...  Darmstadt 

Verein  fiir  Erdkunde        ...               ...               ...  ...         Halle 

Verein  fiir  Naturkunde   ...               ...               ...  ...       Kassel 


Austria. 

K.  K.  Akademie  der  Wissenschaften 
K.  K.  Geologische  Reichsanstalt    ... 


Wien 
Wien 


808      Proceedinf/s  of  the  Royal  Society  of  Victoria. 


K.  K.  Ueographische  Gesellschaft 
K.  K.  Naturliistorisches  Hofmuseuni 
[u.perial  Observatory 


Wien 

Wien 

Prague 


.Switzerland. 

Geographische  Gesellscliaft 
Geogr.  Couimerc.  Gesellscliaft 
Geogr.  Commerc.  Gesellscliaft 
8cliNveizerische  Naturforschendc  Gesellscliaft 
Hociete  de  Physique  et  d'Histoire  Naturelle  ... 


Berne 

St.  G alien 

Aarau 

Berne 

Geneve 


France.  - 

Acadeniie  des  Sciences  et  Belles-Lettres  et  Arts  ...  Lyon 

Feuilles  des  Jeunes  Naturalists       ...  ...  ...  Paris 

Societe  Academique  Indo-Chinoise  ...  ...  Paris 

Societe  de  Geographie     ...               ...  ...  ...  Paris 

Societe  Nationale  de  Cherbourg      ...  ...  Cherbourg 

Societe  Zoologique  de  France           ...  ...  ...  Paris 


Italy. 

Biblioteca  Nazionale  Centrale  Vittorio  Enianuele  Rome 

British  and  American  ArcluiBological  Society  ...  Elome 

Museo  di  Zoologia  ed  Anatomia  Comp.,  R.  Universita  Turin 

Ministero  dei  Lavori  Pubblici         ...              ...  ...  Rome 

Reale  Academia  di  Scienze               ...               ...  ...  Palermo 

Reale  Academia  di  Scienze,  Lettre  ed  Arti  ...  ...  Lucca 

Regia  Academia  di  Scienze,  Lettere  ed  Arti  ...  Medina 

Societa  Geogratica  Italiana               ...              ...  ...  Rome 

Societa  Toscana  di  Scienze  Naturali                ...  ...  Pisa 


Spain  and  Portugal. 

Real  Academia  de  Ciencias  Exactas,  Fisicas  y  Natu  rales 
Sociedade  de  Geographia 


Madrid 
Lisbon 


Holland  and  Belgium. 

Acadeniie  Royale  de  Belgique  ...              ...  Bruxellea 

Bataviaasch     Genootschap    van  Kunsten    en  Weten- 

schappen   ...              ...  ...              ...  ...      Batavia 

Natural  Science  Society  ...  ...              ...  Amsterdani 


List  of  Institutions,  &g. 

Natuurkundig  Genootschap 
Nederlandisch  Botan.  Vereeingiug 
Magnetical  and  Meteorological  Obsei-vatory  ... 
'>ociete  Hollandaise  des  Sciences     ... 
Societe  Macologiqiie  Royale  de  Belgique 
Societe  Provinciale  des  Arts  et  Sciences 


809 

Grouingen 

Nijinegen 

Batavia 

.  Haarlem 
Bruxelles 

.      Utrecht 


Denmark,  Sweden,  and  Nobway. 

Academie  Royale 

Koiigelige  Danske  Videnskabernes  Selskap  ... 

Societe  des  Sciences 


Copenhagen 

Copenhagen 

Christiania 


Russia  and  Roumania. 

Institut  Meteorologique  de  Roumanie            ...  Bucharest 

Jardin  Botanique  Imperial               ...               ...  St.  Petersburg 

'La  Soc.  de  Naturalists  de  I'Universite  de  Kazan  ...        Kazan 

Soc.  de  Naturalistes  Kiew                ...               ...  ...          Kiew 

Societe  des  Naturalistes  de  la  Nouvelle  Russia  ...        Odessa 

Societe  Imperiale  des  Naturalistes                   ...  ..,      Moscow 

Societe  Imjieriale  Russe  de  Geographie           ...  St.  Petersburg 


India  and  Mauritius. 

Royal  Asiatic  Society,  Ceylon  Branch  ...  ...  Colombo 

Geological  Survey  of  India  ...  ...  ...  Calcutta 

Madras  Literary  Society  ...  ...  ...  Madras 

Meteorological  Society     ...  ...  ...  ...  Mauritius 

Natural  History  Society  ...  ...  ...  Bombay 

Royal  Beugal  Asiatic  Society  ...  ...  ...  Calcutta 


China  and  Japan. 

Astronomical  Observatory  ...  ...  Hong  Kong 

China  Branch  of  the  Royal  Asiatic  Society  ...  ...  Shanghai 

Imperial  University  ...  ...  ...  ...         Tokio 

Seismological  Society  of  Japan         ...  ...  ...         Tokio 


Canada. 

Canadian  Institute  ...  ...  ...  ...     Toronto 

Geological  and  Natural  History  Survey  of  Canada  ...      Ottawa 

Royal  Society  of  Canada  .. .  ...  ...  ...Montreal 


810     Proceedings  of  the  Moijal  Society  of  Victoria. 

United  States. 

Academy  of  Natural  bciences           ...  ...  Davenport 

Academy  of  Natural  Sciences          ...  ...  Philadelphia 

Academj'  of  Sciences        ...              ...  ...  San  Francisco 

American  Academy  of  Arts  and  Soiences  ...  ...       Boston 

American  Geographical  Society        ...  ...  New  York 

American  Philosophical  Society       ...  ...  Philadelphia 

Bureau  of  Ethnology        ...              ...  ...  Washington. 

Colorado  Scientific  Society                 ...  ...  ...      Denver 

Cooper  Union  for  the  Advancement  of  Science  and  Art  New  York 

Denison  University           ..                ...  ...  ...          Ohio 

John  Hopkins  Univsr.sity                 ...  ...  Baltimore 

"Kosnios"         ...              ...              ...  ...  San  Francisco 

Maryland  Historical  Society             ...  ...  Baltimore 

Natural  Academy  of  Sciences           ...  ...  Washington 

Ofiice  of  Chief  of  Engineers,  U.S.  Army  ...  Washington 

Philoso])hical  Society        ...               ...  ...  Washington 

"Science"          ...              ...               ...  ...  New  York 

Smithsonian  Institute       ...               ...  ...  Washington 

Society  of  Natural  History               ...  ...  ...      Boston 

Society  of  Natxiral  Sciences              ...  ...  ...      Buffalo 

United  States  Geological  Survey     ...  ...  W^ashington 


Mexico. 

Ministerio  de  Fomento      ...              ...  ...  ...     Mexico 

Observatorio  Meteorologico,  Magnetico  Central  ...     Mexico 

Observatorio  Astronomico  National  ...  ...Tatubaya 

Sociedade  Cientifica,  Antonio  Al.sate  ...  ...     Mexico 

Sociedad  de  Ingenieros  de  Jalisco    ...  ...  Guadalajara 

Secretaria  de  Fomento     ...                ...  ...  Guatemala 


Argentine  Kepublic. 


Academia  de  Ciencias 
La  Museo  di  Plata 


. . .    Cordoba 
Buenos  Ayres 


Australia.  — Victoria. 


"Age" 

"Ai-gus" 

Athenajum 

Astronomical  Observatory 

Australian  Health  Society 

"  Australian  Journal  of  Pharmacy 

Chief  Secretary's  Office     ... 


Melbourne 
Melbourne 
Melbourne 
Melbourne 
Melbourne 
Melbourne 
Melbouine 


List  of  Institutions,  &c. 


311 


Department  of  Mines  and  Water  Supply 

Eclectic  Association  of  Victoria 

Field  Naturalists'  Club  of  Victoria... 

Free  library 

Free  Library 

Free  Library 

Geological  Society  of  Australasia     ... 

German  Association 

Gordon  Technical  College... 

Government  Entomologi.st 

Medical  Society 

Parliamentary  Library 

Pharmaceutical  Society  of  Australasia 

Public  Library 

Office  of  the  Government  Statist 

Royal  Geographical  Society  ..  ...  . 

Railway  Library 

School  of  Mines 

School  of  Mines 

School  of  Mines 

School  of  Mines 

School  of  Mines 

School  of  Mines 

The  Exhibition  Trustees  ... 

University  Library 

Victorian  Chamber  of  Commerce  (Manufactures) 

"  Victorian  Engineer  " 

"  Victorian  Government  Gazette"  ... 

Victorian  Institute  of  Surveyors 

Working  Men's  College,  Latrobe  Street 

Nkw  South  Wales. 

Australian  Museum 

Astronomical  Observatory 

Department  of  Agriculture 

Department  of  Mines 

Linnsean  Society  of  New  South  Wales 

Parliamentary  Library 

Public  Library 

Royal  Geographical  Society 

Royal  Society    ... 

Technological  Museum 

South  Australia. 

Parliamentary  Library 

Public  Library  and  Museum 

Royal  Society  of  South  Australia    ... 


Melbourne 

Melbourne 

Melbourne 

Echuca 

. . .    Geelong 

Bendigo 

Melbourne 

Melbourne 

...  Geelong 
Melbourne 
Melbourne 
Melbourne 
Mell)Ourne 
Melbourne 
Melbourne 
Melbourne 
Melbourne 

. . .    Ballarat 

Castlemaine 

Bendigo 

Mai-y  borough 

Bairnsdale 

...  Stawell 
Melbourne 
Melbourne 
Melbourne 
Melbourne 
Melbourne 
Melbourne 
Melbourne 


Sydney 
Sydney 
Sydney 
Sydney 
Sydney 
Sydney 
Sydney 
Sydney 
Sydney 
Sydney 


Adelaide 
Adelaide 
Adelaide 


ol2      Proceedingti  of  the  Rof/al  Society  of  Victoria. 

QUKKNS;.ANU. 


Purlianientary  Library     ... 
Public  Library ... 
Ro\  al  Geographical  Society 
Roval  Society  of  Queensland 


Brisbane 
Brisbane 
Brisbane 
Brisbane 


Tasmania. 


Parliamentary  Libi-ar}'      ... 

Public  Li])rary  ... 

Royal  Society  of  Tasmania 


Hobart 
HoV)art 
Hobart 


New  Zealand. 

Auckland  Institute  and  Museum     ...  ...  Auckland 

Colonial  Museum  and  Geological  Survey  Department    Wellington 

New  Zealand  Institute  ...  Wellington 

Otago  Institute                   ...                .  ...  Dunedin 

Parliamentary  Library     ...               ...  ...  Wellington 

Public  Library                    ...               ...  ...  Wellington 


«TILI,WEr.L   AND   CO.,    PBINTKKS,    195A    COLLINS   STREET,    MELBOURNE. 


PROCEEDINGS. 


niOCEEDINGS 

OF     THE  -    .      ;  ^_  ;. 

Jlogal  Soact^   of  0utona. 

VOL.    V    (New    Series). 

Edited    under    the    Authority    of   the    Coxincil. 
ISSUED   MAY  1893. 


THE   AUTHORS   OF  THE   SKVERAL   PAPERS   ARE  SOLELY  RESPONSIBLE   FOR    THE    SOUNDNESS   OF 
THE  OPINIONS   GIVKN   AND   fOR   THE   ACCURACY  OF  THE  STATEMENTS  MADE  THEREIN. 


MELBOURNE : 
STILLWELL    AND    CO.,    PEINTERS,    195a    COLLINS    STREET. 


AGENTS    TO    THE   SOCIETY: 

WILLIAMS  &  NORGATE,   14  HENRIETTA  STREET,   COVENT  GARDEN,   LONDON 

To  whom  all  communicaUons  for  transmission  to  the  Roj-al  Society  of  Victoria, 

from  all  parts  of  Europe,  should  be  sent. 

1803. 


,lf]^L^f°>  LIbran 


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